cem1: ProtNLM2 function-description review

Source: pre-release post-processed-2026_02_28k.xml, accession O94297. The exact entry is preserved in cem1-protnlm-source.xml; all original evidence elements and model scores are retained in cem1-protnlm-source.json. This record is currently Swiss-Prot and is absent from the published 26,856-record TrEMBL pilot list. The current public ProtNLM endpoint returns this record; API availability is distinct from membership in the published pilot list. No training-membership inference is made.

Original paragraph 1

Involved in the type II fatty acid elongation cycle. Catalyzes the elongation of a wide range of acyl-ACP by the addition of two carbons from malonyl-ACP to an acyl acceptor. Can efficiently catalyze the conversion of palmitoleoyl-ACP (cis-hexadec-9-enoyl-ACP) to cis-vaccenoyl-ACP (cis-octadec-11-enoyl-ACP), an essential step in the thermal regulation of fatty acid composition.

Original evidence key(s): 2.

Atomic claim Assessment Evidence and limit
Participation in type II fatty-acid elongation CNN Diagnostic ketoacyl-synthase domains and the mitochondrial OXSM/Cem1 assignment support this conserved role. Human OXSM activity and complementation of budding-yeast cem1 establish a transferable mitochondrial mechanism (PMID:15668256).
Two-carbon addition from malonyl-ACP to an acyl acceptor CNN This is the conserved decarboxylative condensation chemistry of the mitochondrial ketoacyl-ACP synthase family. The target retains the annotated catalytic Cys170/His311/His351 residues, and UniProt explicitly transfers the reaction from characterized human OXSM.
Elongation of a wide range of acyl-ACP substrates CNN as broad family-level capability; target range unresolved Human OXSM directly elongates C2–C14 acyl substrates and restores growth of budding-yeast cem1. This supports broad chain-extension capability in a conserved mitochondrial homolog, while it does not measure which chain lengths S. pombe Cem1 efficiently accepts.
Efficient palmitoleoyl-ACP to cis-vaccenoyl-ACP conversion UNC The inspected mitochondrial OXSM experiments establish C2–C14 chain extension and do not establish this exact unsaturated C16 substrate preference in S. pombe. The conserved enzyme family and the presence of a synthase-2 domain label do not resolve target substrate preference. The wording also abbreviates an entire elongation cycle: the synthase itself yields a beta-ketoacyl intermediate, with further reactions required to yield the fully reduced elongated acyl chain.
Essential step in thermal regulation of fatty-acid composition UNC The available mitochondrial OXSM/Cem1 evidence concerns mitochondrial acyl-chain synthesis and lipoate-related metabolism. It neither demonstrates nor excludes an essential thermal-regulation role for S. pombe Cem1. Bacterial FabF thermoregulation cannot establish that target-specific physiological claim.

Primary evidence

Family and feature provenance: cem1-uniprot.txt.