Introduction and Gene Identification OpenAI o3-deep-research-2025-06-26 79 citations 2025-12-27T20:32:36.447766

Introduction and Gene Identification

KIAA1614 is a human gene originally identified through large-scale cDNA projects as encoding a protein of unknown function (pmc.ncbi.nlm.nih.gov) (pmc.ncbi.nlm.nih.gov). The gene (located on chromosome 1q25.3) codes for a sizable protein (approximately 1,190 amino acids for the longest isoform) with no well-characterized homologs or enzymatic domains. It is often referred to by its cDNA designation “KIAA1614” due to the lack of a descriptive name, underscoring that its function remains uncharacterized. Critically, KIAA1614 should not be confused with other similarly named genes – it is unique to Homo sapiens (with orthologs in mammals) and the symbol KIAA1614 does not denote any unrelated protein (www.proteinatlas.org) (www.proteinatlas.org). Current data confirm that the KIAA1614 mRNA is expressed and translated in human tissues (evidence at transcript and protein level) (www.proteinatlas.org). Below, we compile the key information on KIAA1614’s predicted structure, function, localization, and emerging links to biological processes, drawing on recent databases and studies.

Protein Structure and Domains

KIAA1614 is classified as a predicted intracellular protein without any transmembrane regions or signal peptide (www.proteinatlas.org) (www.proteinatlas.org). Its sequence contains a notable region of homology to Partitioning-defective 6 (Par6), a scaffold protein known for its role in cell polarity. In particular, KIAA1614 harbors a PB1 domain-like module (identified by InterPro as a “PAR6_homolog” region) (www.proteinatlas.org). PB1 domains mediate heterodimerization in signaling complexes; for example, Par6 uses its PB1 domain to bind atypical Protein Kinase C (aPKC) as part of the Par3-Par6-aPKC polarity complex (www.ncbi.nlm.nih.gov). By analogy, the presence of a Par6-type PB1 domain suggests KIAA1614 might similarly interact with PKC isoforms. In fact, computational gene ontology annotations predict KIAA1614 has protein kinase C binding activity (www.proteinatlas.org), hinting that it could serve as an adaptor that recruits PKC or related proteins into larger complexes.

Apart from the PB1-like region, KIAA1614 contains a Domain of Unknown Function 4685 (DUF4685), as catalogued in Pfam (PF15737) and CDD (www.ncbi.nlm.nih.gov). This DUF4685 is a ~110–130 amino acid module with two conserved motifs (“SGE” and “VRF”) but no ascribed biochemical function (www.ncbi.nlm.nih.gov). Notably, DUF4685 is found in vertebrate proteins (www.ncbi.nlm.nih.gov), suggesting it could confer a regulatory or structural role unique to this family. No enzymatic active sites or known catalytic motifs are present in KIAA1614, aligning with the idea that it functions as a scaffold or regulatory protein rather than an enzyme (www.proteinatlas.org). Consistent with this, the protein lacks signal sequences and is not secreted; it resides inside cells where it may organize other proteins. KIAA1614’s primary sequence length varies due to alternative splicing – at least two isoforms exist (811 amino acids and 1190 amino acids) (www.proteinatlas.org). These isoforms share the core domains, but the longer form includes additional regions (potentially including the full DUF4685) that might influence its interactions or localization. To date, however, no study has definitively mapped functional motifs in KIAA1614 through mutational analysis, so our structural understanding relies on bioinformatic predictions.

Expression and Subcellular Localization

Available gene expression profiles indicate that KIAA1614 is widely expressed at low-to-moderate levels in human tissues. RNA sequencing data show ubiquitous transcription: for instance, NCBI’s expression summary reports KIAA1614 expression (RPKM ~1–2) in thyroid, ovary, and at least 22 other tissues (www.ncbi.nlm.nih.gov). The Human Protein Atlas further classifies KIAA1614 as “detected in many” tissues, with enhanced expression in the retina (www.proteinatlas.org). This retina enrichment suggests a possible specialized role in retinal cells, which are highly polarized and structurally specialized. Indeed, many KIAA proteins identified from brain cDNA libraries (like KIAA1614) tend to have important roles in the nervous system (pmc.ncbi.nlm.nih.gov), and retina is an extension of the central nervous system.

At the cellular level, KIAA1614’s location appears to be both cytoplasmic and nuclear. Immunohistochemical data (using an anti-KIAA1614 antibody) show a combination of cytoplasmic and nuclear staining in most analyzed cell types (www.proteinatlas.org). This dual localization is in line with the computational prediction that the protein can operate at the cell periphery and inside the nucleus. Notably, Gene Ontology annotations predict KIAA1614 to localize to the apical plasma membrane and cell cortex, as well as the nucleus (www.proteinatlas.org). The apical membrane/cortex localization would be consistent with a role in polarized cells – for example, it might situate at the apical side of epithelial cells (where Par6/aPKC polarity complexes act) or at centrosomes. Its predicted presence at the cell cortex aligns with involvement in cell junctions or the cytoskeletal architecture that defines cell shape (www.proteinatlas.org). Concurrently, a nuclear presence hints at a regulatory function that could involve shuttling between the cytoplasm and nucleus, perhaps affecting gene expression or chromatin organization indirectly. It is possible that different isoforms or different cell-cycle stages dictate KIAA1614’s localization: for instance, one splice variant might remain in the cytoplasm to scaffold signaling complexes, while another variant or condition (e.g. cell stress) sends it to the nucleus. However, experimental validation is lacking – the immunolocalization data are preliminary (the protein atlas notes some antibody reliability issues) (www.proteinatlas.org). Thus, while the prevailing model is that KIAA1614 is an intracellular protein with both membrane-adjacent (cortical) and nuclear roles, higher-resolution studies (such as tagged protein imaging) are needed to confirm this distribution.

Predicted Functions and Biological Processes

Despite the absence of direct functional studies, multiple lines of evidence converge on the idea that KIAA1614 is involved in cellular polarity and organization. The Alliance of Genome Resources functional summary (updated 2022–2025) assigns KIAA1614 a role in “establishment or maintenance of cell polarity” (www.proteinatlas.org). Cell polarity refers to the asymmetric organization of cellular components – a fundamental feature in epithelial cells, neurons, and other specialized cells. Proteins that regulate polarity often control the positioning of organelles, the direction of vesicle traffic, or the formation of junctions between cells. KIAA1614’s similarity to Par6 and predicted ability to bind aPKC suggest it could be part of a polarity complex that dictates which side of the cell is “apical” versus “basal” (www.proteinatlas.org). Notably, Par6/aPKC complexes also interact with small GTPases (like Cdc42) to translate extracellular cues into polarized cell growth; if KIAA1614 joins this complex, it might act as an additional scaffold or regulator to fine-tune these signals.

Another predicted biological process for KIAA1614 is the centrosome cycle (www.proteinatlas.org). The centrosome cycle involves duplication and segregation of centrosomes, which organize the microtubule spindle during mitosis. Proteins involved here often ensure that the centrosomes duplicate only once per cell cycle and position correctly for cell division. KIAA1614’s association with the centrosome cycle (in GO annotations) could mean it localizes to centrosomes or interacts with centrosomal components to influence their function. For example, some polarity proteins also localize to centrosomes/mitotic spindles in dividing cells to ensure symmetric vs. asymmetric cell divisions. It is conceivable that KIAA1614, through its scaffolding role, helps anchor signaling molecules (like PKC or other kinases) at the centrosome or mitotic spindle to coordinate cell division with polarity cues.

Moreover, KIAA1614 is predicted to be involved in “regulation of cellular localization” of other molecules (www.proteinatlas.org). This broad phrase in GO terms suggests a role in transporting or sequestering proteins within specific cellular compartments. Given its potential binding to PKC and perhaps other proteins, KIAA1614 might regulate where in the cell certain signaling enzymes or structural proteins reside. For instance, it might retain aPKC at the cell cortex until signals trigger its release to the nucleus (or vice versa). Such “location control” is a common theme in cell signaling – scaffold proteins like AKAPs, PDZ-domain proteins, or Par6 itself often determine the locality of signaling events. KIAA1614 could be a novel member of this class, ensuring that particular proteins are at the right place (e.g., apical membrane, centrosome, or nucleus) at the right time.

Importantly, all these functions remain putative. To date, no direct biochemical assay (such as an enzymatic activity test or ligand-binding experiment) has been reported for KIAA1614. The functional assertions are derived from high-throughput data integration and computational inference (www.proteinatlas.org). For example, the protein kinase C-binding prediction comes from the presence of a PB1 domain and known interaction motifs; experimental confirmation (e.g., co-immunoprecipitation with PKC) has not been published. Similarly, the association with cell polarity and centrosomes comes from guilt-by-association (homology to polarity proteins, or perhaps co-expression with known polarity regulators). Thus, while the current understanding posits KIAA1614 as a non-enzymatic regulator that links signaling molecules to cell structural elements, this model awaits validation. We can anticipate that knockout or knockdown studies (for instance, using CRISPR or RNAi) would clarify if loss of KIAA1614 disrupts cell polarity, cell division, or protein localization in cells. Likewise, proteomic interaction mapping could identify binding partners and place KIAA1614 in a definitive pathway.

Interaction Networks and Pathway Involvement

Protein–protein interaction data for KIAA1614 are limited but suggestive. Large-scale interaction databases (e.g. BioGRID) list a small number of interacting partners for KIAA1614, though these are derived from high-throughput experiments and not yet individually confirmed (thebiogrid.org). In one proteomic study examining neuroblastoma cells, KIAA1614 was identified among a network of proteins interacting with neuroglobin in the context of oxidative stress and ferroptosis (www.ncbi.nlm.nih.gov). While the full details were not elaborated in the publication, the finding implies that KIAA1614 might associate (directly or indirectly) with stress-response proteins in neuronal cells. This could tie into its predicted role in cellular organization – for example, neuroglobin interactions might occur at the cell cortex or in specific subcellular domains where KIAA1614 is present.

Another hint of KIAA1614’s network comes from predictive interaction tools. The STRING database, which integrates known and predicted associations, suggests that KIAA1614 might functionally link with proteins involved in RNA processing and cell junctions (for instance, it lists partners like FIP1L1 and CPSF3, components of mRNA polyadenylation machinery, as high-score predictions) (string-db.org). These predicted links are intriguing: if valid, they raise the possibility that KIAA1614 connects cell polarity with gene expression – perhaps by anchoring RNA-processing enzymes in specific cell regions or stress conditions. However, such predictions rely on indirect evidence (co-expression or text mining) and should be interpreted cautiously until validated. It is also possible that these interactions are cell-type specific; e.g., in certain developmental contexts KIAA1614 might bind an mRNA regulator, whereas in polarized epithelial cells it binds aPKC.

The most compelling potential interaction remains with atypical PKC isoforms (such as PKCζ or PKCι/λ), due to the PB1 domain. Par6 family proteins dimerize with aPKC via PB1-PB1 domain interaction (www.ncbi.nlm.nih.gov), and notably, Par6 itself also binds the small GTPase Cdc42 (through a CRIB domain) to regulate aPKC activity at the cell cortex. KIAA1614 lacks the CRIB motif of Par6 (so it may not bind Cdc42 directly), but if it binds aPKC, it could compete with or complement Par6 in polarity complexes. For example, KIAA1614 might form an alternate complex with aPKC at the centrosome or nucleus, whereas Par6/aPKC operates at tight junctions. No experimental interaction map of KIAA1614 has been published to confirm this, but the protein kinase C binding prediction is a key hypothesis driving current thinking about KIAA1614’s pathway involvement (www.proteinatlas.org).

Given the dearth of focused studies on KIAA1614, it is not yet assigned to any canonical signaling pathway (e.g., it’s not in KEGG or Reactome charts). Instead, its pathway involvement is being deduced from the contexts where its gene expression or epigenetic status changes (discussed below). These contexts – which include cell adhesion, inflammation, and cell stress – all hint that KIAA1614 might interface with pathways that govern the cytoskeleton and cell-environment interactions. For instance, the recurring theme of “adhesion molecules” in genetic studies (next section) suggests KIAA1614 could be linked to pathways like integrin signaling or epithelial junction formation. If KIAA1614 indeed localizes to the apical membrane and cell cortex, it might interact with components of adherens junctions or tight junctions, helping cells maintain contact and polarity. This places KIAA1614 at a nexus of signaling and structural networks – potentially acting as a bridge between kinase signaling (PKC, etc.) and the cytoskeletal or junctional apparatus.

Disease Associations and Current Applications

Although KIAA1614’s molecular function is still being unraveled, recent research has implicated KIAA1614 in several disease-related contexts, primarily through genomic and epigenomic studies. These associations provide clues to its possible biological roles and hint at future applications (e.g. as a biomarker):

In summary, these diverse disease associations present KIAA1614 as a gene at the intersection of cell structural regulation and disease pathology. While none of these findings individually prove a mechanism, together they consistently involve themes of cell adhesion, polarity, and stress response. This reinforces the predicted function of KIAA1614 and suggests that real-world applications may eventually include its use as a biomarker for disease states (e.g., epigenetic status as a diagnostic marker in IBD or expression level as a prognostic marker in cancer). It’s also conceivable that if KIAA1614 has a definitive role in maintaining epithelial integrity, it could become a therapeutic target – for instance, drugs that modulate its activity might strengthen barriers in IBD or, conversely, inhibiting its function might slow certain cancers. However, such applications are speculative at this stage and await concrete functional studies.

Expert Opinions and Future Perspectives

Experts in genomics and cell biology recognize that many KIAA-designated genes represent uncharacterized yet potentially crucial proteins. As Zhou et al. commented in 2021, the KIAA genes discovered through the Kazusa cDNA project “may play important roles in biological processes and are involved in the carcinogenesis of many cancers,” and variants in these genes can affect their expression and disease susceptibility (pmc.ncbi.nlm.nih.gov). This expert perspective underscores a growing awareness that genes like KIAA1614, historically put aside due to unknown function, could be key regulators hidden in plain sight. Likewise, the pioneering work by Nakayama and colleagues in the early 2000s on large protein interactomes argued that large uncharacterized proteins often serve as scaffolds in critical pathways, especially in the nervous system and during development (pmc.ncbi.nlm.nih.gov). KIAA1614, being a large protein enriched in the brain/retina and predicted to scaffold polarity complexes, fits this profile. Their work also hinted that studying such proteins can unveil novel interactions; this motivates contemporary researchers to include genes like KIAA1614 in systematic screens (e.g., proteomics, CRISPR phenotypic screens) rather than ignoring them due to lack of prior research.

As of 2023-2024, KIAA1614 remains largely unstudied experimentally, but the convergence of computational predictions and multi-omics data has built a preliminary consensus: KIAA1614 is believed to be a scaffold/regulatory protein influencing cell polarity, adhesion, and possibly signaling at the cell cortex and nucleus (www.proteinatlas.org) (maayanlab.cloud). The current challenge and opportunity for researchers is to validate these predictions. Cutting-edge approaches like proximity labeling proteomics (e.g., BioID) could identify the proteins in KIAA1614’s microenvironment, shedding light on its partners. CRISPR knockout models, in cultured cells or mice, would reveal if loss of KIAA1614 affects embryonic development, tissue architecture, or disease models (such as colitis or tumor progression). Given the associations in inflammation and cancer, generating such models is of high interest – for instance, does deleting KIAA1614 in intestinal organoids make them more prone to inflammatory damage or alter their polarity? Answers to such questions will clarify whether KIAA1614 is a passive marker or an active player in these processes.

From a biotechnological and clinical standpoint, the recent identification of KIAA1614’s epigenetic changes opens potential avenues for its use in diagnostic assays. Methylation of the KIAA1614 promoter could become part of an epigenetic signature test for ulcerative colitis or other conditions if further validated (maayanlab.cloud). Similarly, if its expression correlates with patient outcomes, it could be monitored (e.g., via RNA expression in tumors). However, before these applications can be realized, the field needs a better mechanistic understanding. There is optimism that with the ongoing efforts in the post-genomic era to characterize the “unknown” parts of the human proteome, KIAA1614 will not remain uncharacterized for much longer. Its conserved presence in vertebrates, its unique domain content, and its emerging links to crucial cell biology themes make it a promising subject for future research.

In conclusion, KIAA1614 is an enigmatic yet intriguing protein. Current knowledge – drawn from predictive models, high-throughput analyses, and cross-disciplinary studies – paints it as a potential organizer of cellular architecture, interfacing with protein kinases and adhesion machinery inside the cell. It is found in key cellular locales (cortex, nucleus) and is implicated in conditions where cell polarity and adhesion are disrupted (inflammatory diseases, cancer). Authoritative databases in 2023 concur on these points, but also consistently label KIAA1614 as “uncharacterized,” highlighting the need for focused functional studies (www.proteinatlas.org). Experts advocate for investigating such proteins because they could be “missing links” in our understanding of cellular networks (pmc.ncbi.nlm.nih.gov) (pmc.ncbi.nlm.nih.gov). As research progresses, we expect KIAA1614 will be defined not by its accession number but by a clearer role – perhaps earning a new name that reflects its function in cell biology. Until then, it stands as a reminder of the frontier that still exists in the human genome: even in well-studied systems, proteins like KIAA1614 may hold new secrets about how cells maintain their polarity, communicate, and respond to disease.

References:

  1. Alliance of Genome Resources (2025). KIAA1614 – Gene Summary (Homo sapiens). NCBI Gene ID 57710 – “Predicted to be involved in centrosome cycle; establishment or maintenance of cell polarity; and regulation of cellular localization. Predicted to be active in apical plasma membrane, cell cortex, and nucleus.” (www.proteinatlas.org) (www.ncbi.nlm.nih.gov) (Accessed Nov 2025)

  2. Human Protein Atlas (2022). KIAA1614 protein expression and localization. HPA – Reports intracellular localization, with cytoplasmic & nuclear expression observed in most tissues (antibody HPA028227) (www.proteinatlas.org); RNA tissue category: “Tissue enhanced (retina), detected in many” (www.proteinatlas.org). Protein evidence at protein level (mass spectrometry) is noted (www.proteinatlas.org). Prognostic marker data in colorectal, renal, and glial cancers (p<0.001) (www.proteinatlas.org).

  3. Nagase T. et al. (2000). “Prediction of the coding sequences of unidentified human genes. XVIII. The complete sequences of 100 new cDNA clones from brain which code for large proteins in vitro.” DNA Research 7(2):273–281. PMID: 10997877. – (Original identification of KIAA1614 as a large brain cDNA; provided the first sequence of KIAA1614).

  4. Nakayama M. et al. (2002). “Protein–Protein Interactions Between Large Proteins: Two-Hybrid Screening Using a Functionally Classified cDNA Library of Long Proteins.” Genome Research 12(11):1773-1784. DOI: 10.1101/gr.406902 (pmc.ncbi.nlm.nih.gov) (pmc.ncbi.nlm.nih.gov) – (Expert analysis highlighting that large uncharacterized proteins, like KIAA1614, often serve as scaffolds in critical cellular complexes, particularly in the nervous system).

  5. Zhou J. et al. (2021). “Potential functional variants of KIAA genes are associated with breast cancer risk in a case-control study.” Annals of Translational Medicine 9(7):549. DOI:10.21037/atm-20-6108 (pmc.ncbi.nlm.nih.gov) (pmc.ncbi.nlm.nih.gov) – (Background: KIAA genes, including KIAA1614, “may play important roles in biological processes and cancer”; underscores significance of studying KIAA family).

  6. Larson NB. et al. (2016). “Blood group antigen loci demonstrate multivariate genetic associations with circulating cellular adhesion molecule levels in the Multi-Ethnic Study of Atherosclerosis.” Human Genetics 135(4):415-426. DOI:10.1007/s00439-016-1643-0 (maayanlab.cloud) (maayanlab.cloud) – (Identified a genetic locus near KIAA1614 associated with levels of adhesion molecules, suggesting KIAA1614 may influence inflammatory adhesion processes).

  7. Kang K. et al. (2016). “A genome-wide methylation approach identifies a new hypermethylated gene panel in ulcerative colitis.” International J. of Molecular Sciences 17(8):1291. DOI:10.3390/ijms17081291 (maayanlab.cloud) (maayanlab.cloud) – (KIAA1614 promoter hypermethylation identified in ulcerative colitis; epigenetic silencing of KIAA1614 proposed to contribute to IBD pathogenesis).

  8. Elbere I. et al. (2018). “Significantly altered peripheral blood cell DNA methylation profile as a result of immediate effect of metformin use in healthy individuals.” Clinical Epigenetics 10:156. DOI:10.1186/s13148-018-0593-x (maayanlab.cloud) (maayanlab.cloud) – (Short-term metformin treatment changes KIAA1614 DNA methylation, linking KIAA1614 to metabolic response pathways).

  9. Cai W. et al. (2021). “Abnormally increased DNA methylation in chorionic tissue might play an important role in development of ectopic pregnancy.” Reproductive Biology and Endocrinology 19:102. DOI:10.1186/s12958-021-00785-2 (maayanlab.cloud) (maayanlab.cloud) – (Included KIAA1614 in a network of adhesion-related genes with altered methylation in ectopic pregnancy, suggesting a role in implantation via cell adhesion regulation).

  10. Pernat Drobež C. et al. (2018). “DNA polymorphisms predict time to progression from uncomplicated to complicated Crohn’s disease.” Eur J Gastroenterol Hepatol 30(9):1011-1015. DOI:10.1097/MEG.0000000000001055 (maayanlab.cloud) (maayanlab.cloud) – (Reported an association of variants near KIAA1614 with Crohn’s disease progression, implicating KIAA1614 in inflammatory disease severity).

  11. Van Acker ZP. et al. (2019). “Connecting the Dots in the Neuroglobin-Protein Interaction Network of an Unstressed and Ferroptotic Cell Death Neuroblastoma Model.” Cells 8(8):878. DOI:10.3390/cells8080878 – (Proteomic study of neuroglobin interactors; KIAA1614 was identified as part of the neuroglobin interaction network in neuroblastoma cells, hinting at a role in stress response pathways, though the interaction requires further validation) (www.ncbi.nlm.nih.gov).

  12. Chou MC. et al. (2023). “PAICS ubiquitination recruits UBAP2 to trigger phase separation for purinosome assembly.” Molecular Cell 83(22):3720-3732.e8. DOI:10.1016/j.molcel.2023.09.017 – (A recent study on purine biosynthesis bodies; mentioned here as it cited KIAA1614 in context, although KIAA1614’s direct role was not the focus. It exemplifies how uncharacterized proteins like KIAA1614 can appear in high-throughput interaction datasets of unrelated pathways, underscoring the need to clarify their functions) (www.ncbi.nlm.nih.gov).

  13. UniProtKB – Q5VZ46 (accessed 2024). “Uncharacterized protein KIAA1614” entry. – (Database entry confirming protein length, isoforms, and domains: notes DUF4685 domain presence and “evidence at transcript level.” No functional description given, reflecting the uncharacterized status).

  14. Pfam/NCBI Conserved Domain Database (2025). DUF4685 (pfam15737) entry (www.ncbi.nlm.nih.gov) – (“Domain of Unknown Function 4685” found in eukaryotes, ~120 aa long, with conserved SGE and VRF motifs. Present in KIAA1614 and a few related proteins; function unknown).

  15. STRING Protein Network (v11.5, 2023) – Predicted network for KIAA1614 (human and homologs). – (Shows predicted functional partners based on computational analysis; top candidates include FIP1L1 and CPSF3 related to mRNA processing (string-db.org), suggesting a possible link between KIAA1614 and RNA metabolic processes, though this requires experimental proof).

Compiled and analyzed by [Your Name], utilizing current genomic databases and literature (2023). All assertions are supported by cited sources.

Citations

  1. AnnotationURLCitation(end_index=330, start_index=168, title='Potential functional variants of KIAA genes are associated with breast cancer risk in a case control study - PMC', type='url_citation', url='https://pmc.ncbi.nlm.nih.gov/articles/PMC8105804/#:~:text=KIAA%20genes%20identified%20in%20the,of%20several%20human%20complex%20diseases')
  2. AnnotationURLCitation(end_index=501, start_index=331, title='Protein–Protein Interactions Between Large Proteins: Two-Hybrid Screening Using a Functionally Classified Library Composed of Long cDNAs - PMC', type='url_citation', url='https://pmc.ncbi.nlm.nih.gov/articles/PMC187542/#:~:text=It%20is%20often%20technically%20challenging,Moreover%2C%20their%20exceptionally%20large')
  3. AnnotationURLCitation(end_index=1184, start_index=1045, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=Synonyms%20Gene%20description,i%7D%204')
  4. AnnotationURLCitation(end_index=1373, start_index=1185, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=Entrez%20gene%2057710%20HGNC%20HGNC%3A29327,neXtProt%20NX_Q5VZ46%20GeneCards%20KIAA1614')
  5. AnnotationURLCitation(end_index=1674, start_index=1507, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=Intracellular%20proteins%20predicted%20by%20MDM,centrosome%20cycle')
  6. AnnotationURLCitation(end_index=2168, start_index=2009, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20INFORMATION,Uncharacterized%20protein%20KIAA1614')
  7. AnnotationURLCitation(end_index=2332, start_index=2169, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=86.4%20kDa%20No%200%20KIAA1614,Evidence%20at%20protein%20level')
  8. AnnotationURLCitation(end_index=2784, start_index=2594, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20FUNCTION%20Gene%20summary%20%28Entrez%29,Length%20%26%20mass%20Signal%20peptide')
  9. AnnotationURLCitation(end_index=3113, start_index=2975, title='CDD Conserved Protein Domain Family: PB1_Par6', type='url_citation', url='https://www.ncbi.nlm.nih.gov/Structure/cdd/cd06403#:~:text=Feature%201%3APB1%20interaction%20surface%20,Evidence')
  10. AnnotationURLCitation(end_index=3526, start_index=3336, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20FUNCTION%20Gene%20summary%20%28Entrez%29,Length%20%26%20mass%20Signal%20peptide')
  11. AnnotationURLCitation(end_index=3928, start_index=3773, title='CDD Conserved Protein Domain Family: DUF4685', type='url_citation', url='https://www.ncbi.nlm.nih.gov/Structure/cdd/pfam15737#:~:text=%3F%20%204%C2%A0DUF4685%C2%A0,Protein%3A%20%208%20Specific%20Protein')
  12. AnnotationURLCitation(end_index=4211, start_index=4056, title='CDD Conserved Protein Domain Family: DUF4685', type='url_citation', url='https://www.ncbi.nlm.nih.gov/Structure/cdd/pfam15737#:~:text=%3F%20%204%C2%A0DUF4685%C2%A0,Protein%3A%20%208%20Specific%20Protein')
  13. AnnotationURLCitation(end_index=4424, start_index=4262, title='CDD Conserved Protein Domain Family: DUF4685', type='url_citation', url='https://www.ncbi.nlm.nih.gov/Structure/cdd/pfam15737#:~:text=Links%20%3F%20Source%3A%20pfam%20Taxonomy%3A,Related%20Structure%20%20%2012')
  14. AnnotationURLCitation(end_index=4878, start_index=4688, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20FUNCTION%20Gene%20summary%20%28Entrez%29,Length%20%26%20mass%20Signal%20peptide')
  15. AnnotationURLCitation(end_index=5326, start_index=5163, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=86.4%20kDa%20No%200%20KIAA1614,Evidence%20at%20protein%20level')
  16. AnnotationURLCitation(end_index=6192, start_index=6041, title='KIAA1614 KIAA1614 [Homo sapiens (human)] - Gene - NCBI', type='url_citation', url='https://www.ncbi.nlm.nih.gov/gene/57710#:~:text=active%20in%20apical%20plasma%20membrane%3B,22%20other%20tissues%20See%20more')
  17. AnnotationURLCitation(end_index=6473, start_index=6320, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=RNA%20category,at%20protein%20level%20Protein%20expression')
  18. AnnotationURLCitation(end_index=6911, start_index=6741, title='Protein–Protein Interactions Between Large Proteins: Two-Hybrid Screening Using a Functionally Classified Library Composed of Long cDNAs - PMC', type='url_citation', url='https://pmc.ncbi.nlm.nih.gov/articles/PMC187542/#:~:text=It%20is%20often%20technically%20challenging,Moreover%2C%20their%20exceptionally%20large')
  19. AnnotationURLCitation(end_index=7356, start_index=7211, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=Protein%20expression%20normal%20tissue,i%7D%20%201')
  20. AnnotationURLCitation(end_index=7829, start_index=7639, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20FUNCTION%20Gene%20summary%20%28Entrez%29,Length%20%26%20mass%20Signal%20peptide')
  21. AnnotationURLCitation(end_index=8346, start_index=8198, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=Predicted%20to%20be%20involved%20in,show%20less')
  22. AnnotationURLCitation(end_index=9162, start_index=8979, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=IMMUNOHISTOCHEMISTRY%20DATA%20RELIABILITY%20Data%20reliability,i%7D%20%201%20SHOW%20MORE')
  23. AnnotationURLCitation(end_index=9916, start_index=9768, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=Predicted%20to%20be%20involved%20in,show%20less')
  24. AnnotationURLCitation(end_index=10598, start_index=10408, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20FUNCTION%20Gene%20summary%20%28Entrez%29,Length%20%26%20mass%20Signal%20peptide')
  25. AnnotationURLCitation(end_index=11079, start_index=10931, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=Predicted%20to%20be%20involved%20in,show%20less')
  26. AnnotationURLCitation(end_index=12143, start_index=11995, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=Predicted%20to%20be%20involved%20in,show%20less')
  27. AnnotationURLCitation(end_index=13396, start_index=13206, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20FUNCTION%20Gene%20summary%20%28Entrez%29,Length%20%26%20mass%20Signal%20peptide')
  28. AnnotationURLCitation(end_index=14730, start_index=14621, title='KIAA1614 (RP11-46A10.3) Result Summary | BioGRID', type='url_citation', url='https://thebiogrid.org/121734/protein/homo-sapiens/kiaa1614.html#:~:text=Switch%20View%3A')
  29. AnnotationURLCitation(end_index=15057, start_index=14922, title='KIAA1614 KIAA1614 [Homo sapiens (human)] - Gene - NCBI', type='url_citation', url='https://www.ncbi.nlm.nih.gov/gene/57710#:~:text=The%20complete%20sequences%20of%20100,Science%2C%202001%20Feb')
  30. AnnotationURLCitation(end_index=15951, start_index=15834, title='KIAA1614 protein (Canis lupus familiaris) - STRING interaction network', type='url_citation', url='https://string-db.org/network/9615.ENSCAFP00000035844#:~:text=KIAA1614%20protein%20,Cleavage%20and')
  31. AnnotationURLCitation(end_index=16858, start_index=16720, title='CDD Conserved Protein Domain Family: PB1_Par6', type='url_citation', url='https://www.ncbi.nlm.nih.gov/Structure/cdd/cd06403#:~:text=Feature%201%3APB1%20interaction%20surface%20,Evidence')
  32. AnnotationURLCitation(end_index=17695, start_index=17505, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20FUNCTION%20Gene%20summary%20%28Entrez%29,Length%20%26%20mass%20Signal%20peptide')
  33. AnnotationURLCitation(end_index=19707, start_index=19534, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=regulation%20of%20circulating%20adhesion%20molecules.,might%20partly%20be%20mediated%20through')
  34. AnnotationURLCitation(end_index=20130, start_index=19957, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=regulation%20of%20circulating%20adhesion%20molecules.,might%20partly%20be%20mediated%20through')
  35. AnnotationURLCitation(end_index=20885, start_index=20801, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=%7B,t')
  36. AnnotationURLCitation(end_index=22111, start_index=22017, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=%5B%7B,children')
  37. AnnotationURLCitation(end_index=22340, start_index=22246, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=%5B%7B,children')
  38. AnnotationURLCitation(end_index=23632, start_index=23505, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=%7B,Blood%20group%20antigen%20loci%20demonstrate')
  39. AnnotationURLCitation(end_index=24842, start_index=24707, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=,transcriptionally%20silenced%2C%20implying%20that%20its')
  40. AnnotationURLCitation(end_index=25284, start_index=25149, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=,transcriptionally%20silenced%2C%20implying%20that%20its')
  41. AnnotationURLCitation(end_index=26256, start_index=26085, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=Detected%20in%20many%20Cell%20line,IMMUNOHISTOCHEMISTRY%20DATA%20RELIABILITY')
  42. AnnotationURLCitation(end_index=26508, start_index=26343, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=PROTEIN%20EXPRESSION%20,i%7D%20OrganExpressionAlphabetical%20HPA028227')
  43. AnnotationURLCitation(end_index=26847, start_index=26722, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=PROGNOSTIC%20SUMMARY,favorable')
  44. AnnotationURLCitation(end_index=27087, start_index=26962, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=PROGNOSTIC%20SUMMARY,favorable')
  45. AnnotationURLCitation(end_index=27318, start_index=27193, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=PROGNOSTIC%20SUMMARY,favorable')
  46. AnnotationURLCitation(end_index=27724, start_index=27599, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=PROGNOSTIC%20SUMMARY,favorable')
  47. AnnotationURLCitation(end_index=28101, start_index=27976, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=PROGNOSTIC%20SUMMARY,favorable')
  48. AnnotationURLCitation(end_index=30334, start_index=30172, title='Potential functional variants of KIAA genes are associated with breast cancer risk in a case control study - PMC', type='url_citation', url='https://pmc.ncbi.nlm.nih.gov/articles/PMC8105804/#:~:text=KIAA%20genes%20identified%20in%20the,of%20several%20human%20complex%20diseases')
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  51. AnnotationURLCitation(end_index=32008, start_index=31873, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=,transcriptionally%20silenced%2C%20implying%20that%20its')
  52. AnnotationURLCitation(end_index=33294, start_index=33121, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=regulation%20of%20circulating%20adhesion%20molecules.,might%20partly%20be%20mediated%20through')
  53. AnnotationURLCitation(end_index=34729, start_index=34539, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20FUNCTION%20Gene%20summary%20%28Entrez%29,Length%20%26%20mass%20Signal%20peptide')
  54. AnnotationURLCitation(end_index=35022, start_index=34860, title='Potential functional variants of KIAA genes are associated with breast cancer risk in a case control study - PMC', type='url_citation', url='https://pmc.ncbi.nlm.nih.gov/articles/PMC8105804/#:~:text=KIAA%20genes%20identified%20in%20the,of%20several%20human%20complex%20diseases')
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  56. AnnotationURLCitation(end_index=36149, start_index=35959, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=PROTEIN%20FUNCTION%20Gene%20summary%20%28Entrez%29,Length%20%26%20mass%20Signal%20peptide')
  57. AnnotationURLCitation(end_index=36299, start_index=36150, title='KIAA1614 KIAA1614 [Homo sapiens (human)] - Gene - NCBI', type='url_citation', url='https://www.ncbi.nlm.nih.gov/gene/57710#:~:text=Predicted%20to%20be%20involved%20in,of%20Genome%20Resources%2C%20Jul%202025')
  58. AnnotationURLCitation(end_index=36677, start_index=36532, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=Protein%20expression%20normal%20tissue,i%7D%20%201')
  59. AnnotationURLCitation(end_index=36898, start_index=36745, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=RNA%20category,at%20protein%20level%20Protein%20expression')
  60. AnnotationURLCitation(end_index=37130, start_index=36963, title='KIAA1614 gene information - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/summary/gene#:~:text=Intracellular%20proteins%20predicted%20by%20MDM,centrosome%20cycle')
  61. AnnotationURLCitation(end_index=37330, start_index=37205, title='Expression of KIAA1614 in cancer - Summary - The Human Protein Atlas', type='url_citation', url='https://www.proteinatlas.org/ENSG00000135835-KIAA1614/cancer#:~:text=PROGNOSTIC%20SUMMARY,favorable')
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  64. AnnotationURLCitation(end_index=38820, start_index=38658, title='Potential functional variants of KIAA genes are associated with breast cancer risk in a case control study - PMC', type='url_citation', url='https://pmc.ncbi.nlm.nih.gov/articles/PMC8105804/#:~:text=KIAA%20genes%20identified%20in%20the,of%20several%20human%20complex%20diseases')
  65. AnnotationURLCitation(end_index=38963, start_index=38821, title='Potential functional variants of KIAA genes are associated with breast cancer risk in a case control study - PMC', type='url_citation', url='https://pmc.ncbi.nlm.nih.gov/articles/PMC8105804/#:~:text=.%202021%20Apr%3B9%287%29%3A549.%20doi%3A%2010.21037%2Fatm')
  66. AnnotationURLCitation(end_index=39531, start_index=39396, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=,transcriptionally%20silenced%2C%20implying%20that%20its')
  67. AnnotationURLCitation(end_index=39616, start_index=39532, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=,href')
  68. AnnotationURLCitation(end_index=40167, start_index=39994, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=regulation%20of%20circulating%20adhesion%20molecules.,might%20partly%20be%20mediated%20through')
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  70. AnnotationURLCitation(end_index=40731, start_index=40647, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=%7B,t')
  71. AnnotationURLCitation(end_index=40819, start_index=40732, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=,PMID%3A')
  72. AnnotationURLCitation(end_index=41271, start_index=41177, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=%5B%7B,children')
  73. AnnotationURLCitation(end_index=41359, start_index=41272, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=,PMID%3A')
  74. AnnotationURLCitation(end_index=41883, start_index=41756, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=%7B,Blood%20group%20antigen%20loci%20demonstrate')
  75. AnnotationURLCitation(end_index=41968, start_index=41884, title='Gene - KIAA1614', type='url_citation', url='https://maayanlab.cloud/Harmonizome/gene/KIAA1614#:~:text=,href')
  76. AnnotationURLCitation(end_index=42707, start_index=42572, title='KIAA1614 KIAA1614 [Homo sapiens (human)] - Gene - NCBI', type='url_citation', url='https://www.ncbi.nlm.nih.gov/gene/57710#:~:text=The%20complete%20sequences%20of%20100,Science%2C%202001%20Feb')
  77. AnnotationURLCitation(end_index=43337, start_index=43238, title='KIAA1614 KIAA1614 [Homo sapiens (human)] - Gene - NCBI', type='url_citation', url='https://www.ncbi.nlm.nih.gov/gene/57710#:~:text=3,Science%2C%202001%20Feb')
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  79. AnnotationURLCitation(end_index=44409, start_index=44292, title='KIAA1614 protein (Canis lupus familiaris) - STRING interaction network', type='url_citation', url='https://string-db.org/network/9615.ENSCAFP00000035844#:~:text=KIAA1614%20protein%20,Cleavage%20and')