alpha-Man-Ia

UniProt ID: P53624
Organism: Drosophila melanogaster
Review Status: IN PROGRESS
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Gene Description

Alpha-Man-Ia (mas-1) is a Golgi glycoside hydrolase of family GH47 that trims alpha-1,2-linked mannose residues during N-glycan maturation. Its N-terminal membrane anchor presents the catalytic domain to the secretory-pathway lumen. Loss of the enzyme changes glycan proportions and causes Man8GlcNAc2 accumulation, while alternative processing routes retain substantial glycosylation.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0000139 Golgi membrane
IBA
GO_REF:0000033
ACCEPT
Summary: Golgi residence fits the type-II glycan-processing enzyme.
Reason: The mas-1 mutant glycan study identifies the product as Golgi mannosidase I; its N-terminal signal anchor and lumen-facing GH47 catalytic region support Golgi membrane association. The direct biochemical work independently establishes glycan trimming.
Supporting Evidence:
PMID:9654102
The mas-1 gene of Drosophila melanogaster encodes Golgi mannosidase I (MAS-1)
GO:0000139 Golgi membrane
IEA
GO_REF:0000044
ACCEPT
Summary: Golgi residence fits the type-II glycan-processing enzyme.
Reason: The mas-1 mutant glycan study identifies the product as Golgi mannosidase I; its N-terminal signal anchor and lumen-facing GH47 catalytic region support Golgi membrane association. The direct biochemical work independently establishes glycan trimming.
Supporting Evidence:
PMID:9654102
The mas-1 gene of Drosophila melanogaster encodes Golgi mannosidase I (MAS-1)
GO:0000139 Golgi membrane
ISS
GO_REF:0000024
ACCEPT
Summary: Golgi residence fits the type-II glycan-processing enzyme.
Reason: The mas-1 mutant glycan study identifies the product as Golgi mannosidase I; its N-terminal signal anchor and lumen-facing GH47 catalytic region support Golgi membrane association. The direct biochemical work independently establishes glycan trimming.
Supporting Evidence:
PMID:9654102
The mas-1 gene of Drosophila melanogaster encodes Golgi mannosidase I (MAS-1)
GO:0004571 mannosyl-oligosaccharide 1,2-alpha-mannosidase activity
IBA
GO_REF:0000033
ACCEPT
Summary: The enzyme removes alpha-1,2-linked mannose from oligomannosidic N-glycans.
Reason: Recombinant mas-1 is active toward oligomannosidic glycans, and mas-1 deletion causes Man8GlcNAc2 accumulation. Together these support the catalytic annotation on alpha-Man-Ia rather than transfer solely from a GH47 family label.
Supporting Evidence:
PMID:23979800
Some of the recombinant mannosidases were demonstrably active towards oligomannosidic glycans, specifically, the Co(II)-requiring ManIIb, two 'acidic' mannosidases and the class I mas-1 mannosidase.
PMID:9654102
There is an accumulation of the Man8GlcNAc2 which is one of the substrates for the MAS-1 enzyme.
GO:0004571 mannosyl-oligosaccharide 1,2-alpha-mannosidase activity
IDA
PMID:23979800
Characterisation of class I and II Ξ±-mannosidases from Droso...
ACCEPT
Summary: The enzyme removes alpha-1,2-linked mannose from oligomannosidic N-glycans.
Reason: Recombinant mas-1 is active toward oligomannosidic glycans, and mas-1 deletion causes Man8GlcNAc2 accumulation. Together these support the catalytic annotation on alpha-Man-Ia rather than transfer solely from a GH47 family label.
Supporting Evidence:
PMID:23979800
Some of the recombinant mannosidases were demonstrably active towards oligomannosidic glycans, specifically, the Co(II)-requiring ManIIb, two 'acidic' mannosidases and the class I mas-1 mannosidase.
PMID:9654102
There is an accumulation of the Man8GlcNAc2 which is one of the substrates for the MAS-1 enzyme.
GO:0004571 mannosyl-oligosaccharide 1,2-alpha-mannosidase activity
IEA
GO_REF:0000120
ACCEPT
Summary: The enzyme removes alpha-1,2-linked mannose from oligomannosidic N-glycans.
Reason: Recombinant mas-1 is active toward oligomannosidic glycans, and mas-1 deletion causes Man8GlcNAc2 accumulation. Together these support the catalytic annotation on alpha-Man-Ia rather than transfer solely from a GH47 family label.
Supporting Evidence:
PMID:23979800
Some of the recombinant mannosidases were demonstrably active towards oligomannosidic glycans, specifically, the Co(II)-requiring ManIIb, two 'acidic' mannosidases and the class I mas-1 mannosidase.
PMID:9654102
There is an accumulation of the Man8GlcNAc2 which is one of the substrates for the MAS-1 enzyme.
GO:0004571 mannosyl-oligosaccharide 1,2-alpha-mannosidase activity
IMP
PMID:9654102
Mutant analysis reveals an alternative pathway for N-linked ...
ACCEPT
Summary: The enzyme removes alpha-1,2-linked mannose from oligomannosidic N-glycans.
Reason: Recombinant mas-1 is active toward oligomannosidic glycans, and mas-1 deletion causes Man8GlcNAc2 accumulation. Together these support the catalytic annotation on alpha-Man-Ia rather than transfer solely from a GH47 family label.
Supporting Evidence:
PMID:23979800
Some of the recombinant mannosidases were demonstrably active towards oligomannosidic glycans, specifically, the Co(II)-requiring ManIIb, two 'acidic' mannosidases and the class I mas-1 mannosidase.
PMID:9654102
There is an accumulation of the Man8GlcNAc2 which is one of the substrates for the MAS-1 enzyme.
GO:0004571 mannosyl-oligosaccharide 1,2-alpha-mannosidase activity
ISS
GO_REF:0000024
ACCEPT
Summary: The enzyme removes alpha-1,2-linked mannose from oligomannosidic N-glycans.
Reason: Recombinant mas-1 is active toward oligomannosidic glycans, and mas-1 deletion causes Man8GlcNAc2 accumulation. Together these support the catalytic annotation on alpha-Man-Ia rather than transfer solely from a GH47 family label.
Supporting Evidence:
PMID:23979800
Some of the recombinant mannosidases were demonstrably active towards oligomannosidic glycans, specifically, the Co(II)-requiring ManIIb, two 'acidic' mannosidases and the class I mas-1 mannosidase.
PMID:9654102
There is an accumulation of the Man8GlcNAc2 which is one of the substrates for the MAS-1 enzyme.
GO:0005509 calcium ion binding
IEA
GO_REF:0000002
KEEP AS NON CORE
Summary: Calcium binding is a cofactor feature of the GH47 catalytic apparatus.
Reason: The calcium-dependent GH47 mechanism makes this binding annotation reasonable; it is ancillary to glycan hydrolysis rather than a distinct calcium-signaling function.
Supporting Evidence:
PMID:23979800
Some of the recombinant mannosidases were demonstrably active towards oligomannosidic glycans, specifically, the Co(II)-requiring ManIIb, two 'acidic' mannosidases and the class I mas-1 mannosidase.
GO:0005783 endoplasmic reticulum
IBA
GO_REF:0000033
UNDECIDED
Summary: ER residence requires distinction from the established Golgi role.
Reason: The experimentally supported role is Golgi N-glycan trimming. ER entry during biosynthesis does not establish persistent ER residence; the inherited ER annotation may reflect another GH47 branch, but the exact PAINT node and target trafficking evidence do not resolve that distinction here.
Supporting Evidence:
PMID:9654102
The mas-1 gene of Drosophila melanogaster encodes Golgi mannosidase I (MAS-1)
GO:0005975 carbohydrate metabolic process
IEA
GO_REF:0000002
MODIFY
Summary: N-glycan mannose trimming specifies the broad carbohydrate process.
Reason: Man8GlcNAc2 accumulation in the mutant identifies a direct glycoprotein-processing role rather than undifferentiated carbohydrate metabolism.
Supporting Evidence:
PMID:9654102
There is an accumulation of the Man8GlcNAc2 which is one of the substrates for the MAS-1 enzyme.
GO:0012505 endomembrane system
HDA
PMID:19317464
Mapping organelle proteins and protein complexes in Drosophi...
KEEP AS NON CORE
Summary: The endomembrane-system assignment is consistent but broad.
Reason: The organellar proteomics annotation is compatible with the independently established Golgi glycan-processing role. It does not resolve a vesicle lumen versus a membrane anchor.
Supporting Evidence:
PMID:9654102
The mas-1 gene of Drosophila melanogaster encodes Golgi mannosidase I (MAS-1)
GO:0016020 membrane
IEA
GO_REF:0000002
KEEP AS NON CORE
Summary: Broad membrane membership is supported by the signal anchor.
Reason: The type-II Golgi mannosidase has a membrane anchor; the specific Golgi membrane term better expresses its core location.
Supporting Evidence:
PMID:9654102
The mas-1 gene of Drosophila melanogaster encodes Golgi mannosidase I (MAS-1)
GO:0036503 ERAD quality control pathway
IBA
GO_REF:0000033
UNDECIDED
Summary: A dedicated ERAD role is not established for the Golgi-trimming enzyme.
Reason: GH47 includes ER quality-control mannosidases as well as Golgi-processing enzymes. The target mutant and recombinant assays demonstrate N-glycan trimming but do not directly show degradation commitment or ERAD participation. This is a branch-specific functional uncertainty, not evidence that all GH47 proteins lack ERAD roles.
Supporting Evidence:
PMID:23979800
Some of the recombinant mannosidases were demonstrably active towards oligomannosidic glycans, specifically, the Co(II)-requiring ManIIb, two 'acidic' mannosidases and the class I mas-1 mannosidase.
PMID:9654102
There is an accumulation of the Man8GlcNAc2 which is one of the substrates for the MAS-1 enzyme.
GO:1904381 Golgi apparatus N-glycan mannose trimming
IMP
PMID:9654102
Mutant analysis reveals an alternative pathway for N-linked ...
ACCEPT
Summary: Golgi N-glycan mannose trimming is the direct biological process.
Reason: The mutant glycan profile accumulates an enzyme substrate while retaining compensatory processing routes, establishing a normal contribution to mannose trimming without requiring lethality.
Supporting Evidence:
PMID:9654102
There is an accumulation of the Man8GlcNAc2 which is one of the substrates for the MAS-1 enzyme.

Core Functions

Trims mannose from N-linked glycans in the Golgi.

Supporting Evidence:
  • PMID:23979800
    Some of the recombinant mannosidases were demonstrably active towards oligomannosidic glycans, specifically, the Co(II)-requiring ManIIb, two 'acidic' mannosidases and the class I mas-1 mannosidase.
  • PMID:9654102
    There is an accumulation of the Man8GlcNAc2 which is one of the substrates for the MAS-1 enzyme.

References

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Deep Research

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