sns

UniProt ID: Q0E9F2
Organism: Drosophila melanogaster
Review Status: IN PROGRESS
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Gene Description

Sticks and stones (Sns) is a nephrin-related immunoglobulin-superfamily adhesion protein. It recognizes Kirre/Roughest on apposed cells, enabling myoblast fusion and organizing nephrocyte filtration diaphragms. Its extracellular Ig and fibronectin-type domains and intracellular signaling region couple cell recognition to tissue morphogenesis. Sns also contributes to eye organization, sensory-organ spacing and reproductive support-cell functions.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0001745 compound eye morphogenesis
IMP
PMID:20599904
Preferential adhesion maintains separation of ommatidia in t...
KEEP AS NON CORE
Summary: Sns participates in the differential adhesion and cellular arrangement of the developing compound eye. Retain this tissue-specific developmental association as noncore relative to its cell-adhesion mechanism; the evidence does not establish an independent photoreceptor fate-specification activity.
Reason: Sns participates in the differential adhesion and cellular arrangement of the developing compound eye. Retain this tissue-specific developmental association as noncore relative to its cell-adhesion mechanism; the evidence does not establish an independent photoreceptor fate-specification activity.
Supporting Evidence:
PMID:20599904
These results strongly suggest expression of Hbs and Sns is sufficient to render ommatidia less adhesive to other ommatidia than to IOCs.
GO:0001751 compound eye photoreceptor cell differentiation
NAS
PMID:20599904
Preferential adhesion maintains separation of ommatidia in t...
KEEP AS NON CORE
Summary: Sns participates in the differential adhesion and cellular arrangement of the developing compound eye. Retain this tissue-specific developmental association as noncore relative to its cell-adhesion mechanism; the evidence does not establish an independent photoreceptor fate-specification activity.
Reason: Sns participates in the differential adhesion and cellular arrangement of the developing compound eye. Retain this tissue-specific developmental association as noncore relative to its cell-adhesion mechanism; the evidence does not establish an independent photoreceptor fate-specification activity.
Supporting Evidence:
PMID:20599904
These results strongly suggest expression of Hbs and Sns is sufficient to render ommatidia less adhesive to other ommatidia than to IOCs.
GO:0005515 protein binding
IPI
PMID:18971929
The insect nephrocyte is a podocyte-like cell with a filtrat...
KEEP AS NON CORE
Summary: The primary study demonstrates binding of the Sns intracellular region to Mec2/podocin in the nephrocyte diaphragm complex. Preserve the experimentally supported interaction as noncore; generic protein binding does not describe the principal extracellular adhesion activity.
Reason: The primary study demonstrates binding of the Sns intracellular region to Mec2/podocin in the nephrocyte diaphragm complex. Preserve the experimentally supported interaction as noncore; generic protein binding does not describe the principal extracellular adhesion activity.
Supporting Evidence:
PMID:18971929
Sns interacts with Mec-2 (podocin) and Duf interacts with Pyd (ZO-1) (Fig. 4j).
GO:0005515 protein binding
IPI
PMID:23827685
An extracellular interactome of immunoglobulin and LRR prote...
MODIFY
Summary: Sns binds the extracellular adhesion protein Roughest; the interaction is part of the conserved nephrin/NEPH adhesion system. Cell adhesion molecule binding is more informative than generic protein binding, and the Drosophila heterocomplex is directly measured.
Reason: Sns binds the extracellular adhesion protein Roughest; the interaction is part of the conserved nephrin/NEPH adhesion system. Cell adhesion molecule binding is more informative than generic protein binding, and the Drosophila heterocomplex is directly measured.
Proposed replacements: cell adhesion molecule binding
Supporting Evidence:
PMID:24485456
theDrosophila homologs of SYG-1(Rst and Duf/Kirre) and of SYG-2(SNS and Hbs)all form hetero-complexes withaffinities between 1 to 4 ΞΌM
GO:0005515 protein binding
IPI
PMID:23827685
An extracellular interactome of immunoglobulin and LRR prote...
KEEP AS NON CORE
Summary: The extracellular interactome records an interaction with Unc-5. Retain the binding observation as noncore rather than infer a particular guidance pathway or ligand activity from this screen alone.
Reason: The extracellular interactome records an interaction with Unc-5. Retain the binding observation as noncore rather than infer a particular guidance pathway or ligand activity from this screen alone.
Supporting Evidence:
PMID:23827685
Our study reports the complete interactome of extracellular IgSF, FnIII and LRR protein families
GO:0005515 protein binding
IPI
PMID:24485456
Extracellular architecture of the SYG-1/SYG-2 adhesion compl...
MODIFY
Summary: Sns binds the extracellular adhesion protein Roughest; the interaction is part of the conserved nephrin/NEPH adhesion system. Cell adhesion molecule binding is more informative than generic protein binding, and the Drosophila heterocomplex is directly measured.
Reason: Sns binds the extracellular adhesion protein Roughest; the interaction is part of the conserved nephrin/NEPH adhesion system. Cell adhesion molecule binding is more informative than generic protein binding, and the Drosophila heterocomplex is directly measured.
Proposed replacements: cell adhesion molecule binding
Supporting Evidence:
PMID:24485456
theDrosophila homologs of SYG-1(Rst and Duf/Kirre) and of SYG-2(SNS and Hbs)all form hetero-complexes withaffinities between 1 to 4 ΞΌM
GO:0005886 plasma membrane
IBA
GO_REF:0000033
ACCEPT
Summary: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Reason: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Supporting Evidence:
PMID:10859168
SNS is enriched at the membrane and clusters at discrete sites, coincident with the occurrence of myoblast fusion.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0005886 plasma membrane
IDA
PMID:10859168
Drosophila SNS, a member of the immunoglobulin superfamily t...
ACCEPT
Summary: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Reason: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Supporting Evidence:
PMID:10859168
SNS is enriched at the membrane and clusters at discrete sites, coincident with the occurrence of myoblast fusion.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0005886 plasma membrane
IDA
PMID:18971929
The insect nephrocyte is a podocyte-like cell with a filtrat...
ACCEPT
Summary: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Reason: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Supporting Evidence:
PMID:10859168
SNS is enriched at the membrane and clusters at discrete sites, coincident with the occurrence of myoblast fusion.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0005911 cell-cell junction
IBA
GO_REF:0000033
ACCEPT
Summary: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Reason: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Supporting Evidence:
PMID:10859168
SNS is enriched at the membrane and clusters at discrete sites, coincident with the occurrence of myoblast fusion.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0005917 nephrocyte diaphragm
IDA
PMID:18971929
The insect nephrocyte is a podocyte-like cell with a filtrat...
ACCEPT
Summary: Sns is a structural cell-adhesion component of the nephrocyte diaphragm. Direct localization and loss-of-function studies support its presence and role at this filtration junction.
Reason: Sns is a structural cell-adhesion component of the nephrocyte diaphragm. Direct localization and loss-of-function studies support its presence and role at this filtration junction.
Supporting Evidence:
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0005917 nephrocyte diaphragm
IDA
PMID:19515699
Sns and Kirre, the Drosophila orthologs of Nephrin and Neph1...
ACCEPT
Summary: Sns is a structural cell-adhesion component of the nephrocyte diaphragm. Direct localization and loss-of-function studies support its presence and role at this filtration junction.
Reason: Sns is a structural cell-adhesion component of the nephrocyte diaphragm. Direct localization and loss-of-function studies support its presence and role at this filtration junction.
Supporting Evidence:
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0005917 nephrocyte diaphragm
IDA
PMID:38878170
Phospholipid scramblase 1: an essential component of the nep...
ACCEPT
Summary: Sns is a structural cell-adhesion component of the nephrocyte diaphragm. Direct localization and loss-of-function studies support its presence and role at this filtration junction.
Reason: Sns is a structural cell-adhesion component of the nephrocyte diaphragm. Direct localization and loss-of-function studies support its presence and role at this filtration junction.
Supporting Evidence:
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0007156 homophilic cell-cell adhesion
IGI
PMID:19515699
Sns and Kirre, the Drosophila orthologs of Nephrin and Neph1...
UNDECIDED
Summary: The accessible primary abstract establishes Sns-dependent adhesion between garland nephrocytes, but does not resolve whether the required molecular interaction is homophilic rather than Sns–Kirre heterophilic binding. Preserve the experimental IGI assertion unresolved until the full mechanistic evidence can be checked.
Reason: The accessible primary abstract establishes Sns-dependent adhesion between garland nephrocytes, but does not resolve whether the required molecular interaction is homophilic rather than Sns–Kirre heterophilic binding. Preserve the experimental IGI assertion unresolved until the full mechanistic evidence can be checked.
Supporting Evidence:
PMID:19515699
they contribute to adhesion of GCNs in the ;garland' and to their fusion into binucleate cells.
GO:0007157 heterophilic cell-cell adhesion
IDA
PMID:19270174
The immunoglobulin superfamily member Hbs functions redundan...
ACCEPT
Summary: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Reason: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Supporting Evidence:
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0007157 heterophilic cell-cell adhesion
IMP
PMID:15955848
A positive feedback loop between Dumbfounded and Rolling peb...
ACCEPT
Summary: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Reason: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Supporting Evidence:
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0007286 spermatid development
IMP
PMID:21652648
The WASp-based actin polymerization machinery is required in...
KEEP AS NON CORE
Summary: Sns participates in the somatic support-cell machinery required for spermatid maturation/release. Retain this developmental role without implying a sperm-intrinsic enzymatic function.
Reason: Sns participates in the somatic support-cell machinery required for spermatid maturation/release. Retain this developmental role without implying a sperm-intrinsic enzymatic function.
Supporting Evidence:
PMID:21652648
Involvement of the cell surface protein Sticks and stones and the Wsp adapter protein D-WIP (Vrp1) is also crucial.
GO:0007520 myoblast fusion
IMP
PMID:15955848
A positive feedback loop between Dumbfounded and Rolling peb...
ACCEPT
Summary: Sns is required on fusion-competent myoblasts for their adhesion to founder cells and subsequent muscle-fiber formation. The defining unfused-myoblast phenotype and receptor-ligand mechanism independently establish the gene-level myoblast-fusion role.
Reason: Sns is required on fusion-competent myoblasts for their adhesion to founder cells and subsequent muscle-fiber formation. The defining unfused-myoblast phenotype and receptor-ligand mechanism independently establish the gene-level myoblast-fusion role.
Supporting Evidence:
PMID:10859168
The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
GO:0007520 myoblast fusion
IMP
PMID:18003739
SCAR/WAVE and Arp2/3 are crucial for cytoskeletal remodeling...
ACCEPT
Summary: Sns is required on fusion-competent myoblasts for their adhesion to founder cells and subsequent muscle-fiber formation. The defining unfused-myoblast phenotype and receptor-ligand mechanism independently establish the gene-level myoblast-fusion role.
Reason: Sns is required on fusion-competent myoblasts for their adhesion to founder cells and subsequent muscle-fiber formation. The defining unfused-myoblast phenotype and receptor-ligand mechanism independently establish the gene-level myoblast-fusion role.
Supporting Evidence:
PMID:10859168
The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
GO:0007520 myoblast fusion
IMP
PMID:19270174
The immunoglobulin superfamily member Hbs functions redundan...
ACCEPT
Summary: Sns is required on fusion-competent myoblasts for their adhesion to founder cells and subsequent muscle-fiber formation. The defining unfused-myoblast phenotype and receptor-ligand mechanism independently establish the gene-level myoblast-fusion role.
Reason: Sns is required on fusion-competent myoblasts for their adhesion to founder cells and subsequent muscle-fiber formation. The defining unfused-myoblast phenotype and receptor-ligand mechanism independently establish the gene-level myoblast-fusion role.
Supporting Evidence:
PMID:10859168
The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
GO:0007523 larval visceral muscle development
IMP
PMID:11744371
The formation of syncytia within the visceral musculature of...
KEEP AS NON CORE
Summary: Mutations in sns disrupt fusion-dependent visceral muscle development. This is a tissue-specific consequence of its conserved adhesive myoblast-fusion mechanism.
Reason: Mutations in sns disrupt fusion-dependent visceral muscle development. This is a tissue-specific consequence of its conserved adhesive myoblast-fusion mechanism.
Supporting Evidence:
PMID:11744371
Mutations of genes that are involved in somatic myoblast fusion, such as sns, dumbfounded (duf) or myoblast city (mbc), also cause severe defects within the visceral musculature.
GO:0008407 chaeta morphogenesis
IGI
PMID:26053791
The Cell Adhesion Molecules Roughest, Hibris, Kin of Irre an...
KEEP AS NON CORE
Summary: Sns contributes to positioning of sensory-bristle precursors through tissue adhesion. This specific morphogenesis role is retained as noncore relative to its underlying cell-recognition mechanism.
Reason: Sns contributes to positioning of sensory-bristle precursors through tissue adhesion. This specific morphogenesis role is retained as noncore relative to its underlying cell-recognition mechanism.
Supporting Evidence:
PMID:26053791
In this paper we explore the roles of the IRM-proteins Rst, Hbs, Kirre and SNS in the spacing of bristles of the anterior wing margin
GO:0009653 anatomical structure morphogenesis
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: The broad developmental annotation is consistent with gene-specific muscle-fusion, eye-patterning and nephrocyte-development evidence. These are correct umbrella descriptions but less informative than the specific adhesion and junction-assembly functions; automated provenance itself is not their biological support.
Reason: The broad developmental annotation is consistent with gene-specific muscle-fusion, eye-patterning and nephrocyte-development evidence. These are correct umbrella descriptions but less informative than the specific adhesion and junction-assembly functions; automated provenance itself is not their biological support.
Supporting Evidence:
PMID:10859168
The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
PMID:20599904
These results strongly suggest expression of Hbs and Sns is sufficient to render ommatidia less adhesive to other ommatidia than to IOCs.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0016020 membrane
IEA
GO_REF:0000044
ACCEPT
Summary: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Reason: Sns is a cell-surface adhesion molecule enriched at myoblast contacts and nephrocyte junctions. Primary localization supports the membrane/junction assignment independently of automated location propagation.
Supporting Evidence:
PMID:10859168
SNS is enriched at the membrane and clusters at discrete sites, coincident with the occurrence of myoblast fusion.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0016202 regulation of striated muscle tissue development
NAS
PMID:20186342
The intracellular domain of Dumbfounded affects myoblast fus...
KEEP AS NON CORE
Summary: The gene participates in formation of striated muscle through myoblast recognition and fusion. Retain this broad regulatory developmental description as noncore; the more specific fusion annotations capture the direct role.
Reason: The gene participates in formation of striated muscle through myoblast recognition and fusion. Retain this broad regulatory developmental description as noncore; the more specific fusion annotations capture the direct role.
Supporting Evidence:
PMID:10859168
The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
GO:0016324 apical plasma membrane
IDA
PMID:26053791
The Cell Adhesion Molecules Roughest, Hibris, Kin of Irre an...
ACCEPT
Summary: Sns is localized in the apical contact region of sensory-organ precursor cells in the wing-disc analysis. This is consistent with its membrane-adhesion role, without implying universal apical localization in every tissue.
Reason: Sns is localized in the apical contact region of sensory-organ precursor cells in the wing-disc analysis. This is consistent with its membrane-adhesion role, without implying universal apical localization in every tissue.
Supporting Evidence:
PMID:26053791
The IRM-proteins mediated contact between these cell types on the apical side in the adherens junction (Fig 1D–1E).
PMID:26053791
SNS was in contrast to the other three IRM-proteins only detected in the membranes surrounding the SOPs
GO:0030036 actin cytoskeleton organization
IMP
PMID:18003739
SCAR/WAVE and Arp2/3 are crucial for cytoskeletal remodeling...
UNDECIDED
Summary: The cited study addresses actin foci at myoblast-fusion sites, but its cached abstract does not resolve the Sns-specific actin-remodeling result. Sns-dependent fusion is firmly established; this particular downstream cytoskeletal assertion remains unresolved pending access to the full experimental analysis.
Reason: The cited study addresses actin foci at myoblast-fusion sites, but its cached abstract does not resolve the Sns-specific actin-remodeling result. Sns-dependent fusion is firmly established; this particular downstream cytoskeletal assertion remains unresolved pending access to the full experimental analysis.
Supporting Evidence:
PMID:10859168
The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
GO:0030154 cell differentiation
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: The broad developmental annotation is consistent with gene-specific muscle-fusion, eye-patterning and nephrocyte-development evidence. These are correct umbrella descriptions but less informative than the specific adhesion and junction-assembly functions; automated provenance itself is not their biological support.
Reason: The broad developmental annotation is consistent with gene-specific muscle-fusion, eye-patterning and nephrocyte-development evidence. These are correct umbrella descriptions but less informative than the specific adhesion and junction-assembly functions; automated provenance itself is not their biological support.
Supporting Evidence:
PMID:10859168
The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
PMID:20599904
These results strongly suggest expression of Hbs and Sns is sufficient to render ommatidia less adhesive to other ommatidia than to IOCs.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0036059 nephrocyte diaphragm assembly
IMP
PMID:18971929
The insect nephrocyte is a podocyte-like cell with a filtrat...
ACCEPT
Summary: Loss of Sns reduces or abolishes the nephrocyte diaphragm, demonstrating a direct structural adhesion contribution to assembly of the filtration junction.
Reason: Loss of Sns reduces or abolishes the nephrocyte diaphragm, demonstrating a direct structural adhesion contribution to assembly of the filtration junction.
Supporting Evidence:
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0036059 nephrocyte diaphragm assembly
IMP
PMID:19515699
Sns and Kirre, the Drosophila orthologs of Nephrin and Neph1...
ACCEPT
Summary: Loss of Sns reduces or abolishes the nephrocyte diaphragm, demonstrating a direct structural adhesion contribution to assembly of the filtration junction.
Reason: Loss of Sns reduces or abolishes the nephrocyte diaphragm, demonstrating a direct structural adhesion contribution to assembly of the filtration junction.
Supporting Evidence:
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0050839 cell adhesion molecule binding
IBA
GO_REF:0000033
ACCEPT
Summary: The binding of Sns to Kirre/Roughest is supported by cell-recognition experiments and direct measurements of Drosophila heterocomplexes. This is a specific core molecular activity.
Reason: The binding of Sns to Kirre/Roughest is supported by cell-recognition experiments and direct measurements of Drosophila heterocomplexes. This is a specific core molecular activity.
Supporting Evidence:
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
PMID:24485456
theDrosophila homologs of SYG-1(Rst and Duf/Kirre) and of SYG-2(SNS and Hbs)all form hetero-complexes withaffinities between 1 to 4 ΞΌM
GO:0061321 garland nephrocyte differentiation
IMP
PMID:19515699
Sns and Kirre, the Drosophila orthologs of Nephrin and Neph1...
KEEP AS NON CORE
Summary: Sns mediates adhesion and fusion of garland nephrocytes into binucleate cells and supports their diaphragm formation. Retain the corresponding differentiation phenotype as a context-specific developmental consequence.
Reason: Sns mediates adhesion and fusion of garland nephrocytes into binucleate cells and supports their diaphragm formation. Retain the corresponding differentiation phenotype as a context-specific developmental consequence.
Supporting Evidence:
PMID:19515699
they contribute to adhesion of GCNs in the ;garland' and to their fusion into binucleate cells.
GO:0097206 nephrocyte filtration
IMP
PMID:18971929
The insect nephrocyte is a podocyte-like cell with a filtrat...
ACCEPT
Summary: Sns is an essential structural component of the nephrocyte filtration diaphragm; its depletion disrupts the junction and abolishes protein uptake. It participates in filtration without acting as an active solute pump.
Reason: Sns is an essential structural component of the nephrocyte filtration diaphragm; its depletion disrupts the junction and abolishes protein uptake. It participates in filtration without acting as an active solute pump.
Supporting Evidence:
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
PMID:23291470
Knockdown of sns or duf completely abolished the accumulation of the fluorescent protein in pericardial nephrocytes.
GO:0097206 nephrocyte filtration
IMP
PMID:23291470
An in vivo functional analysis system for renal gene discove...
ACCEPT
Summary: Sns is an essential structural component of the nephrocyte filtration diaphragm; its depletion disrupts the junction and abolishes protein uptake. It participates in filtration without acting as an active solute pump.
Reason: Sns is an essential structural component of the nephrocyte filtration diaphragm; its depletion disrupts the junction and abolishes protein uptake. It participates in filtration without acting as an active solute pump.
Supporting Evidence:
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
PMID:23291470
Knockdown of sns or duf completely abolished the accumulation of the fluorescent protein in pericardial nephrocytes.
GO:0098609 cell-cell adhesion
IBA
GO_REF:0000033
ACCEPT
Summary: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Reason: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Supporting Evidence:
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0098609 cell-cell adhesion
NAS
PMID:19515699
Sns and Kirre, the Drosophila orthologs of Nephrin and Neph1...
ACCEPT
Summary: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Reason: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Supporting Evidence:
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0098631 cell adhesion mediator activity
IPI
PMID:19270174
The immunoglobulin superfamily member Hbs functions redundan...
ACCEPT
Summary: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Reason: Sns on fusion-competent myoblasts engages Kirre/Roughest on founder cells, directly supporting heterophilic recognition and adhesion. This adhesive mechanism also supports specialized nephrocyte contacts.
Supporting Evidence:
PMID:19270174
Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:0098636 protein complex involved in cell adhesion
IPI
PMID:23827685
An extracellular interactome of immunoglobulin and LRR prote...
ACCEPT
Summary: The extracellular Sns–Kirre/Roughest adhesion complex is physically supported by the interactome and independent heterocomplex affinity measurements. This provides a specific complex context for its cell-adhesion activity.
Reason: The extracellular Sns–Kirre/Roughest adhesion complex is physically supported by the interactome and independent heterocomplex affinity measurements. This provides a specific complex context for its cell-adhesion activity.
Supporting Evidence:
PMID:24485456
theDrosophila homologs of SYG-1(Rst and Duf/Kirre) and of SYG-2(SNS and Hbs)all form hetero-complexes withaffinities between 1 to 4 ΞΌM
GO:0160108 animal gross anatomical part developmental process
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: The broad developmental annotation is consistent with gene-specific muscle-fusion, eye-patterning and nephrocyte-development evidence. These are correct umbrella descriptions but less informative than the specific adhesion and junction-assembly functions; automated provenance itself is not their biological support.
Reason: The broad developmental annotation is consistent with gene-specific muscle-fusion, eye-patterning and nephrocyte-development evidence. These are correct umbrella descriptions but less informative than the specific adhesion and junction-assembly functions; automated provenance itself is not their biological support.
Supporting Evidence:
PMID:10859168
The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
PMID:20599904
These results strongly suggest expression of Hbs and Sns is sufficient to render ommatidia less adhesive to other ommatidia than to IOCs.
PMID:19515699
Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
GO:1901739 regulation of myoblast fusion
NAS
PMID:11684662
Characterization of Drosophila hibris, a gene related to hum...
ACCEPT
Summary: Sns has a well-established gene-level role in myoblast fusion, with its cytoplasmic domain required for precursor formation. The Hibris-focused reference alone is not used to infer the Sns mechanism; direct Sns experiments support the process.
Reason: Sns has a well-established gene-level role in myoblast fusion, with its cytoplasmic domain required for precursor formation. The Hibris-focused reference alone is not used to infer the Sns mechanism; direct Sns experiments support the process.
Supporting Evidence:
PMID:10859168
The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
PMID:19270174
either an Hbs or Sns cytodomain is essential for muscle precursor formation

Core Functions

Mediates heterophilic cell recognition and adhesion during myoblast fusion.

Directly Involved In:
Cellular Locations:
Supporting Evidence:
  • PMID:10859168
    The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts.
  • PMID:19270174
    Kin-of-IrreC (Kirre) and Roughest (Rst) functioning redundantly in the founder cell and Sticks-and-stones (Sns) serving as their ligand in the FCMs.

Provides the adhesive structural scaffold required for nephrocyte diaphragm assembly and filtration.

Supporting Evidence:
  • PMID:19515699
    Sns and Kirre proteins localize to the ND at the entry point into the labyrinthine channels and, like their vertebrate counterparts, are essential for its formation.
  • PMID:23291470
    Knockdown of sns or duf completely abolished the accumulation of the fluorescent protein in pericardial nephrocytes.

References

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External Prediction Reviews

These computational predictions are reviewed separately from the GOA annotation set used for this review. The assessments below are from this project and do not constitute official GO annotations or endorsement by GO/UniProt. They are not included in the existing annotation review above.

ProtNLM2 External predictions

View prediction review YAML Β· sns-protnlm-predictions-review.yaml Β· Review status: COMPLETE

Sns has established gene-level developmental roles, but the selected isoform B contains an extra 63-residue insertion with a predicted membrane-spanning segment. Gene-level Sns-dependent muscle and tissue morphogenesis is strongly supported. Membrane association is supported by the retained hydrophobic regions and direct Sns cell-surface localization.

Source documents: genes/DROME/sns/sns-predictions-source.json Β· genes/DROME/sns/sns-uniprot-source.json

Review score: 2 = concordant with evidence; 1 = uncertain; 0 = discordant with evidence. This is an assessment score, not a model probability.

GO:0030154 cell differentiation GO_BP
UNC β€” Uncertain Review score: 1/2
Prediction method: ProtNLM2 Β· Version: UniProt API snapshot 2026-09-08 Β· file:DROME/sns/sns-predictions-source.json
Review rationale: Sns has established gene-level developmental roles, but the selected isoform B contains an extra 63-residue insertion with a predicted membrane-spanning segment. All shared residues are conserved, yet an additional membrane crossing could change presentation of the signaling tail required in fusion and differentiation. The accessible experiments do not resolve this topology or establish function of this exact longer form. Thus the prediction remains uncertain for the selected sequence, without disputing Sns gene-level differentiation biology.
Supporting Evidence:
  • PMID:10859168: "The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts."
  • file:DROME/sns/sns-bioinformatics/RESULTS.md: "The sole difference is a 63-residue insertion at the reference 1059/1060 junction, corresponding to target 1060–1122."
  • file:DROME/sns/sns-bioinformatics/RESULTS.md: "The selected record predicts transmembrane segments 1089–1112 (within the insertion) and 1141–1166 (in the shared sequence)."
GO:0009653 anatomical structure morphogenesis GO_BP
UNC β€” Uncertain Review score: 1/2
Prediction method: ProtNLM2 Β· Version: UniProt API snapshot 2026-09-08 Β· file:DROME/sns/sns-predictions-source.json
Review rationale: Gene-level Sns-dependent muscle and tissue morphogenesis is strongly supported. The exact selected isoform has a specific extra hydrophobic insertion that could alter the membrane topology required to couple extracellular adhesion to intracellular signaling. Its consequence is untested, so the morphology claim is not refuted but remains uncertain for this sequence. Mere agreement with the existing broad automated annotation would not resolve that issue.
Supporting Evidence:
  • PMID:10859168: "The Drosophila sticks-and-stones (sns) locus was identified on the basis of its mutant phenotype, the complete absence of body wall muscles and corresponding presence of unfused myoblasts."
  • PMID:20599904: "These results strongly suggest expression of Hbs and Sns is sufficient to render ommatidia less adhesive to other ommatidia than to IOCs."
  • file:DROME/sns/sns-bioinformatics/RESULTS.md: "The sole difference is a 63-residue insertion at the reference 1059/1060 junction, corresponding to target 1060–1122."
  • file:DROME/sns/sns-bioinformatics/RESULTS.md: "The selected record predicts transmembrane segments 1089–1112 (within the insertion) and 1141–1166 (in the shared sequence)."
GO:0016020 membrane GO_CC
LSP β€” Less precise than existing annotation Review score: 2/2
Prediction method: ProtNLM2 Β· Version: UniProt API snapshot 2026-09-08 Β· file:DROME/sns/sns-predictions-source.json
Review rationale: Membrane association is supported by the retained hydrophobic regions and direct Sns cell-surface localization. The extra hydrophobic insertion raises signaling-topology questions but does not undermine membrane association itself. The prediction is broader than the existing plasma-membrane and nephrocyte-diaphragm annotations.
Supporting Evidence:
  • PMID:10859168: "SNS is enriched at the membrane and clusters at discrete sites, coincident with the occurrence of myoblast fusion."
  • file:DROME/sns/sns-bioinformatics/RESULTS.md: "The selected record predicts transmembrane segments 1089–1112 (within the insertion) and 1141–1166 (in the shared sequence)."

Deep Research

Falcon

(sns-deep-research-falcon.md)

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Manual

(sns-deep-research-manual.md)

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πŸ“š Additional Documentation

Notes

(sns-notes.md)

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Bioinformatics Results

(RESULTS.md)

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πŸ“„ View Raw YAML

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