NCU08595

UniProt ID: Q7SCN3
Organism: Neurospora crassa (strain ATCC 24698 / 74-OR23-1A / CBS 708.71 / DSM 1257 / FGSC 987)
Review Status: COMPLETE
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Gene Description

NCU08595 is a Rpf2-family ribosome assembly factor with a Brix RNA-binding domain. By conservation with characterized fungal Rpf2, it associates with Rrs1, binds 5S rRNA and promotes incorporation of the 5S ribonucleoprotein particle into nucleolar large-subunit precursors. This assembly step supports large-subunit rRNA maturation.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0000027 ribosomal large subunit assembly
IEA
GO_REF:0000002
ACCEPT
Summary: NCU08595 has the Rpf2-specific family assignment in addition to its Brix domain.
Reason: NCU08595 has the Rpf2-specific family assignment in addition to its Brix domain. Characterized fungal Rpf2-Rrs1 recruits 5S ribonucleoprotein components into pre-ribosomes and supports subsequent large-subunit rRNA maturation. This is conserved assembly-factor participation, not intrinsic rRNA cleavage.
Supporting Evidence:
PMID:25855814
Proteins Rpf2 and Rrs1 are required for 60S ribosomal subunit maturation.
PMID:25855814
gel shift analysis revealed that the Rpf2-Rrs1 complex binds directly to 5S rRNA.
GO:0000463 maturation of LSU-rRNA from tricistronic rRNA transcript (SSU-rRNA, 5.8S rRNA, LSU-rRNA)
IBA
GO_REF:0000033
ACCEPT
Summary: NCU08595 has the Rpf2-specific family assignment in addition to its Brix domain.
Reason: NCU08595 has the Rpf2-specific family assignment in addition to its Brix domain. Characterized fungal Rpf2-Rrs1 recruits 5S ribonucleoprotein components into pre-ribosomes and supports subsequent large-subunit rRNA maturation. This is conserved assembly-factor participation, not intrinsic rRNA cleavage.
Supporting Evidence:
PMID:25855814
Proteins Rpf2 and Rrs1 are required for 60S ribosomal subunit maturation.
PMID:25855814
gel shift analysis revealed that the Rpf2-Rrs1 complex binds directly to 5S rRNA.
GO:0000470 maturation of LSU-rRNA
IEA
GO_REF:0000002
ACCEPT
Summary: NCU08595 has the Rpf2-specific family assignment in addition to its Brix domain.
Reason: NCU08595 has the Rpf2-specific family assignment in addition to its Brix domain. Characterized fungal Rpf2-Rrs1 recruits 5S ribonucleoprotein components into pre-ribosomes and supports subsequent large-subunit rRNA maturation. This is conserved assembly-factor participation, not intrinsic rRNA cleavage.
Supporting Evidence:
PMID:25855814
Proteins Rpf2 and Rrs1 are required for 60S ribosomal subunit maturation.
PMID:25855814
gel shift analysis revealed that the Rpf2-Rrs1 complex binds directly to 5S rRNA.
GO:0005730 nucleolus
IBA
GO_REF:0000033
ACCEPT
Summary: Rpf2 assembles 5S ribonucleoprotein components into nuclear pre-ribosomes and supports early large-subunit maturation.
Reason: Rpf2 assembles 5S ribonucleoprotein components into nuclear pre-ribosomes and supports early large-subunit maturation. The Rpf2-specific family and curator-assessed ancestral localization justify nucleolar residence by conservation.
Supporting Evidence:
PMID:25855814
Proteins Rpf2 and Rrs1 are required for 60S ribosomal subunit maturation.
GO:0005730 nucleolus
IEA
GO_REF:0000120
ACCEPT
Summary: Rpf2 assembles 5S ribonucleoprotein components into nuclear pre-ribosomes and supports early large-subunit maturation.
Reason: Rpf2 assembles 5S ribonucleoprotein components into nuclear pre-ribosomes and supports early large-subunit maturation. The Rpf2-specific family and curator-assessed ancestral localization justify nucleolar residence by conservation.
Supporting Evidence:
PMID:25855814
Proteins Rpf2 and Rrs1 are required for 60S ribosomal subunit maturation.
GO:0006364 rRNA processing
IEA
GO_REF:0000002
MODIFY
Summary: The broad ribosome-biogenesis or RNA-processing assertion is supported but can be made precise: Rpf2 supports large-subunit rRNA maturation through recruitment and assembly of the 5S ribonucleoprotein particle..
Reason: The broad ribosome-biogenesis or RNA-processing assertion is supported but can be made precise: Rpf2 supports large-subunit rRNA maturation through recruitment and assembly of the 5S ribonucleoprotein particle.
Supporting Evidence:
PMID:25855814
Proteins Rpf2 and Rrs1 are required for 60S ribosomal subunit maturation.
PMID:25855814
gel shift analysis revealed that the Rpf2-Rrs1 complex binds directly to 5S rRNA.
GO:0019843 rRNA binding
IBA
GO_REF:0000033
ACCEPT
Summary: Gel-shift and mutational analyses of fungal Rpf2-Rrs1 establish direct 5S rRNA binding with a contribution from Rpf2.
Reason: Gel-shift and mutational analyses of fungal Rpf2-Rrs1 establish direct 5S rRNA binding with a contribution from Rpf2. The target Rpf2-specific domain supports transfer of this RNA-binding property beyond the generic Brix fold.
Supporting Evidence:
PMID:25855814
gel shift analysis revealed that the Rpf2-Rrs1 complex binds directly to 5S rRNA.
GO:0019843 rRNA binding
IEA
GO_REF:0000120
ACCEPT
Summary: Gel-shift and mutational analyses of fungal Rpf2-Rrs1 establish direct 5S rRNA binding with a contribution from Rpf2.
Reason: Gel-shift and mutational analyses of fungal Rpf2-Rrs1 establish direct 5S rRNA binding with a contribution from Rpf2. The target Rpf2-specific domain supports transfer of this RNA-binding property beyond the generic Brix fold.
Supporting Evidence:
PMID:25855814
gel shift analysis revealed that the Rpf2-Rrs1 complex binds directly to 5S rRNA.
GO:0042254 ribosome biogenesis
IEA
GO_REF:0000117
MODIFY
Summary: The broad ribosome-biogenesis or RNA-processing assertion is supported but can be made precise: Rpf2 supports large-subunit rRNA maturation through recruitment and assembly of the 5S ribonucleoprotein particle..
Reason: The broad ribosome-biogenesis or RNA-processing assertion is supported but can be made precise: Rpf2 supports large-subunit rRNA maturation through recruitment and assembly of the 5S ribonucleoprotein particle.
Supporting Evidence:
PMID:25855814
Proteins Rpf2 and Rrs1 are required for 60S ribosomal subunit maturation.
PMID:25855814
gel shift analysis revealed that the Rpf2-Rrs1 complex binds directly to 5S rRNA.

Core Functions

NCU08595 is a Rpf2-family ribosome assembly factor with a Brix RNA-binding domain. By conservation with characterized fungal Rpf2, it associates with Rrs1, binds 5S rRNA and promotes incorporation of the 5S ribonucleoprotein particle into nucleolar large-subunit precursors. This assembly step supports large-subunit rRNA maturation.

Supporting Evidence:
  • PMID:25855814
    Proteins Rpf2 and Rrs1 are required for 60S ribosomal subunit maturation.
  • PMID:25855814
    gel shift analysis revealed that the Rpf2-Rrs1 complex binds directly to 5S rRNA.
  • file:NEUCR/NCU08595/NCU08595-uniprot.txt
    DR InterPro; IPR039770; Rpf2.

References

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External Prediction Reviews

These computational predictions are reviewed separately from the GOA annotation set used for this review. The assessments below are from this project and do not constitute official GO annotations or endorsement by GO/UniProt. They are not included in the existing annotation review above.

ProtNLM2 External predictions

View prediction review YAML Β· NCU08595-protnlm-predictions-review.yaml Β· Review status: COMPLETE

NCU08595 is a Rpf2-family ribosome assembly factor with a Brix RNA-binding domain. By conservation with characterized fungal Rpf2, it associates with Rrs1, binds 5S rRNA and promotes incorporation of the 5S ribonucleoprotein particle into nucleolar large-subunit precursors. This assembly step supports large-subunit rRNA maturation.

Source documents: genes/NEUCR/NCU08595/NCU08595-protnlm-source.json Β· genes/NEUCR/NCU08595/NCU08595-uniprot.txt Β· genes/NEUCR/NCU08595/NCU08595-ai-review.yaml

Review score: 2 = concordant with evidence; 1 = uncertain; 0 = discordant with evidence. This is an assessment score, not a model probability.

GO:0000470 maturation of LSU-rRNA GO_BP
LSP β€” Less precise than existing annotation Review score: 2/2
Prediction method: ProtNLM2 Β· Version: UniProt API snapshot 2026-09-09 Β· file:NEUCR/NCU08595/NCU08595-protnlm-source.json
Review rationale: The Rpf2-specific family assignment, beyond the generic Brix domain, supports transfer from characterized fungal Rpf2-Rrs1 complexes. Structural and biochemical studies establish 5S rRNA recruitment and downstream large-subunit maturation. GOA already includes maturation of LSU-rRNA from the tricistronic precursor (GO:0000463), making this general LSU-rRNA maturation prediction less precise. Participation is through ribosome assembly and does not imply intrinsic nuclease chemistry.
Supporting Evidence:

Deep Research

Falcon

(NCU08595-deep-research-falcon.md)

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πŸ“š Additional Documentation

Notes

(NCU08595-notes.md)

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πŸ“„ View Raw YAML

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