rpo41

UniProt ID: O13993
Organism: Schizosaccharomyces pombe (strain 972 / ATCC 24843)
Review Status: COMPLETE
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Gene Description

Rpo41 is the mitochondrial DNA-directed RNA polymerase of Schizosaccharomyces pombe. It binds mitochondrial promoters and synthesizes RNA from DNA templates; association with the transcription factor Mtf1 greatly enhances promoter-directed initiation. Its transcriptional activity sustains mitochondrial gene expression.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0001018 mitochondrial promoter sequence-specific DNA binding
EXP
PMID:21357609
Identification and characterization of the mitochondrial RNA...
ACCEPT
Summary: mitochondrial promoter sequence-specific DNA binding is supported.
Reason: Purified fission-yeast Rpo41 binds mitochondrial promoters in electrophoretic mobility-shift assays, including activity without Mtf1; Mtf1 enhances productive promoter recognition.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0001018 mitochondrial promoter sequence-specific DNA binding
IBA
GO_REF:0000033
ACCEPT
Summary: mitochondrial promoter sequence-specific DNA binding is supported.
Reason: Purified fission-yeast Rpo41 binds mitochondrial promoters in electrophoretic mobility-shift assays, including activity without Mtf1; Mtf1 enhances productive promoter recognition.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0003677 DNA binding
IEA
GO_REF:0000002
MODIFY
Summary: DNA binding is reviewed in the context of rpo41.
Reason: The broad DNA binding annotation is compatible with the biology, but mitochondrial promoter sequence-specific DNA binding captures the supported rpo41 function more precisely.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0003899 DNA-directed RNA polymerase activity
EXP
PMID:21357609
Identification and characterization of the mitochondrial RNA...
ACCEPT
Summary: DNA-directed RNA polymerase activity is supported.
Reason: Reconstituted transcription with purified fission-yeast Rpo41 directly establishes DNA-directed RNA synthesis. This is an intrinsic catalytic polymerase, with Mtf1 enhancing initiation.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0003899 DNA-directed RNA polymerase activity
IBA
GO_REF:0000033
ACCEPT
Summary: DNA-directed RNA polymerase activity is supported.
Reason: Reconstituted transcription with purified fission-yeast Rpo41 directly establishes DNA-directed RNA synthesis. This is an intrinsic catalytic polymerase, with Mtf1 enhancing initiation.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0003899 DNA-directed RNA polymerase activity
IEA
GO_REF:0000120
ACCEPT
Summary: DNA-directed RNA polymerase activity is supported.
Reason: Reconstituted transcription with purified fission-yeast Rpo41 directly establishes DNA-directed RNA synthesis. This is an intrinsic catalytic polymerase, with Mtf1 enhancing initiation.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0005739 mitochondrion
IDA
PMID:21727087
A genome wide study in fission yeast reveals nine PPR protei...
ACCEPT
Summary: mitochondrion is supported.
Reason: The mitochondrial localization is consistent with the direct mitochondrial-promoter transcription experiments and the PPR localization study.
Supporting Evidence:
file:SCHPO/rpo41/rpo41-uniprot.txt
DR GO; GO:0005739; C:mitochondrion; IDA:PomBase.
GO:0005739 mitochondrion
IEA
GO_REF:0000044
ACCEPT
Summary: mitochondrion is supported.
Reason: The mitochondrial localization is consistent with the direct mitochondrial-promoter transcription experiments and the PPR localization study.
Supporting Evidence:
file:SCHPO/rpo41/rpo41-uniprot.txt
DR GO; GO:0005739; C:mitochondrion; IDA:PomBase.
GO:0005759 mitochondrial matrix
ISS
GO_REF:0000024
ACCEPT
Summary: mitochondrial matrix is supported.
Reason: Matrix residence is a curator-mediated similarity inference consistent with access to mitochondrial DNA and the conserved mitochondrial transcription machinery. No target-specific divergence is evident.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0006351 DNA-templated transcription
IEA
GO_REF:0000002
MODIFY
Summary: DNA-templated transcription is reviewed in the context of rpo41.
Reason: The broad DNA-templated transcription annotation is compatible with the biology, but mitochondrial transcription captures the supported rpo41 function more precisely.
Proposed replacements: mitochondrial transcription
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0006390 mitochondrial transcription
IBA
GO_REF:0000033
ACCEPT
Summary: mitochondrial transcription is supported.
Reason: Rpo41 depletion and overexpression change mitochondrial transcripts, and purified Rpo41 transcribes mitochondrial promoters (PMID:21357609).
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0006390 mitochondrial transcription
IEA
GO_REF:0000117
ACCEPT
Summary: mitochondrial transcription is supported.
Reason: Rpo41 depletion and overexpression change mitochondrial transcripts, and purified Rpo41 transcribes mitochondrial promoters (PMID:21357609).
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0006391 transcription initiation at mitochondrial promoter
IMP
PMID:21357609
Identification and characterization of the mitochondrial RNA...
ACCEPT
Summary: transcription initiation at mitochondrial promoter is supported.
Reason: The reconstituted system initiates at mitochondrial promoters and recapitulates the in-vivo start sites.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0034062 5'-3' RNA polymerase activity
IEA
GO_REF:0000116
ACCEPT
Summary: 5'-3' RNA polymerase activity is supported.
Reason: The direct DNA-templated RNA synthesis assay supports this broader directional RNA polymerization activity; it is compatible with the specific DNA-directed polymerase annotation.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0034245 mitochondrial DNA-directed RNA polymerase complex
IBA
GO_REF:0000033
ACCEPT
Summary: mitochondrial DNA-directed RNA polymerase complex is supported.
Reason: Mtf1 and Rpo41 form the functional mitochondrial transcription machinery; purified factors bind promoters more efficiently together and support transcription.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0034245 mitochondrial DNA-directed RNA polymerase complex
IDA
PMID:21357609
Identification and characterization of the mitochondrial RNA...
ACCEPT
Summary: mitochondrial DNA-directed RNA polymerase complex is supported.
Reason: Mtf1 and Rpo41 form the functional mitochondrial transcription machinery; purified factors bind promoters more efficiently together and support transcription.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.
GO:0034245 mitochondrial DNA-directed RNA polymerase complex
IEA
GO_REF:0000117
ACCEPT
Summary: mitochondrial DNA-directed RNA polymerase complex is supported.
Reason: Mtf1 and Rpo41 form the functional mitochondrial transcription machinery; purified factors bind promoters more efficiently together and support transcription.
Supporting Evidence:
PMID:21357609
We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.

Core Functions

Catalyzes mitochondrial promoter-directed RNA synthesis in cooperation with Mtf1.

Supporting Evidence:
  • PMID:21357609
    We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro.

References

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External Prediction Reviews

These computational predictions are reviewed separately from the GOA annotation set used for this review. The assessments below are from this project and do not constitute official GO annotations or endorsement by GO/UniProt. They are not included in the existing annotation review above.

ProtNLM2 External predictions

View prediction review YAML Β· rpo41-protnlm-predictions-review.yaml Β· Review status: COMPLETE

The broad GO predictions describe established biology; all are less specific than supported curated annotations (LSP).

Source documents: genes/SCHPO/rpo41/rpo41-protnlm-source.xml Β· genes/SCHPO/rpo41/rpo41-protnlm-source.json Β· genes/SCHPO/rpo41/rpo41-uniprot.txt

Review score: 2 = concordant with evidence; 1 = uncertain; 0 = discordant with evidence. This is an assessment score, not a model probability.

GO:0003677 DNA binding GO_MF
LSP β€” Less precise than existing annotation Review score: 2/2
Prediction method: ProtNLM2 Β· Version: pre-release post-processed-2026_02_28k.xml Β· file:SCHPO/rpo41/rpo41-protnlm-source.xml
Review rationale: Purified S. pombe Rpo41 binds mitochondrial promoters and supports transcription together with Mtf1 (PMID:21357609). The predicted DNA binding is biologically supported. The existing mitochondrial promoter sequence-specific DNA binding annotation is more specific within the same GO aspect, so the prediction is LSP.
Supporting Evidence:
  • PMID:21357609: "We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro."
GO:0006351 DNA-templated transcription GO_BP
LSP β€” Less precise than existing annotation Review score: 2/2
Prediction method: ProtNLM2 Β· Version: pre-release post-processed-2026_02_28k.xml Β· file:SCHPO/rpo41/rpo41-protnlm-source.xml
Review rationale: Purified S. pombe Rpo41 binds mitochondrial promoters and supports transcription together with Mtf1 (PMID:21357609). The predicted DNA-templated transcription is biologically supported. The existing mitochondrial transcription annotation is more specific within the same GO aspect, so the prediction is LSP.
Supporting Evidence:
  • PMID:21357609: "We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro."
GO:0000428 DNA-directed RNA polymerase complex GO_CC
LSP β€” Less precise than existing annotation Review score: 2/2
Prediction method: ProtNLM2 Β· Version: pre-release post-processed-2026_02_28k.xml Β· file:SCHPO/rpo41/rpo41-protnlm-source.xml
Review rationale: Purified S. pombe Rpo41 binds mitochondrial promoters and supports transcription together with Mtf1 (PMID:21357609). The predicted DNA-directed RNA polymerase complex is biologically supported. The existing mitochondrial DNA-directed RNA polymerase complex annotation is more specific within the same GO aspect, so the prediction is LSP.
Supporting Evidence:
  • PMID:21357609: "We show biochemically, that purified Mtf1 and Rpo41 together can bind to the S. pombe mitochondrial promoters and can support transcription from the S. pombe mitochondrial promoters in vitro."

Deep Research

Falcon

(rpo41-deep-research-falcon.md)

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πŸ“š Additional Documentation

Notes

(rpo41-notes.md)

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Protnlm Function Review

(rpo41-protnlm-function-review.md)

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πŸ“„ View Raw YAML

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