wss1

UniProt ID: Q9P7B5
Organism: Schizosaccharomyces pombe (strain 972 / ATCC 24843)
Review Status: COMPLETE
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Gene Description

Wss1 is a nuclear Wss1-like metalloprotease with a conserved WLM catalytic domain and a C-terminal SUMO-interaction motif. Its role in proteolytic repair of covalent DNA-protein cross-links is inferred from the characterized budding-yeast Wss1 family and curated phylogenetic evidence. Removal of the cross-linked protein allows downstream DNA repair or lesion bypass. Direct biochemical characterization of the fission-yeast protein remains limited.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0004222 metalloendopeptidase activity
ISO
GO_REF:0000024
ACCEPT
Summary: The target carries the diagnostic WLM domain and conserved metal-binding/catalytic residues.
Reason: The target carries the diagnostic WLM domain and conserved metal-binding/catalytic residues. Curated orthology and PAINT ancestry support inheritance of the Wss1 protease/DNA-repair function, whose mechanistic foundation is DNA-dependent proteolysis of cross-linked proteins in the characterized yeast homolog. This is a justified family inference, not target-specific experimental proof.
Supporting Evidence:
PMID:24998930
Notably, in vitro assays indicate that substrates such as topoisomerase 1 are processed by the metalloprotease directly and in a DNA-dependent manner.
file:SCHPO/wss1/wss1-uniprot.txt
DR Pfam; PF08325; WLM; 1.
GO:0005634 nucleus
HDA
PMID:16823372
ORFeome cloning and global analysis of protein localization ...
ACCEPT
Summary: Nuclear localization is supported by the reviewed record citing target localization measurements (PMID:16823372) and is consistent with conserved Wss1 action on DNA-protein cross-links.
Reason: Nuclear localization is supported by the reviewed record citing target localization measurements (PMID:16823372) and is consistent with conserved Wss1 action on DNA-protein cross-links. The IBA adds a curated ancestral localization inference.
Supporting Evidence:
file:SCHPO/wss1/wss1-uniprot.txt
CC -!- SUBCELLULAR LOCATION: Nucleus {ECO:0000269|PubMed:16823372}.
GO:0005634 nucleus
IBA
GO_REF:0000033
ACCEPT
Summary: Nuclear localization is supported by the reviewed record citing target localization measurements (PMID:16823372) and is consistent with conserved Wss1 action on DNA-protein cross-links.
Reason: Nuclear localization is supported by the reviewed record citing target localization measurements (PMID:16823372) and is consistent with conserved Wss1 action on DNA-protein cross-links. The IBA adds a curated ancestral localization inference.
Supporting Evidence:
file:SCHPO/wss1/wss1-uniprot.txt
CC -!- SUBCELLULAR LOCATION: Nucleus {ECO:0000269|PubMed:16823372}.
GO:0005634 nucleus
IEA
GO_REF:0000044
ACCEPT
Summary: Nuclear localization is supported by the reviewed record citing target localization measurements (PMID:16823372) and is consistent with conserved Wss1 action on DNA-protein cross-links.
Reason: Nuclear localization is supported by the reviewed record citing target localization measurements (PMID:16823372) and is consistent with conserved Wss1 action on DNA-protein cross-links. The IBA adds a curated ancestral localization inference.
Supporting Evidence:
file:SCHPO/wss1/wss1-uniprot.txt
CC -!- SUBCELLULAR LOCATION: Nucleus {ECO:0000269|PubMed:16823372}.
GO:0006281 DNA repair
IBA
GO_REF:0000033
ACCEPT
Summary: The target carries the diagnostic WLM domain and conserved metal-binding/catalytic residues.
Reason: The target carries the diagnostic WLM domain and conserved metal-binding/catalytic residues. Curated orthology and PAINT ancestry support inheritance of the Wss1 protease/DNA-repair function, whose mechanistic foundation is DNA-dependent proteolysis of cross-linked proteins in the characterized yeast homolog. This is a justified family inference, not target-specific experimental proof.
Supporting Evidence:
PMID:24998930
Notably, in vitro assays indicate that substrates such as topoisomerase 1 are processed by the metalloprotease directly and in a DNA-dependent manner.
file:SCHPO/wss1/wss1-uniprot.txt
DR Pfam; PF08325; WLM; 1.
GO:0008237 metallopeptidase activity
IBA
GO_REF:0000033
ACCEPT
Summary: The target carries the diagnostic WLM domain and conserved metal-binding/catalytic residues.
Reason: The target carries the diagnostic WLM domain and conserved metal-binding/catalytic residues. Curated orthology and PAINT ancestry support inheritance of the Wss1 protease/DNA-repair function, whose mechanistic foundation is DNA-dependent proteolysis of cross-linked proteins in the characterized yeast homolog. This is a justified family inference, not target-specific experimental proof.
Supporting Evidence:
PMID:24998930
Notably, in vitro assays indicate that substrates such as topoisomerase 1 are processed by the metalloprotease directly and in a DNA-dependent manner.
file:SCHPO/wss1/wss1-uniprot.txt
DR Pfam; PF08325; WLM; 1.
GO:0019985 translesion synthesis
ISO
GO_REF:0000024
ACCEPT
Summary: Wss1-mediated proteolytic removal of DNA-cross-linked proteins can permit downstream lesion bypass; participation in translesion synthesis does not assert that Wss1 is a DNA polymerase.
Reason: Wss1-mediated proteolytic removal of DNA-cross-linked proteins can permit downstream lesion bypass; participation in translesion synthesis does not assert that Wss1 is a DNA polymerase. Retain the curated orthology inference at that pathway-participation scope.
Supporting Evidence:
PMID:24998930
Notably, in vitro assays indicate that substrates such as topoisomerase 1 are processed by the metalloprotease directly and in a DNA-dependent manner.
GO:0106300 protein-DNA covalent cross-linking repair
ISO
GO_REF:0000024
ACCEPT
Summary: The target carries the diagnostic WLM domain and conserved metal-binding/catalytic residues.
Reason: The target carries the diagnostic WLM domain and conserved metal-binding/catalytic residues. Curated orthology and PAINT ancestry support inheritance of the Wss1 protease/DNA-repair function, whose mechanistic foundation is DNA-dependent proteolysis of cross-linked proteins in the characterized yeast homolog. This is a justified family inference, not target-specific experimental proof.
Supporting Evidence:
PMID:24998930
Notably, in vitro assays indicate that substrates such as topoisomerase 1 are processed by the metalloprotease directly and in a DNA-dependent manner.
file:SCHPO/wss1/wss1-uniprot.txt
DR Pfam; PF08325; WLM; 1.

Core Functions

WLM-family metalloendopeptidase inferred to remove protein components of DNA-protein cross-links and permit downstream repair.

Cellular Locations:
Supporting Evidence:
  • PMID:24998930
    Notably, in vitro assays indicate that substrates such as topoisomerase 1 are processed by the metalloprotease directly and in a DNA-dependent manner.
  • file:SCHPO/wss1/wss1-uniprot.txt
    DR Pfam; PF08325; WLM; 1.

References

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Suggested Questions for Experts

Q: Does Wss1 associate with any nuclear membrane or membrane-bound complex, or is its nuclear signal restricted to soluble chromatin-associated pools?

External Prediction Reviews

These computational predictions are reviewed separately from the GOA annotation set used for this review. The assessments below are from this project and do not constitute official GO annotations or endorsement by GO/UniProt. They are not included in the existing annotation review above.

ProtNLM2 External predictions

View prediction review YAML Β· wss1-protnlm-predictions-review.yaml Β· Review status: COMPLETE

Membrane residence remains unresolved: the source hydrates it from the Nucleus keyword, which does not establish membrane association.

Source documents: genes/SCHPO/wss1/wss1-protnlm-source.xml Β· genes/SCHPO/wss1/wss1-uniprot-source.json Β· genes/SCHPO/wss1/wss1-uniprot.txt

Review score: 2 = concordant with evidence; 1 = uncertain; 0 = discordant with evidence. This is an assessment score, not a model probability.

GO:0016020 membrane GO_CC
UNC β€” Uncertain Review score: 1/2
Prediction method: ProtNLM2 Β· Version: pre-release post-processed-2026_02_28k.xml Β· file:SCHPO/wss1/wss1-protnlm-source.xml
Review rationale: The reviewed target record places Wss1 in the nucleus and identifies a soluble WLM metalloprotease domain; it does not establish membrane residence. The original XML hydrates the membrane claim from KW-0539, which is the UniProt keyword Nucleus, not a membrane-specific observation. Nuclear residence alone does not entail association with the nuclear envelope, and the family DNA-protein-crosslink repair mechanism does not require a membrane. Nevertheless, lack of a transmembrane helix would not exclude peripheral membrane association, so the broad claim cannot be conclusively refuted solely from these data. Direct membrane fractionation, colocalization or an established membrane-associated complex is missing.
Supporting Evidence:

Deep Research

Falcon

(wss1-deep-research-falcon.md)

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πŸ“š Additional Documentation

Notes

(wss1-notes.md)

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πŸ“„ View Raw YAML

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