AGO3

UniProt ID: Q9H9G7
Organism: Homo sapiens
Review Status: COMPLETE
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Gene Description

AGO3 encodes human Argonaute-3, a small-RNA-binding RISC effector with a context-restricted slicer activity. Its core function is Argonaute-mediated small-RNA-guided gene silencing, with biochemical evidence for guide- and target-context-dependent RNA endonuclease activity and a non-core nuclear splicing role reported in immune cells.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0005634 nucleus
IBA
GO_REF:0000033
KEEP AS NON CORE
Summary: nucleus is a plausible or reported location/context but is not the core functional site annotation for AGO3.
Reason: The core location is cytoplasmic RISC/RNP granules; nuclear, membrane, synaptic, exosomal, or other compartment annotations should remain non-core unless tied to a specific curated mechanism.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005737 cytoplasm
IBA
GO_REF:0000033
ACCEPT
Summary: cytoplasm is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0004521 RNA endonuclease activity
IBA
GO_REF:0000033
ACCEPT
Summary: RNA endonuclease activity is supported for AGO3 as a conditional, guide-dependent slicer activity.
Reason: AGO3 is not the canonical slicer like AGO2, but biochemical studies support guide- and target-context-dependent RNA cleavage.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035194 regulatory ncRNA-mediated post-transcriptional gene silencing
IBA
GO_REF:0000033
ACCEPT
Summary: regulatory ncRNA-mediated post-transcriptional gene silencing is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0016442 RISC complex
IBA
GO_REF:0000033
ACCEPT
Summary: RISC complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0036464 cytoplasmic ribonucleoprotein granule
IBA
GO_REF:0000033
ACCEPT
Summary: cytoplasmic ribonucleoprotein granule is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0031054 pre-miRNA processing
IBA
GO_REF:0000033
MARK AS OVER ANNOTATED
Summary: pre-miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
Reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing, not as the Drosha/Dicer processing enzyme for pre-miRNAs.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035198 miRNA binding
IBA
GO_REF:0000033
ACCEPT
Summary: miRNA binding is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0003727 single-stranded RNA binding
IBA
GO_REF:0000033
MARK AS OVER ANNOTATED
Summary: single-stranded RNA binding is too generic for AGO3.
Reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC rather than a broad nucleic-acid or RNA-binding label.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
IEA
GO_REF:0000044
ACCEPT
Summary: P-body is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0003676 nucleic acid binding
IEA
GO_REF:0000002
MARK AS OVER ANNOTATED
Summary: nucleic acid binding is too generic for AGO3.
Reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC rather than a broad nucleic-acid or RNA-binding label.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0003723 RNA binding
IEA
GO_REF:0000120
MARK AS OVER ANNOTATED
Summary: RNA binding is too generic for AGO3.
Reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC rather than a broad nucleic-acid or RNA-binding label.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0003725 double-stranded RNA binding
IEA
GO_REF:0000117
MARK AS OVER ANNOTATED
Summary: double-stranded RNA binding is too generic for AGO3.
Reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC rather than a broad nucleic-acid or RNA-binding label.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0003727 single-stranded RNA binding
IEA
GO_REF:0000117
MARK AS OVER ANNOTATED
Summary: single-stranded RNA binding is too generic for AGO3.
Reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC rather than a broad nucleic-acid or RNA-binding label.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0016442 RISC complex
IEA
GO_REF:0000120
ACCEPT
Summary: RISC complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0031047 regulatory ncRNA-mediated gene silencing
IEA
GO_REF:0000002
ACCEPT
Summary: regulatory ncRNA-mediated gene silencing is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0031054 pre-miRNA processing
IEA
GO_REF:0000117
MARK AS OVER ANNOTATED
Summary: pre-miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
Reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing, not as the Drosha/Dicer processing enzyme for pre-miRNAs.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035198 miRNA binding
IEA
GO_REF:0000120
ACCEPT
Summary: miRNA binding is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035278 miRNA-mediated gene silencing by inhibition of translation
IEA
GO_REF:0000120
ACCEPT
Summary: miRNA-mediated gene silencing by inhibition of translation is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035279 miRNA-mediated gene silencing by mRNA destabilization
IEA
GO_REF:0000108
KEEP AS NON CORE
Summary: miRNA-mediated gene silencing by mRNA destabilization is plausible for AGO3 only as a context-dependent non-core process, not as an unqualified core AGO3 function.
Reason: AGO3 is primarily a small-RNA-binding RISC effector, and later biochemical work supports guide- and target-dependent slicing. Endogenous AGO3 cleavage targets remain unclear, so this broad IEA process should not be treated as a core positive annotation.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Importantly, this review emphasizes a major open question: **endogenous physiological target RNAs cleaved by AGO3 remain unknown**.
GO:0036464 cytoplasmic ribonucleoprotein granule
IEA
GO_REF:0000117
ACCEPT
Summary: cytoplasmic ribonucleoprotein granule is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0070578 RISC-loading complex
IEA
GO_REF:0000117
ACCEPT
Summary: RISC-loading complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0070922 RISC complex assembly
IEA
GO_REF:0000117
ACCEPT
Summary: RISC complex assembly is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0072091 regulation of stem cell proliferation
IEA
GO_REF:0000002
KEEP AS NON CORE
Summary: regulation of stem cell proliferation is downstream or context-specific for AGO3.
Reason: This process may be supported in a particular experiment or pathway model, but it is peripheral to the core Argonaute/RISC molecular role.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005515 protein binding
IPI
PMID:19167051
Importin 8 is a gene silencing factor that targets argonaute...
MARK AS OVER ANNOTATED
Summary: Generic protein binding does not capture the specific Argonaute/RISC function.
Reason: The cited interactions are best interpreted as RISC/cofactor context for small-RNA-guided silencing, not as a standalone core function.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005515 protein binding
IPI
PMID:19324964
The C-terminal half of human Ago2 binds to multiple GW-rich ...
MARK AS OVER ANNOTATED
Summary: Generic protein binding does not capture the specific Argonaute/RISC function.
Reason: The cited interactions are best interpreted as RISC/cofactor context for small-RNA-guided silencing, not as a standalone core function.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005515 protein binding
IPI
PMID:19383768
The C-terminal domains of human TNRC6A, TNRC6B, and TNRC6C s...
MARK AS OVER ANNOTATED
Summary: Generic protein binding does not capture the specific Argonaute/RISC function.
Reason: The cited interactions are best interpreted as RISC/cofactor context for small-RNA-guided silencing, not as a standalone core function.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005515 protein binding
IPI
PMID:23064648
DICER- and AGO3-dependent generation of retinoic acid-induce...
MARK AS OVER ANNOTATED
Summary: Generic protein binding does not capture the specific Argonaute/RISC function.
Reason: The cited interactions are best interpreted as RISC/cofactor context for small-RNA-guided silencing, not as a standalone core function.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005515 protein binding
IPI
PMID:28330616
Systematic Analysis of Human Protein Phosphatase Interaction...
MARK AS OVER ANNOTATED
Summary: Generic protein binding does not capture the specific Argonaute/RISC function.
Reason: The cited interactions are best interpreted as RISC/cofactor context for small-RNA-guided silencing, not as a standalone core function.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005515 protein binding
IPI
PMID:28683311
Argonaute Utilization for miRNA Silencing Is Determined by P...
MARK AS OVER ANNOTATED
Summary: Generic protein binding does not capture the specific Argonaute/RISC function.
Reason: The cited interactions are best interpreted as RISC/cofactor context for small-RNA-guided silencing, not as a standalone core function.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0000794 condensed nuclear chromosome
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: condensed nuclear chromosome is a plausible or reported location/context but is not the core functional site annotation for AGO3.
Reason: The core location is cytoplasmic RISC/RNP granules; nuclear, membrane, synaptic, exosomal, or other compartment annotations should remain non-core unless tied to a specific curated mechanism.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0010628 positive regulation of gene expression
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of gene expression reflects a reported nuclear or regulatory context rather than the main conserved AGO3 function.
Reason: Argonaute paralogs have reported nuclear/regulatory interactions, but the dominant conserved function remains small-RNA-guided post-transcriptional repression.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:1901224 positive regulation of non-canonical NF-kappaB signal transduction
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of non-canonical NF-kappaB signal transduction reflects a reported nuclear or regulatory context rather than the main conserved AGO3 function.
Reason: Argonaute paralogs have reported nuclear/regulatory interactions, but the dominant conserved function remains small-RNA-guided post-transcriptional repression.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035196 miRNA processing
NAS
PMID:28781232
Multivalent Recruitment of Human Argonaute by GW182.
MARK AS OVER ANNOTATED
Summary: miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
Reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing, not as the Drosha/Dicer processing enzyme for pre-miRNAs.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0036464 cytoplasmic ribonucleoprotein granule
IDA
GO_REF:0000052
ACCEPT
Summary: cytoplasmic ribonucleoprotein granule is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0004521 RNA endonuclease activity
IMP
PMID:29040713
Human Argonaute3 has slicer activity.
ACCEPT
Summary: RNA endonuclease activity is supported for AGO3 as a conditional, guide-dependent slicer activity.
Reason: AGO3 is not the canonical slicer like AGO2, but biochemical studies support guide- and target-context-dependent RNA cleavage.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0090624 endoribonuclease activity, cleaving miRNA-paired mRNA
IMP
PMID:29040713
Human Argonaute3 has slicer activity.
ACCEPT
Summary: endoribonuclease activity, cleaving miRNA-paired mRNA is supported for AGO3 as a conditional, guide-dependent slicer activity.
Reason: AGO3 is not the canonical slicer like AGO2, but biochemical studies support guide- and target-context-dependent RNA cleavage.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035196 miRNA processing
IMP
PMID:22795694
Slicing-independent RISC activation requires the argonaute P...
MARK AS OVER ANNOTATED
Summary: miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
Reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing, not as the Drosha/Dicer processing enzyme for pre-miRNAs.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0031054 pre-miRNA processing
IDA
PMID:19966796
ATP-dependent human RISC assembly pathways.
MARK AS OVER ANNOTATED
Summary: pre-miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
Reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing, not as the Drosha/Dicer processing enzyme for pre-miRNAs.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0003725 double-stranded RNA binding
IDA
PMID:19966796
ATP-dependent human RISC assembly pathways.
MARK AS OVER ANNOTATED
Summary: double-stranded RNA binding is too generic for AGO3.
Reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC rather than a broad nucleic-acid or RNA-binding label.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0003727 single-stranded RNA binding
IDA
PMID:19966796
ATP-dependent human RISC assembly pathways.
MARK AS OVER ANNOTATED
Summary: single-stranded RNA binding is too generic for AGO3.
Reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC rather than a broad nucleic-acid or RNA-binding label.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0016442 RISC complex
IDA
PMID:19966796
ATP-dependent human RISC assembly pathways.
ACCEPT
Summary: RISC complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0070578 RISC-loading complex
IDA
PMID:19966796
ATP-dependent human RISC assembly pathways.
ACCEPT
Summary: RISC-loading complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0070922 RISC complex assembly
IDA
PMID:19966796
ATP-dependent human RISC assembly pathways.
ACCEPT
Summary: RISC complex assembly is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0090625 siRNA-mediated gene silencing by mRNA destabilization
IDA NOT
PMID:15260970
Human Argonaute2 mediates RNA cleavage targeted by miRNAs an...
ACCEPT
Summary: This NOT annotation is appropriate for the PMID:15260970 assay context; AGO3 did not behave as the AGO2-like siRNA-guided target-cleavage Argonaute in that study.
Reason: PMID:15260970 supports AGO3 association with miRNAs/RISC but reports target-cleavage activity only for AGO2 in the tested assays. Later work supports conditional AGO3 slicing, so this NOT call should be treated as scoped to the 2004 AGO2-specific siRNA-cleavage context.
Supporting Evidence:
PMID:15260970
Purification of the FLAG/HA-epitope-tagged Ago containing complexes from different human cell lines revealed that endonuclease activity is exclusively associated with Ago2.
file:human/AGO3/AGO3-deep-research-falcon.md
Historically, mammalian **AGO2** was regarded as the only catalytically competent slicer, while AGO1/AGO3/AGO4 were categorized as non-slicer AGOs for most miRNA-mediated repression.
GO:0005737 cytoplasm
IDA
PMID:15260970
Human Argonaute2 mediates RNA cleavage targeted by miRNAs an...
ACCEPT
Summary: cytoplasm is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0016442 RISC complex
IDA
PMID:15260970
Human Argonaute2 mediates RNA cleavage targeted by miRNAs an...
ACCEPT
Summary: RISC complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035198 miRNA binding
IDA
PMID:15260970
Human Argonaute2 mediates RNA cleavage targeted by miRNAs an...
ACCEPT
Summary: miRNA binding is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035279 miRNA-mediated gene silencing by mRNA destabilization
IDA NOT
PMID:15260970
Human Argonaute2 mediates RNA cleavage targeted by miRNAs an...
REMOVE
Summary: This older NOT annotation should be removed because later AGO3-specific evidence supports guide-dependent miRNA-paired target cleavage.
Reason: PMID:15260970 did not detect AGO3-dependent target cleavage in the tested 2004 assay, but PMID:29040713 supersedes that broad negative by demonstrating guide- and target-dependent AGO3 slicer activity.
Supporting Evidence:
PMID:15260970
The specific role of Ago2 in guiding target RNA cleavage was confirmed independently by siRNA-based depletion of individual Ago members in combination with a sensitive positive-readout reporter assay.
PMID:29040713
recombinant AGO3 loaded with miR-20a cleaves complementary target RNAs, whereas AGO3 loaded with let-7a, miR-19b or miR-16 does not, indicating that AGO3 has slicer activity but that this activity depends on the guide RNA.
GO:0016020 membrane
HDA
PMID:19946888
Defining the membrane proteome of NK cells.
KEEP AS NON CORE
Summary: membrane is a plausible or reported location/context but is not the core functional site annotation for AGO3.
Reason: The core location is cytoplasmic RISC/RNP granules; nuclear, membrane, synaptic, exosomal, or other compartment annotations should remain non-core unless tied to a specific curated mechanism.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0003723 RNA binding
HDA
PMID:22658674
Insights into RNA biology from an atlas of mammalian mRNA-bi...
MARK AS OVER ANNOTATED
Summary: RNA binding is too generic for AGO3.
Reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC rather than a broad nucleic-acid or RNA-binding label.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0003723 RNA binding
HDA
PMID:22681889
The mRNA-bound proteome and its global occupancy profile on ...
MARK AS OVER ANNOTATED
Summary: RNA binding is too generic for AGO3.
Reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC rather than a broad nucleic-acid or RNA-binding label.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-1606561
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-1606682
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-1912362
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-1912363
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-1912364
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-1912366
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-1912367
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-1912368
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-1912406
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-2318752
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-3209151
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-426489
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-426522
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-4518575
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-5687103
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-8935766
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-8937097
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-8937134
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-8938440
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-8938487
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-8938507
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-8944650
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-8944684
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-8944706
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9011958
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9012203
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9012208
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9038545
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9618392
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9618486
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9624925
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9657791
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9858289
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9858309
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9924921
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9925095
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9925158
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9925159
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005829 cytosol
TAS
Reactome:R-HSA-9925324
ACCEPT
Summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0072091 regulation of stem cell proliferation
IMP
PMID:23064648
DICER- and AGO3-dependent generation of retinoic acid-induce...
KEEP AS NON CORE
Summary: regulation of stem cell proliferation is downstream or context-specific for AGO3.
Reason: This process may be supported in a particular experiment or pathway model, but it is peripheral to the core Argonaute/RISC molecular role.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0005515 protein binding
IPI
PMID:22915799
HIV-1 replication and APOBEC3 antiviral activity are not reg...
MARK AS OVER ANNOTATED
Summary: Generic protein binding does not capture the specific Argonaute/RISC function.
Reason: The cited interactions are best interpreted as RISC/cofactor context for small-RNA-guided silencing, not as a standalone core function.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0006402 mRNA catabolic process
IDA
PMID:18771919
Importance of translation and nonnucleolytic ago proteins fo...
KEEP AS NON CORE
Summary: mRNA catabolic process is downstream or context-specific for AGO3.
Reason: This process may be supported in a particular experiment or pathway model, but it is peripheral to the core Argonaute/RISC molecular role.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**
GO:0035278 miRNA-mediated gene silencing by inhibition of translation
IDA
PMID:18771919
Importance of translation and nonnucleolytic ago proteins fo...
ACCEPT
Summary: miRNA-mediated gene silencing by inhibition of translation is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
Reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression and related RISC complexes or cytoplasmic RNP compartments.
Supporting Evidence:
file:human/AGO3/AGO3-deep-research-falcon.md
Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**

Core Functions

Small-RNA-guided Argonaute/RISC effector function in post-transcriptional gene silencing.

Supporting Evidence:
  • file:human/AGO3/AGO3-deep-research-falcon.md
    Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**

Conditional RNA-guided endonuclease activity that depends on guide length/sequence and target context.

Supporting Evidence:
  • file:human/AGO3/AGO3-deep-research-falcon.md
    AGO3 cleavage is **guide-sequence and guide-length dependent**. miR-20a activates AGO3, whereas let-7a, miR-19b, and miR-16 do not in the reported assays.

References

Gene Ontology annotation through association of InterPro records with GO terms
Annotation inferences using phylogenetic trees
Gene Ontology annotation based on UniProtKB/Swiss-Prot Subcellular Location vocabulary mapping, accompanied by conservative changes to GO terms applied by UniProt
Gene Ontology annotation based on curation of immunofluorescence data
Automatic transfer of experimentally verified manual GO annotation data to orthologs using Ensembl Compara
Automatic assignment of GO terms using logical inference, based on on inter-ontology links
Electronic Gene Ontology annotations created by ARBA machine learning models
Combined Automated Annotation using Multiple IEA Methods
Human Argonaute2 mediates RNA cleavage targeted by miRNAs and siRNAs.
Importance of translation and nonnucleolytic ago proteins for on-target RNA interference.
Importin 8 is a gene silencing factor that targets argonaute proteins to distinct mRNAs.
The C-terminal half of human Ago2 binds to multiple GW-rich regions of GW182 and requires GW182 to mediate silencing.
The C-terminal domains of human TNRC6A, TNRC6B, and TNRC6C silence bound transcripts independently of Argonaute proteins.
Defining the membrane proteome of NK cells.
ATP-dependent human RISC assembly pathways.
Insights into RNA biology from an atlas of mammalian mRNA-binding proteins.
The mRNA-bound proteome and its global occupancy profile on protein-coding transcripts.
Slicing-independent RISC activation requires the argonaute PAZ domain.
HIV-1 replication and APOBEC3 antiviral activity are not regulated by P bodies.
DICER- and AGO3-dependent generation of retinoic acid-induced DR2 Alu RNAs regulates human stem cell proliferation.
Systematic Analysis of Human Protein Phosphatase Interactions and Dynamics.
Argonaute Utilization for miRNA Silencing Is Determined by Phosphorylation-Dependent Recruitment of LIM-Domain-Containing Proteins.
Multivalent Recruitment of Human Argonaute by GW182.
Human Argonaute3 has slicer activity.
Reactome:R-HSA-1606561
MIR449 microRNAs bind 3'UTR of NOTCH1 mRNA
Reactome:R-HSA-1606682
MIR34 microRNAs bind 3'UTR of NOTCH1 mRNA
Reactome:R-HSA-1912362
MIR150 microRNA binds 3'UTR of NOTCH3 mRNA
Reactome:R-HSA-1912363
MIR200B/C microRNAs bind NOTCH1 mRNA
Reactome:R-HSA-1912364
MIR181C microRNA binds 3'UTR of NOTCH4 mRNA
Reactome:R-HSA-1912366
MIR206 microRNA binds 3'UTR of NOTCH3 mRNA
Reactome:R-HSA-1912367
MIR34 microRNAs bind 3'UTR of NOTCH2 mRNA
Reactome:R-HSA-1912368
MIR302A microRNA binds 3'UTR of NOTCH4 mRNA
Reactome:R-HSA-1912406
p53 positively regulates transcription of MIR34 microRNAs
Reactome:R-HSA-2318752
miR-26A microRNAs bind PTEN mRNA
Reactome:R-HSA-3209151
miR-24 binds p16INK4A and p14ARF mRNAs
Reactome:R-HSA-426489
RISC binds inexactly matching target RNAs
Reactome:R-HSA-426522
Nonendonucleolytic RISC binds exactly matching target RNAs
Reactome:R-HSA-4518575
miR-92b binds 3'UTR of NLK mRNA
Reactome:R-HSA-5687103
FOXO3 regulates MIR34B,C expression
Reactome:R-HSA-8935766
miR-378 binds RUNX1 mRNA
Reactome:R-HSA-8937097
MIR27A gene transcription is stimulated by the complex containing RUNX1, PRMT1 and GATA1 and inhibited by the complex of RUNX1, SIN3A and PRMT6
Reactome:R-HSA-8937134
miR-27a binds RUNX1 mRNA
Reactome:R-HSA-8938440
miR-17 binds RUNX1 mRNA
Reactome:R-HSA-8938487
miR-20a binds RUNX1 mRNA
Reactome:R-HSA-8938507
miR-106a binds RUNX1 mRNA
Reactome:R-HSA-8944650
miR-214 microRNA binds PTEN mRNA
Reactome:R-HSA-8944684
miR-205 microRNA binds PTEN mRNA
Reactome:R-HSA-8944706
miR-21 nonendonucleolytic RISC binds PTEN mRNA
Reactome:R-HSA-9011958
miR-26A and B bind to the 3'UTR of the GREB1 mRNA
Reactome:R-HSA-9012203
miR-26A and B bind to the 3'UTR of the CHD11 mRNA
Reactome:R-HSA-9012208
miR-26A and B bind to the 3'UTR of the KPNA2 mRNA
Reactome:R-HSA-9038545
miR-613 binds to the 3'UTR of the NR1H3 mRNA
Reactome:R-HSA-9618392
miR-26 binds to the 3'UTR of the ARL4C mRNA
Reactome:R-HSA-9618486
miR-26 binds to the 3'UTR of the ABCA1 mRNA
Reactome:R-HSA-9624925
miR-33 binds to the 3'UTR of the ABCA1 mRNA
Reactome:R-HSA-9657791
miR-144 binds to the 3'UTR of the ABCA1 mRNA
Reactome:R-HSA-9858289
MITF-M-dependent miR-211 expression
Reactome:R-HSA-9858309
miR-211 RISC binds POU3F2 mRNA
Reactome:R-HSA-9924921
CD274 mRNA binds miR-34 RISC
Reactome:R-HSA-9925095
CD274 mRNA binds miR-340 RISC
Reactome:R-HSA-9925158
CD274 mRNA binds miR-152 RISC
Reactome:R-HSA-9925159
CD274 mRNA binds miR-140 RISC
Reactome:R-HSA-9925324
CD274 mRNA binds miR-200B/C RISC
file:human/AGO3/AGO3-uniprot.txt
UniProt text export for AGO3
  • UniProt identifies AGO3 and its protein family/domain context.
    "Belongs to the argonaute family. Ago subfamily."
file:human/AGO3/AGO3-deep-research-falcon.md
Falcon deep research report for AGO3
  • Falcon research supports the reviewed core function of AGO3.
    "Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding Argonaute effector**"

Suggested Questions for Experts

Q: Which endogenous human tissues or stress states require AGO3-specific RISC activity rather than redundant activity from other AGO paralogs?

Q: Which protein cofactors determine guide loading, localization, and non-core nuclear or granule-associated functions of AGO3?

Suggested Experiments

Experiment: Endogenous tagging of AGO3 followed by small-RNA CLIP, target RNA profiling, and paralog-specific rescue in AGO-depleted human cells.

Hypothesis: AGO3 has paralog-specific guide, target, and compartment preferences within human RISC biology.

Type: genome editing/RNA-protein interaction profiling

Experiment: Biochemical reconstitution of AGO3-RISC with representative miRNA and siRNA guides to compare target repression, target cleavage, and TNRC6 recruitment.

Hypothesis: AGO3 activity is best explained by guide-loaded RISC function rather than generic RNA or protein binding.

Type: biochemical reconstitution

Deep Research

Falcon

(AGO3-deep-research-falcon.md)
Comprehensive Research Report: Human **AGO3 / EIF2C3** (UniProt **Q9H9G7**) Functional Annotation Falcon Edison Scientific Literature 31 citations 2026-05-02T23:07:50.846094

The research report should be a detailed narrative explaining the function, biological processes, and localization of the gene product. Citations should be given for all claims.

You should prioritize authoritative reviews and primary scientific literature when conducting research. You can supplement
this with annotations you find in gene/protein databases, but these can be outdated or inaccurate.

We are specifically interested in the primary function of the gene - for enzymes, what reaction is catalyzed, and what is the substrate specificity? For transporters, what is the substrate? For structural proteins or adapters, what is the broader structural role? For signaling molecules, what is the role in the pathway.

We are interested in where in or outside the cell the gene product carries out its function.

We are also interested in the signaling or biochemical pathways in which the gene functions. We are less interested in broad pleiotropic effects, except where these elucidate the precise role.

Include evidence where possible. We are interested in both experimental evidence as well as inference from structure, evolution, or bioinformatic analysis. Precise studies should be prioritized over high-throughput, where available.

Comprehensive Research Report: Human AGO3 / EIF2C3 (UniProt Q9H9G7) Functional Annotation

0. Target identity verification (mandatory)

All evidence summarized below pertains to Homo sapiens Argonaute-3 (AGO3; synonym EIF2C3), consistent with UniProt accession Q9H9G7 and membership in the eukaryotic Argonaute (Ago) subfamily that binds small RNAs in the miRNA/siRNA pathway. Primary studies explicitly purify and characterize human AGO3 protein and/or compare it directly to other human AGO paralogs (AGO1–AGO4), avoiding symbol ambiguity with non-human “AGO3” proteins. (park2017humanargonaute3has pages 1-1, park2020humanargonaute2and pages 1-2)

1. Key concepts and definitions (current understanding)

1.1 Argonaute proteins and miRISC

In mammals, Argonaute (AGO) proteins are the core effector proteins of the microRNA-induced silencing complex (miRISC), in which a loaded small RNA (typically a ~22-nt miRNA) guides the AGO-containing complex to complementary RNA targets to repress gene expression post-transcriptionally. RISC loading is coupled to DICER processing of small-RNA duplexes and involves chaperones HSC70/HSP90; canonical models emphasize ATP-dependent loading and passenger-strand removal to form a mature guide-loaded AGO. (narasimhan2025partnersinsilencing pages 1-2)

1.2 “Slicer” activity and catalytic center

“Slicer” refers to RNA-guided endonuclease cleavage of a target RNA by an AGO protein’s PIWI domain (an RNase H-like fold). Historically, mammalian AGO2 was regarded as the only catalytically competent slicer, while AGO1/AGO3/AGO4 were categorized as non-slicer AGOs for most miRNA-mediated repression. However, human AGO3 retains the canonical catalytic tetrad and can cleave targets under specific conditions (guide-dependent), indicating that “non-slicer” is context-dependent for AGO3. (narasimhan2025partnersinsilencing pages 1-2, park2017humanargonaute3has pages 1-1)

1.3 Domain architecture relevant to function

Eukaryotic AGOs share a conserved architecture consisting of N, PAZ, MID, and PIWI domains connected by linker regions. In this framework, the PAZ domain anchors the guide RNA 3′ end and the MID domain binds the 5′ end, while the PIWI domain contains the catalytic center for cleavage when slicing occurs; structural features of the nucleic-acid binding channel shape guide/target geometry and substrate requirements. (martinmerchan2023domainorganizationexpression pages 5-6, park2017humanargonaute3has pages 7-8)

2. Molecular function of human AGO3 (what reaction; substrate specificity)

2.1 Primary enzymatic function: conditional RNA-guided endonuclease

Reaction: When activated, AGO3 functions as an RNA-guided endoribonuclease, cleaving a complementary target RNA directed by a loaded guide RNA (miRNA/siRNA-like). (park2017humanargonaute3has pages 1-1)

Substrate specificity constraints: Experimental evidence indicates AGO3 slicing is highly dependent on the guide RNA identity and length and additionally requires extended target context (flanking regions) not required for AGO2. (park2017humanargonaute3has pages 1-1, park2017humanargonaute3has pages 7-8)

2.2 Evidence that human AGO3 has slicer activity (2017 structural/biochemical breakthrough)

A foundational biochemical and structural study purified recombinant human AGO3 and showed:

  • Guide dependence: AGO3 loaded with miR-20a cleaved a complementary RNA target, whereas AGO3 loaded with let-7a, miR-19b, or miR-16 did not, demonstrating that AGO3 slicing depends on the guide RNA sequence. (park2017humanargonaute3has pages 1-1)
  • Target-context dependence: AGO3 cleaved a 60-nt miR-20a target but not a 23-nt target; deletion of either 5′ or 3′ flanking sequence around the binding site strongly reduced cleavage, indicating a requirement for both 5′ and 3′ flanking regions for efficient slicing. (park2017humanargonaute3has pages 7-8)
  • Catalytic-site requirement: Cleavage is abolished by a catalytic mutation (E638A), consistent with bona fide enzymatic slicing. (park2017humanargonaute3has pages 1-2, park2017humanargonaute3has media c985c1a4)

Quantitative examples from figures: In the reported assays, AGO3 slicing with miR-20a showed measurable cleavage (e.g., ~7.0% in one guide condition), and concentration-dependent assays showed ~9.2–18.7% cleavage; the catalytic mutant eliminated cleavage. (park2017humanargonaute3has media c985c1a4)

2.3 Structural basis for AGO3’s constrained slicing

The same study reported a 3.28 Å crystal structure of human AGO3-RISC (PDB referenced in text) showing:

  • A complete catalytic tetrad (DEDH) in AGO3’s PIWI domain; the inset identifies residues D598, E638, D670, H808. (park2017humanargonaute3has pages 7-8, park2017humanargonaute3has media c985c1a4)
  • An “immature/imperfect” nucleic-acid binding channel relative to AGO2, consistent with the empirical requirement for target flanking sequences and the notion that accessory factors could modulate AGO3’s channel and function in cells. (park2017humanargonaute3has pages 1-1, park2017humanargonaute3has pages 7-8)

These structural observations support a model in which AGO3 is catalytically competent but geometrically constrained, making cleavage difficult unless specific guide/target configurations are met. (park2017humanargonaute3has pages 1-1, park2017humanargonaute3has pages 7-8)

3. Small-RNA partners and guide-dependent activation (cityRNAs and tyRNAs)

3.1 CityRNAs: unusually short guides that strongly activate AGO3 (2020)

A subsequent study established that AGO3 can become a highly competent slicer when programmed with specific 14-nt guide RNAs, termed cleavage-inducing tiny guide RNAs (cityRNAs):

  • 14-nt 3′-shortened variants of let-7a, miR-27a, and miR-17–92 family miRNAs can activate AGO3, increasing target cleavage up to ~82-fold in some instances. (park2020humanargonaute2and pages 1-2)
  • In lysate-based assays, FLAG-AGO3 achieved ~50% target cleavage when supplied with 14-nt single-stranded miR-20a, whereas FLAG-AGO2 produced ~30% cleavage with a canonical 23-nt duplex miR-20a in the compared setup, supporting distinct optimal guide lengths for AGO2 vs AGO3. (park2020humanargonaute2and pages 1-2)
  • A catalytically dead AGO3 mutant (E638A) was used as a control to support that cleavage is catalytic. (park2020humanargonaute2and pages 1-2)

These results revise the operational definition of AGO3 as strictly “non-slicer,” showing length- and sequence-conditional activation.

3.2 2024 synthesis: a proposed physiological route to AGO3 activation via guide trimming

A 2024 expert review integrates these findings into a mechanistic model in which 3′→5′ exonucleases trim AGO-associated miRNAs into ~13–14 nt tiny RNAs capable of activating AGO3:

  • ISG20, TREX1, and ERI1 are discussed as enzymes that can trim AGO-associated miRNAs to short lengths. (nakanishi2024whenargonautetakes pages 9-11)
  • The review highlights evidence that ISG20-mediated trimming can convert AGO3-RISC into a slicer for particular miRNAs (e.g., trimmed miR-20a and let-7a), while other miRNAs (e.g., miR-16, miR-19b) do not activate AGO3 even when shortened, consistent with sequence dependence. (nakanishi2024whenargonautetakes pages 9-11)
  • A proposed physiological trigger is innate immune activation, since ISG20 is interferon-stimulated and may be upregulated in viral infection/stress; the review also notes a requirement for Mn2+ (not Mg2+) for ISG20 trimming and suggests mitochondria may release manganese into the cytoplasm during viral infection, potentially enabling the trimming reaction. (nakanishi2024whenargonautetakes pages 9-11)

Importantly, this review emphasizes a major open question: endogenous physiological target RNAs cleaved by AGO3 remain unknown. (nakanishi2024whenargonautetakes pages 1-3)

4. Biological processes, pathways, and cellular roles

4.1 Canonical role: miRNA/siRNA pathway effector (miRISC)

AGO3 participates in the canonical miRISC framework shared by mammalian AGOs, including assembly via the RISC loading complex and interactions with core miRISC cofactors (review-supported general mechanisms). (narasimhan2025partnersinsilencing pages 1-2, narasimhan2025partnersinsilencing pages 4-5)

4.2 Non-canonical AGO3-specific nuclear function: regulation of pre-mRNA splicing

Beyond cytoplasmic miRISC repression, AGO3 has been reported (via authoritative review synthesis of primary literature) to regulate pre-mRNA splicing in Th2 cells through a specific, RNA-independent nuclear interaction with SF3B3:

  • The AGO3–SF3B3 interaction is described as controlling global pre-mRNA splicing in T cells and regulating IL-13 expression via Nisch pre-mRNA. (narasimhan2025partnersinsilencing pages 10-11)
  • This indicates AGO3 can act as a nuclear regulator of RNA processing, distinct from its guide-mediated slicing function. (narasimhan2025partnersinsilencing pages 10-11)

5. Subcellular localization (where AGO3 acts)

Direct AGO3 localization mapping is less extensive in the available sources than for AGO2, but two evidence-supported localization inferences are robust:

  1. Nuclear presence/function in Th2 cells: AGO3 performs a nuclear function through interaction with the spliceosome-associated factor SF3B3, implying nuclear localization in that context. (narasimhan2025partnersinsilencing pages 10-11)
  2. Cytoplasmic RISC activity under appropriate guide conditions: The biochemical and lysate-based slicing assays demonstrate that AGO3 can perform cytosolic RISC-like cleavage when provided with activating guides (miR-20a; 14-nt cityRNAs), consistent with cytoplasmic activity when the correct small RNAs are available. (park2020humanargonaute2and pages 1-2, park2017humanargonaute3has pages 1-1)

More generally, mammalian Argonaute proteins can accumulate in nuclei under particular conditions (e.g., environmental/cellular-state changes), highlighting compartmental plasticity of AGO biology; however, these data are AGO2-focused and should not be over-interpreted as AGO3-specific without direct AGO3 measurements. (chen2026decodingargonautespecificity pages 7-10)

6. Recent developments (prioritizing 2023–2024)

6.1 2024: Mechanistic consolidation of AGO3 activation and innate immunity linkage

The 2024 Journal of Biological Chemistry review frames AGO3 as a conditionally deployable slicer whose activation may be coupled to interferon-stimulated RNA metabolism (ISG20) and the generation of tiny RNAs/cityRNAs, potentially shifting slicing capacity toward AGO3 in antiviral states. It also highlights key unknowns, including the identity of endogenous AGO3 cleavage targets and when AGO3 slicing occurs in vivo. (nakanishi2024whenargonautetakes pages 9-11, nakanishi2024whenargonautetakes pages 1-3)

6.2 2023–2024: Clinical studies measuring AGO3 autoantibodies

While not a direct functional assay, two 2023–2024 clinical lines of evidence incorporate AGO3 into real-world measurements:

  • SLE autoantigen discovery (2023): A proteome-wide screen (>21,000 proteins) identified AGO1/AGO2/AGO3 among IgG/IgA-reactive antigens in SLE, with ELISA validation across two cohorts (n=49 and n=46 SLE; controls including n=48 healthy plus multiple rheumatic-disease control groups). Positivity thresholds used the 95th percentile of healthy donors. However, AGO3-specific prevalence/clinical correlations were not clearly quantified in the extracted text. (moadab2023argonautevaultand pages 1-3, moadab2023argonautevaultand pages 3-4)
  • HBV-related cirrhosis/ACLF (2024): A conformation-stabilizing ELISA study screened AGO1–3 autoantibodies and reported AGO3-Ab medians (IQR) across disease stages (e.g., HC 12.3 (10.5–23.4); CHB 15.3 (11.5–29.3); CLC 19.7 (14.6–31.6); DLC 19.7 (12.2–24.2); ACLF 17.5 (13.9–25.2), mg/ml). The primary prognostic biomarker was AGO2-Ab, whereas AGO3-Ab showed weaker/variable associations. (wang2024argonaute2autoantibodiesa pages 2-3, wang2024argonaute2autoantibodiesa pages 3-6)

7. Current applications and real-world implementations

7.1 Diagnostics/biomarkers: AGO3 autoantibodies as measured analytes

Implementation status: AGO3 is being measured in human cohorts as an autoantigen/autoantibody target, but current evidence supports it more as part of broader Argonaute autoantibody panels than as a validated standalone biomarker.

  • In SLE, AGO3 is one of the microRNA-processing proteins recognized by autoantibodies in proteome-wide screening and follow-up ELISA workflows, but quantitative performance metrics for AGO3 specifically (sensitivity/specificity) are not provided in the available excerpts. (moadab2023argonautevaultand pages 1-3, moadab2023argonautevaultand pages 9-15)
  • In HBV-related cirrhosis/ACLF, AGO3-Ab levels were quantified by ELISA, but the study emphasizes AGO2-Ab as prognostic and suggests AGO3-Ab may be intermittently detected or lack severity correlation. (wang2024argonaute2autoantibodiesa pages 2-3, wang2024argonaute2autoantibodiesa pages 9-10)

7.2 Translational mechanistic concept: leveraging AGO3 cityRNA activation

Although not yet a clinical implementation, the discovery that 14-nt cityRNAs can strongly activate AGO3 slicing suggests potential future applications in:

  • Programmable RNA cleavage systems using AGO3 with engineered short guides.
  • Antiviral RNAi augmentation if interferon-induced trimming indeed generates endogenous cityRNAs that activate AGO3 in vivo.

However, the key limitation remains that physiological AGO3 cleavage targets have not been identified, and thus therapeutic exploitation is currently speculative and preclinical. (nakanishi2024whenargonautetakes pages 9-11, nakanishi2024whenargonautetakes pages 1-3)

8. Expert opinions and analysis (authoritative synthesis)

8.1 Consensus view evolving: AGO3 is catalytically competent but context-restricted

Recent expert synthesis recognizes AGO3 as retaining the catalytic machinery (DEDH tetrad) and having the capacity for slicing under defined guide conditions, challenging earlier categorical statements that only AGO2 can slice. (nakanishi2024whenargonautetakes pages 1-3, narasimhan2025partnersinsilencing pages 4-5)

8.2 Mechanistic interpretation: structural constraints and regulatory gating

Structural data indicate AGO3’s active site is intact, but its nucleic-acid binding channel is less optimized than AGO2’s, providing a plausible mechanistic rationale for why AGO3 requires (i) specific guides (miR-20a/cityRNAs) and (ii) extended target flanks or specialized conformations to achieve cleavage. This supports a “regulated latent nuclease” model for AGO3. (park2017humanargonaute3has pages 1-1, park2017humanargonaute3has pages 7-8)

8.3 Open questions emphasized by reviews

Authoritative reviews highlight outstanding gaps that limit complete functional annotation:

  • Which endogenous targets (if any) are cleaved by AGO3 in vivo? (nakanishi2024whenargonautetakes pages 1-3)
  • Under what physiological conditions are cityRNAs/tyRNAs produced at sufficient abundance to activate AGO3? (nakanishi2024whenargonautetakes pages 9-11)
  • How do accessory proteins and cellular compartmentalization modulate AGO3 slicing versus non-slicing functions? (park2017humanargonaute3has pages 1-1, narasimhan2025partnersinsilencing pages 10-11)

9. Key statistics and data (from recent studies and primary mechanistic work)

9.1 Enzymology / mechanistic statistics

  • Up to ~82-fold increase in AGO3 target cleavage with 14-nt cityRNAs (PNAS 2020; URL: https://doi.org/10.1073/pnas.2015026117; published Oct 2020). (park2020humanargonaute2and pages 1-2)
  • ~50% cleavage by FLAG-AGO3 in lysate assay with 14-nt miR-20a in the described setup (vs ~30% for FLAG-AGO2 with 23-nt duplex miR-20a in the compared condition). (park2020humanargonaute2and pages 1-2)
  • Measurable recombinant AGO3 cleavage with miR-20a, including ~7.0% in one condition and ~9.2–18.7% across concentration conditions; catalytic mutant (E638A) abrogates cleavage (NAR 2017; URL: https://doi.org/10.1093/nar/gkx916; published online 11 Oct 2017). (park2017humanargonaute3has media c985c1a4)

9.2 Clinical cohort statistics where AGO3 is measured

  • SLE autoantigen discovery (2023; J Rheumatol; URL: https://doi.org/10.3899/jrheum.2022-1327; May 2023): initial screen n=12 SLE and n=6 healthy; validation cohorts include n=49 and n=46 SLE; healthy controls include n=48 plus disease controls (RA n=20; SjS n=12; AAV n=11; LVV n=9; PsA n=8). AGO3 included among antigens tested, but AGO3-specific prevalence values are not explicitly stated in the extracted text. (moadab2023argonautevaultand pages 1-3, moadab2023argonautevaultand pages 3-4)
  • HBV-related cirrhosis/ACLF (2024; Front Cell Infect Microbiol; URL: https://doi.org/10.3389/fcimb.2024.1407064; Jul 2024): AGO3 autoantibodies reported as medians (IQR) across clinical strata, including 12.3 (10.5–23.4) in healthy controls and 17.5 (13.9–25.2) in ACLF (mg/ml), with intermediate values for CHB/CLC/DLC. (wang2024argonaute2autoantibodiesa pages 2-3, wang2024argonaute2autoantibodiesa pages 3-6)

10. Summary of functional annotation (evidence-weighted)

Human AGO3 (EIF2C3; Q9H9G7) is best annotated as a small-RNA-binding Argonaute effector with:

  1. Canonical miRISC-associated roles shared with mammalian AGO paralogs in miRNA-mediated gene silencing (repression/decay) (narasimhan2025partnersinsilencing pages 1-2, narasimhan2025partnersinsilencing pages 4-5),
  2. A conditionally active RNA-guided endonuclease (“slicer”) function that requires specific guide RNAs (miR-20a or 14-nt cityRNAs) and extended target context, supported by biochemical assays, catalytic mutagenesis, and crystal structures (park2017humanargonaute3has pages 1-1, park2020humanargonaute2and pages 1-2, park2017humanargonaute3has media c985c1a4), and
  3. A distinct non-canonical nuclear role in pre-mRNA splicing via interaction with SF3B3 in Th2 cells, indicating AGO3 can contribute to immune regulation outside canonical cytoplasmic slicing. (narasimhan2025partnersinsilencing pages 10-11)

11. Evidence summary table

Aspect Key findings Evidence type Key sources (author year journal) with DOI URLs Quantitative data/statistics (if any)
identity/domains Target verified as human AGO3 / EIF2C3 / Argonaute-3, one of four human AGO paralogs. AGO3 has canonical Argonaute architecture (N, PAZ, MID, PIWI with linkers) and retains the catalytic DEDH tetrad in PIWI; crystal structure showed a complete active site but a less well-defined nucleic-acid binding channel than AGO2. (park2017humanargonaute3has pages 1-1, park2017humanargonaute3has pages 7-8, martinmerchan2023domainorganizationexpression pages 5-6) structural, biochemical, review Park et al. 2017 Nucleic Acids Res. https://doi.org/10.1093/nar/gkx916; Martín-Merchán et al. 2023 J Exp Bot. https://doi.org/10.1093/jxb/erad030 Crystal structure resolved to 3.28 Å; catalytic residues visualized as D598, E638, D670, H808. (park2017humanargonaute3has pages 7-8, park2017humanargonaute3has media c985c1a4)
catalytic activity AGO3 is not constitutively inactive: recombinant human AGO3 can cleave RNA when loaded with certain guides, establishing bona fide slicer activity under restricted conditions. This revises the older simplification that only AGO2 slices in humans. (park2017humanargonaute3has pages 1-1, park2020humanargonaute2and pages 1-2, narasimhan2025partnersinsilencing pages 4-5) biochemical, structural, review Park et al. 2017 Nucleic Acids Res. https://doi.org/10.1093/nar/gkx916; Park et al. 2020 PNAS https://doi.org/10.1073/pnas.2015026117; Narasimhan & Erkeland 2025 Non-Coding RNA https://doi.org/10.3390/ncrna11040062 In Park 2017, AGO3 loaded with miR-20a showed measurable cleavage, including ~7.0% in one assay and 9.2–18.7% across concentration-dependent assays; catalytic mutant E638A abolished activity. (park2017humanargonaute3has media c985c1a4)
guide requirements AGO3 cleavage is guide-sequence and guide-length dependent. miR-20a activates AGO3, whereas let-7a, miR-19b, and miR-16 do not in the reported assays. Later work showed specific 14-nt tyRNAs/cityRNAs (e.g., shortened miR-20a, let-7a, miR-27a, miR-17-92-family guides) strongly activate AGO3. (park2017humanargonaute3has pages 1-1, park2020humanargonaute2and pages 1-2, park2020humanargonaute2and pages 2-3) biochemical Park et al. 2017 Nucleic Acids Res. https://doi.org/10.1093/nar/gkx916; Park et al. 2020 PNAS https://doi.org/10.1073/pnas.2015026117 14-nt cityRNAs increased AGO3 cleavage up to ~82-fold; in lysate-based assays, FLAG-AGO3 reached ~50% cleavage with 14-nt miR-20a, whereas FLAG-AGO2 favored 23-nt duplex miR-20a at ~30% in the compared setup. (park2020humanargonaute2and pages 1-2)
target requirements AGO3 has stricter substrate requirements than AGO2. Cleavage depends on the post-seed region of the guide and requires target RNAs with both 5′- and 3′-flanking regions around the complementary site; short minimal targets are poorly cleaved. (park2017humanargonaute3has pages 1-1, park2017humanargonaute3has pages 4-5, park2017humanargonaute3has pages 7-8) biochemical, structural Park et al. 2017 Nucleic Acids Res. https://doi.org/10.1093/nar/gkx916 AGO3 cleaved a 60-nt miR-20a target but not a 23-nt target; deleting either 5′ or 3′ flanks drastically reduced activity. Seed substitutions at g7/g8 lowered cleavage. (park2017humanargonaute3has pages 7-8, park2017humanargonaute3has pages 4-5)
non-canonical roles AGO3 also has an AGO3-specific nuclear role in pre-mRNA splicing in type 2 T cells, acting through an RNA-independent interaction with SF3B3 rather than through canonical cytoplasmic slicing alone. Reported consequences include control of global splicing and regulation of IL-13 via Nisch pre-mRNA. (narasimhan2025partnersinsilencing pages 10-11) cellular, molecular, review Narasimhan & Erkeland 2025 Non-Coding RNA https://doi.org/10.3390/ncrna11040062 No direct effect size quoted in available context, but review describes AGO3 as essential for type 2 T-cell responses through this pathway. (narasimhan2025partnersinsilencing pages 10-11)
localization Direct AGO3 localization evidence is limited in the available source set. AGO3 is clearly implicated in the nucleus for SF3B3-linked splicing in Th2 cells; broader human AGO literature supports nuclear AGO family presence, but AGO3-specific compartment maps remain less well resolved than for AGO2. (narasimhan2025partnersinsilencing pages 10-11, chen2026decodingargonautespecificity pages 7-10) cellular, review Narasimhan & Erkeland 2025 Non-Coding RNA https://doi.org/10.3390/ncrna11040062; Chen & Phillips 2026 RNA https://doi.org/10.1261/rna.080816.125 Qualitative only in available context: nuclear AGO3 function inferred from the documented AGO3-SF3B3 splicing complex. (narasimhan2025partnersinsilencing pages 10-11)
interactions/complexes In canonical RNA silencing, AGO3 participates in miRISC and likely shares core interactors such as GW182/TNRC6 and regulatory post-translational modification pathways with other mammalian AGOs. AGO3 additionally forms an RNA-independent nuclear complex with SF3B3 for splicing control. (narasimhan2025partnersinsilencing pages 10-11, narasimhan2025partnersinsilencing pages 1-2, narasimhan2025partnersinsilencing pages 4-5) review, cellular Narasimhan & Erkeland 2025 Non-Coding RNA https://doi.org/10.3390/ncrna11040062 No AGO3-specific stoichiometric values in available context. Review notes AGO1/3/4 are generally considered non-slicer during passenger-strand removal, but AGO3 is an exception under specific guide conditions. (narasimhan2025partnersinsilencing pages 1-2, narasimhan2025partnersinsilencing pages 4-5)
disease/clinical associations Available recent evidence links AGO3 to immune regulation and autoimmunity-related seroreactivity rather than a validated AGO3-specific therapy. AGO3 has been identified as an autoantigen target in systemic lupus erythematosus screens, and broader clinical literature/reviews suggest paralog-specific roles in disease, but AGO3-specific translational applications remain limited. (narasimhan2025partnersinsilencing pages 10-11) clinical proteomic/screening context, review Narasimhan & Erkeland 2025 Non-Coding RNA https://doi.org/10.3390/ncrna11040062 No robust AGO3-only clinical performance metric available in the cited AGO3-specific context; evidence is currently more mechanistic than implementational. (narasimhan2025partnersinsilencing pages 10-11)

Table: This table summarizes validated functional annotation evidence for human AGO3 (UniProt Q9H9G7/EIF2C3), separating AGO3-specific findings from broader Argonaute-family statements. It highlights where evidence is strong (conditional slicer activity, structural features, nuclear splicing role) and where AGO3-specific data remain limited (localization, clinical implementation).

12. Key URLs (most relevant)

  • Park et al., 2017-10-11 (online), Nucleic Acids Research: “Human Argonaute3 has slicer activity.” https://doi.org/10.1093/nar/gkx916 (park2017humanargonaute3has pages 1-1)
  • Park et al., 2020-10 (PNAS), “Human Argonaute2 and Argonaute3 are catalytically activated by different lengths of guide RNA.” https://doi.org/10.1073/pnas.2015026117 (park2020humanargonaute2and pages 1-2)
  • Nakanishi, 2024-01 (J Biol Chem), “When Argonaute takes out the ribonuclease sword.” https://doi.org/10.1016/j.jbc.2023.105499 (nakanishi2024whenargonautetakes pages 1-3)
  • Moadab et al., 2023-05 (J Rheumatol), “Argonaute, Vault, and Ribosomal Proteins Targeted by Autoantibodies in Systemic Lupus Erythematosus.” https://doi.org/10.3899/jrheum.2022-1327 (moadab2023argonautevaultand pages 1-3)
  • Wang et al., 2024-07 (Front Cell Infect Microbiol), “Argonaute-2 autoantibodies…” (includes AGO3-Ab quantification). https://doi.org/10.3389/fcimb.2024.1407064 (wang2024argonaute2autoantibodiesa pages 1-2)

References

  1. (park2017humanargonaute3has pages 1-1): Mi Seul Park, Hong-Duc Phan, Florian Busch, Samantha H. Hinckley, James A. Brackbill, Vicki H. Wysocki, and Kotaro Nakanishi. Human argonaute3 has slicer activity. Nucleic Acids Research, 45:11867-11877, Oct 2017. URL: https://doi.org/10.1093/nar/gkx916, doi:10.1093/nar/gkx916. This article has 148 citations and is from a highest quality peer-reviewed journal.

  2. (park2020humanargonaute2and pages 1-2): Mi Seul Park, GeunYoung Sim, Audrey C. Kehling, and Kotaro Nakanishi. Human argonaute2 and argonaute3 are catalytically activated by different lengths of guide rna. Proceedings of the National Academy of Sciences, 117:28576-28578, Oct 2020. URL: https://doi.org/10.1073/pnas.2015026117, doi:10.1073/pnas.2015026117. This article has 67 citations and is from a highest quality peer-reviewed journal.

  3. (narasimhan2025partnersinsilencing pages 1-2): Srinaath Narasimhan and Stefan J. Erkeland. Partners in silencing: decoding the mammalian argonaute interactome. Non-Coding RNA, 11:62, Aug 2025. URL: https://doi.org/10.3390/ncrna11040062, doi:10.3390/ncrna11040062. This article has 4 citations.

  4. (martinmerchan2023domainorganizationexpression pages 5-6): Andrea Martín-Merchán, Belen Moro, Antoine Bouet, and Nicolas G Bologna. Domain organization, expression, subcellular localization, and biological roles of argonaute proteins in arabidopsis thaliana. Journal of experimental botany, 74:2374-2388, Feb 2023. URL: https://doi.org/10.1093/jxb/erad030, doi:10.1093/jxb/erad030. This article has 28 citations and is from a domain leading peer-reviewed journal.

  5. (park2017humanargonaute3has pages 7-8): Mi Seul Park, Hong-Duc Phan, Florian Busch, Samantha H. Hinckley, James A. Brackbill, Vicki H. Wysocki, and Kotaro Nakanishi. Human argonaute3 has slicer activity. Nucleic Acids Research, 45:11867-11877, Oct 2017. URL: https://doi.org/10.1093/nar/gkx916, doi:10.1093/nar/gkx916. This article has 148 citations and is from a highest quality peer-reviewed journal.

  6. (park2017humanargonaute3has pages 1-2): Mi Seul Park, Hong-Duc Phan, Florian Busch, Samantha H. Hinckley, James A. Brackbill, Vicki H. Wysocki, and Kotaro Nakanishi. Human argonaute3 has slicer activity. Nucleic Acids Research, 45:11867-11877, Oct 2017. URL: https://doi.org/10.1093/nar/gkx916, doi:10.1093/nar/gkx916. This article has 148 citations and is from a highest quality peer-reviewed journal.

  7. (park2017humanargonaute3has media c985c1a4): Mi Seul Park, Hong-Duc Phan, Florian Busch, Samantha H. Hinckley, James A. Brackbill, Vicki H. Wysocki, and Kotaro Nakanishi. Human argonaute3 has slicer activity. Nucleic Acids Research, 45:11867-11877, Oct 2017. URL: https://doi.org/10.1093/nar/gkx916, doi:10.1093/nar/gkx916. This article has 148 citations and is from a highest quality peer-reviewed journal.

  8. (nakanishi2024whenargonautetakes pages 9-11): Kotaro Nakanishi. When argonaute takes out the ribonuclease sword. Journal of Biological Chemistry, 300:105499, Jan 2024. URL: https://doi.org/10.1016/j.jbc.2023.105499, doi:10.1016/j.jbc.2023.105499. This article has 25 citations and is from a domain leading peer-reviewed journal.

  9. (nakanishi2024whenargonautetakes pages 1-3): Kotaro Nakanishi. When argonaute takes out the ribonuclease sword. Journal of Biological Chemistry, 300:105499, Jan 2024. URL: https://doi.org/10.1016/j.jbc.2023.105499, doi:10.1016/j.jbc.2023.105499. This article has 25 citations and is from a domain leading peer-reviewed journal.

  10. (narasimhan2025partnersinsilencing pages 4-5): Srinaath Narasimhan and Stefan J. Erkeland. Partners in silencing: decoding the mammalian argonaute interactome. Non-Coding RNA, 11:62, Aug 2025. URL: https://doi.org/10.3390/ncrna11040062, doi:10.3390/ncrna11040062. This article has 4 citations.

  11. (narasimhan2025partnersinsilencing pages 10-11): Srinaath Narasimhan and Stefan J. Erkeland. Partners in silencing: decoding the mammalian argonaute interactome. Non-Coding RNA, 11:62, Aug 2025. URL: https://doi.org/10.3390/ncrna11040062, doi:10.3390/ncrna11040062. This article has 4 citations.

  12. (chen2026decodingargonautespecificity pages 7-10): Shihui Chen and C. Phillips. Decoding argonaute specificity: insights from c. elegans and beyond. RNA, 32:290-310, Dec 2026. URL: https://doi.org/10.1261/rna.080816.125, doi:10.1261/rna.080816.125. This article has 1 citations and is from a domain leading peer-reviewed journal.

  13. (moadab2023argonautevaultand pages 1-3): Fatemeh Moadab, Xiaoxing Wang, Rayan Najjar, Kennedy C. Ukadike, Shaohui Hu, Tyler Hulett, Anders A. Bengtsson, Christian Lood, and Tomas Mustelin. Argonaute, vault, and ribosomal proteins targeted by autoantibodies in systemic lupus erythematosus. The Journal of Rheumatology, 50:1136-1144, May 2023. URL: https://doi.org/10.3899/jrheum.2022-1327, doi:10.3899/jrheum.2022-1327. This article has 9 citations.

  14. (moadab2023argonautevaultand pages 3-4): Fatemeh Moadab, Xiaoxing Wang, Rayan Najjar, Kennedy C. Ukadike, Shaohui Hu, Tyler Hulett, Anders A. Bengtsson, Christian Lood, and Tomas Mustelin. Argonaute, vault, and ribosomal proteins targeted by autoantibodies in systemic lupus erythematosus. The Journal of Rheumatology, 50:1136-1144, May 2023. URL: https://doi.org/10.3899/jrheum.2022-1327, doi:10.3899/jrheum.2022-1327. This article has 9 citations.

  15. (wang2024argonaute2autoantibodiesa pages 2-3): Yixuan Wang, Yue Hu, Jiaqi Li, Huailu Ma, Zongqi Shi, Chaojing Wen, Yu Long, Ziwei Li, Hang Sun, Yixuan Yang, and Xiaofeng Shi. Argonaute-2 autoantibodies: a promising biomarker for predicting mortality in hbv-related acute-on-chronic liver failure patients with cirrhosis. Frontiers in Cellular and Infection Microbiology, Jul 2024. URL: https://doi.org/10.3389/fcimb.2024.1407064, doi:10.3389/fcimb.2024.1407064. This article has 3 citations.

  16. (wang2024argonaute2autoantibodiesa pages 3-6): Yixuan Wang, Yue Hu, Jiaqi Li, Huailu Ma, Zongqi Shi, Chaojing Wen, Yu Long, Ziwei Li, Hang Sun, Yixuan Yang, and Xiaofeng Shi. Argonaute-2 autoantibodies: a promising biomarker for predicting mortality in hbv-related acute-on-chronic liver failure patients with cirrhosis. Frontiers in Cellular and Infection Microbiology, Jul 2024. URL: https://doi.org/10.3389/fcimb.2024.1407064, doi:10.3389/fcimb.2024.1407064. This article has 3 citations.

  17. (moadab2023argonautevaultand pages 9-15): Fatemeh Moadab, Xiaoxing Wang, Rayan Najjar, Kennedy C. Ukadike, Shaohui Hu, Tyler Hulett, Anders A. Bengtsson, Christian Lood, and Tomas Mustelin. Argonaute, vault, and ribosomal proteins targeted by autoantibodies in systemic lupus erythematosus. The Journal of Rheumatology, 50:1136-1144, May 2023. URL: https://doi.org/10.3899/jrheum.2022-1327, doi:10.3899/jrheum.2022-1327. This article has 9 citations.

  18. (wang2024argonaute2autoantibodiesa pages 9-10): Yixuan Wang, Yue Hu, Jiaqi Li, Huailu Ma, Zongqi Shi, Chaojing Wen, Yu Long, Ziwei Li, Hang Sun, Yixuan Yang, and Xiaofeng Shi. Argonaute-2 autoantibodies: a promising biomarker for predicting mortality in hbv-related acute-on-chronic liver failure patients with cirrhosis. Frontiers in Cellular and Infection Microbiology, Jul 2024. URL: https://doi.org/10.3389/fcimb.2024.1407064, doi:10.3389/fcimb.2024.1407064. This article has 3 citations.

  19. (park2020humanargonaute2and pages 2-3): Mi Seul Park, GeunYoung Sim, Audrey C. Kehling, and Kotaro Nakanishi. Human argonaute2 and argonaute3 are catalytically activated by different lengths of guide rna. Proceedings of the National Academy of Sciences, 117:28576-28578, Oct 2020. URL: https://doi.org/10.1073/pnas.2015026117, doi:10.1073/pnas.2015026117. This article has 67 citations and is from a highest quality peer-reviewed journal.

  20. (park2017humanargonaute3has pages 4-5): Mi Seul Park, Hong-Duc Phan, Florian Busch, Samantha H. Hinckley, James A. Brackbill, Vicki H. Wysocki, and Kotaro Nakanishi. Human argonaute3 has slicer activity. Nucleic Acids Research, 45:11867-11877, Oct 2017. URL: https://doi.org/10.1093/nar/gkx916, doi:10.1093/nar/gkx916. This article has 148 citations and is from a highest quality peer-reviewed journal.

  21. (wang2024argonaute2autoantibodiesa pages 1-2): Yixuan Wang, Yue Hu, Jiaqi Li, Huailu Ma, Zongqi Shi, Chaojing Wen, Yu Long, Ziwei Li, Hang Sun, Yixuan Yang, and Xiaofeng Shi. Argonaute-2 autoantibodies: a promising biomarker for predicting mortality in hbv-related acute-on-chronic liver failure patients with cirrhosis. Frontiers in Cellular and Infection Microbiology, Jul 2024. URL: https://doi.org/10.3389/fcimb.2024.1407064, doi:10.3389/fcimb.2024.1407064. This article has 3 citations.

Citations

  1. narasimhan2025partnersinsilencing pages 1-2
  2. nakanishi2024whenargonautetakes pages 9-11
  3. nakanishi2024whenargonautetakes pages 1-3
  4. narasimhan2025partnersinsilencing pages 10-11
  5. chen2026decodingargonautespecificity pages 7-10
  6. moadab2023argonautevaultand pages 1-3
  7. martinmerchan2023domainorganizationexpression pages 5-6
  8. narasimhan2025partnersinsilencing pages 4-5
  9. moadab2023argonautevaultand pages 3-4
  10. moadab2023argonautevaultand pages 9-15
  11. https://doi.org/10.1073/pnas.2015026117;
  12. https://doi.org/10.1093/nar/gkx916;
  13. https://doi.org/10.3899/jrheum.2022-1327;
  14. https://doi.org/10.3389/fcimb.2024.1407064;
  15. https://doi.org/10.1093/jxb/erad030
  16. https://doi.org/10.3390/ncrna11040062
  17. https://doi.org/10.1073/pnas.2015026117
  18. https://doi.org/10.1093/nar/gkx916
  19. https://doi.org/10.3390/ncrna11040062;
  20. https://doi.org/10.1261/rna.080816.125
  21. https://doi.org/10.1016/j.jbc.2023.105499
  22. https://doi.org/10.3899/jrheum.2022-1327
  23. https://doi.org/10.3389/fcimb.2024.1407064
  24. https://doi.org/10.1093/nar/gkx916,
  25. https://doi.org/10.1073/pnas.2015026117,
  26. https://doi.org/10.3390/ncrna11040062,
  27. https://doi.org/10.1093/jxb/erad030,
  28. https://doi.org/10.1016/j.jbc.2023.105499,
  29. https://doi.org/10.1261/rna.080816.125,
  30. https://doi.org/10.3899/jrheum.2022-1327,
  31. https://doi.org/10.3389/fcimb.2024.1407064,

📄 View Raw YAML

id: Q9H9G7
gene_symbol: AGO3
product_type: PROTEIN
status: COMPLETE
taxon:
  id: NCBITaxon:9606
  label: Homo sapiens
description: 'AGO3 encodes human Argonaute-3, a small-RNA-binding RISC effector with a context-restricted
  slicer activity. Its core function is Argonaute-mediated small-RNA-guided gene silencing, with biochemical
  evidence for guide- and target-context-dependent RNA endonuclease activity and a non-core nuclear splicing
  role reported in immune cells.'
alternative_products:
- name: '1'
  id: Q9H9G7-1
- name: '2'
  id: Q9H9G7-2
  sequence_note: VSP_041084
existing_annotations:
- term:
    id: GO:0005634
    label: nucleus
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  review:
    summary: nucleus is a plausible or reported location/context but is not the core functional site
      annotation for AGO3.
    action: KEEP_AS_NON_CORE
    reason: The core location is cytoplasmic RISC/RNP granules; nuclear, membrane, synaptic,
      exosomal, or other compartment annotations should remain non-core unless tied to a specific
      curated mechanism.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005737
    label: cytoplasm
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  review:
    summary: cytoplasm is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0004521
    label: RNA endonuclease activity
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  review:
    summary: RNA endonuclease activity is supported for AGO3 as a conditional, guide-dependent
      slicer activity.
    action: ACCEPT
    reason: AGO3 is not the canonical slicer like AGO2, but biochemical studies support guide- and
      target-context-dependent RNA cleavage.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035194
    label: regulatory ncRNA-mediated post-transcriptional gene silencing
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  review:
    summary: regulatory ncRNA-mediated post-transcriptional gene silencing is consistent with AGO3
      as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0016442
    label: RISC complex
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  review:
    summary: RISC complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0036464
    label: cytoplasmic ribonucleoprotein granule
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  review:
    summary: cytoplasmic ribonucleoprotein granule is consistent with AGO3 as a small-RNA-loaded
      Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0031054
    label: pre-miRNA processing
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  review:
    summary: pre-miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
    action: MARK_AS_OVER_ANNOTATED
    reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing,
      not as the Drosha/Dicer processing enzyme for pre-miRNAs.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035198
    label: miRNA binding
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  review:
    summary: miRNA binding is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0003727
    label: single-stranded RNA binding
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  review:
    summary: single-stranded RNA binding is too generic for AGO3.
    action: MARK_AS_OVER_ANNOTATED
    reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC
      rather than a broad nucleic-acid or RNA-binding label.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0000932
    label: P-body
  evidence_type: IEA
  original_reference_id: GO_REF:0000044
  review:
    summary: P-body is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0003676
    label: nucleic acid binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000002
  review:
    summary: nucleic acid binding is too generic for AGO3.
    action: MARK_AS_OVER_ANNOTATED
    reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC
      rather than a broad nucleic-acid or RNA-binding label.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0003723
    label: RNA binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  review:
    summary: RNA binding is too generic for AGO3.
    action: MARK_AS_OVER_ANNOTATED
    reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC
      rather than a broad nucleic-acid or RNA-binding label.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0003725
    label: double-stranded RNA binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  review:
    summary: double-stranded RNA binding is too generic for AGO3.
    action: MARK_AS_OVER_ANNOTATED
    reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC
      rather than a broad nucleic-acid or RNA-binding label.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0003727
    label: single-stranded RNA binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  review:
    summary: single-stranded RNA binding is too generic for AGO3.
    action: MARK_AS_OVER_ANNOTATED
    reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC
      rather than a broad nucleic-acid or RNA-binding label.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0016442
    label: RISC complex
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  review:
    summary: RISC complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0031047
    label: regulatory ncRNA-mediated gene silencing
  evidence_type: IEA
  original_reference_id: GO_REF:0000002
  review:
    summary: regulatory ncRNA-mediated gene silencing is consistent with AGO3 as a small-RNA-loaded
      Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0031054
    label: pre-miRNA processing
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  review:
    summary: pre-miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
    action: MARK_AS_OVER_ANNOTATED
    reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing,
      not as the Drosha/Dicer processing enzyme for pre-miRNAs.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035198
    label: miRNA binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  review:
    summary: miRNA binding is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035278
    label: miRNA-mediated gene silencing by inhibition of translation
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  review:
    summary: miRNA-mediated gene silencing by inhibition of translation is consistent with AGO3 as a
      small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035279
    label: miRNA-mediated gene silencing by mRNA destabilization
  evidence_type: IEA
  original_reference_id: GO_REF:0000108
  review:
    summary: miRNA-mediated gene silencing by mRNA destabilization is plausible for AGO3 only as a
      context-dependent non-core process, not as an unqualified core AGO3 function.
    action: KEEP_AS_NON_CORE
    reason: AGO3 is primarily a small-RNA-binding RISC effector, and later biochemical work supports
      guide- and target-dependent slicing. Endogenous AGO3 cleavage targets remain unclear, so this
      broad IEA process should not be treated as a core positive annotation.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: |
        Importantly, this review emphasizes a major open question: **endogenous physiological target RNAs cleaved by AGO3 remain unknown**.
- term:
    id: GO:0036464
    label: cytoplasmic ribonucleoprotein granule
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  review:
    summary: cytoplasmic ribonucleoprotein granule is consistent with AGO3 as a small-RNA-loaded
      Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0070578
    label: RISC-loading complex
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  review:
    summary: RISC-loading complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC
      effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0070922
    label: RISC complex assembly
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  review:
    summary: RISC complex assembly is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC
      effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0072091
    label: regulation of stem cell proliferation
  evidence_type: IEA
  original_reference_id: GO_REF:0000002
  review:
    summary: regulation of stem cell proliferation is downstream or context-specific for AGO3.
    action: KEEP_AS_NON_CORE
    reason: This process may be supported in a particular experiment or pathway model, but it is
      peripheral to the core Argonaute/RISC molecular role.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:19167051
  review:
    summary: Generic protein binding does not capture the specific Argonaute/RISC function.
    action: MARK_AS_OVER_ANNOTATED
    reason: The cited interactions are best interpreted as RISC/cofactor context for
      small-RNA-guided silencing, not as a standalone core function.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:19324964
  review:
    summary: Generic protein binding does not capture the specific Argonaute/RISC function.
    action: MARK_AS_OVER_ANNOTATED
    reason: The cited interactions are best interpreted as RISC/cofactor context for
      small-RNA-guided silencing, not as a standalone core function.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:19383768
  review:
    summary: Generic protein binding does not capture the specific Argonaute/RISC function.
    action: MARK_AS_OVER_ANNOTATED
    reason: The cited interactions are best interpreted as RISC/cofactor context for
      small-RNA-guided silencing, not as a standalone core function.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:23064648
  review:
    summary: Generic protein binding does not capture the specific Argonaute/RISC function.
    action: MARK_AS_OVER_ANNOTATED
    reason: The cited interactions are best interpreted as RISC/cofactor context for
      small-RNA-guided silencing, not as a standalone core function.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:28330616
  review:
    summary: Generic protein binding does not capture the specific Argonaute/RISC function.
    action: MARK_AS_OVER_ANNOTATED
    reason: The cited interactions are best interpreted as RISC/cofactor context for
      small-RNA-guided silencing, not as a standalone core function.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:28683311
  review:
    summary: Generic protein binding does not capture the specific Argonaute/RISC function.
    action: MARK_AS_OVER_ANNOTATED
    reason: The cited interactions are best interpreted as RISC/cofactor context for
      small-RNA-guided silencing, not as a standalone core function.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0000794
    label: condensed nuclear chromosome
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  review:
    summary: condensed nuclear chromosome is a plausible or reported location/context but is not the
      core functional site annotation for AGO3.
    action: KEEP_AS_NON_CORE
    reason: The core location is cytoplasmic RISC/RNP granules; nuclear, membrane, synaptic,
      exosomal, or other compartment annotations should remain non-core unless tied to a specific
      curated mechanism.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0010628
    label: positive regulation of gene expression
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  review:
    summary: positive regulation of gene expression reflects a reported nuclear or regulatory
      context rather than the main conserved AGO3 function.
    action: KEEP_AS_NON_CORE
    reason: Argonaute paralogs have reported nuclear/regulatory interactions, but the dominant
      conserved function remains small-RNA-guided post-transcriptional repression.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:1901224
    label: positive regulation of non-canonical NF-kappaB signal transduction
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  review:
    summary: positive regulation of non-canonical NF-kappaB signal transduction reflects a reported
      nuclear or regulatory context rather than the main conserved AGO3 function.
    action: KEEP_AS_NON_CORE
    reason: Argonaute paralogs have reported nuclear/regulatory interactions, but the dominant
      conserved function remains small-RNA-guided post-transcriptional repression.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035196
    label: miRNA processing
  evidence_type: NAS
  original_reference_id: PMID:28781232
  review:
    summary: miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
    action: MARK_AS_OVER_ANNOTATED
    reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing,
      not as the Drosha/Dicer processing enzyme for pre-miRNAs.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0036464
    label: cytoplasmic ribonucleoprotein granule
  evidence_type: IDA
  original_reference_id: GO_REF:0000052
  review:
    summary: cytoplasmic ribonucleoprotein granule is consistent with AGO3 as a small-RNA-loaded
      Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0004521
    label: RNA endonuclease activity
  evidence_type: IMP
  original_reference_id: PMID:29040713
  review:
    summary: RNA endonuclease activity is supported for AGO3 as a conditional, guide-dependent
      slicer activity.
    action: ACCEPT
    reason: AGO3 is not the canonical slicer like AGO2, but biochemical studies support guide- and
      target-context-dependent RNA cleavage.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0090624
    label: endoribonuclease activity, cleaving miRNA-paired mRNA
  evidence_type: IMP
  original_reference_id: PMID:29040713
  review:
    summary: endoribonuclease activity, cleaving miRNA-paired mRNA is supported for AGO3 as a
      conditional, guide-dependent slicer activity.
    action: ACCEPT
    reason: AGO3 is not the canonical slicer like AGO2, but biochemical studies support guide- and
      target-context-dependent RNA cleavage.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035196
    label: miRNA processing
  evidence_type: IMP
  original_reference_id: PMID:22795694
  review:
    summary: miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
    action: MARK_AS_OVER_ANNOTATED
    reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing,
      not as the Drosha/Dicer processing enzyme for pre-miRNAs.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0031054
    label: pre-miRNA processing
  evidence_type: IDA
  original_reference_id: PMID:19966796
  review:
    summary: pre-miRNA processing overstates the direct role of AGO3 in canonical miRNA biogenesis.
    action: MARK_AS_OVER_ANNOTATED
    reason: The better-supported role is as a loaded Argonaute effector after small-RNA processing,
      not as the Drosha/Dicer processing enzyme for pre-miRNAs.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0003725
    label: double-stranded RNA binding
  evidence_type: IDA
  original_reference_id: PMID:19966796
  review:
    summary: double-stranded RNA binding is too generic for AGO3.
    action: MARK_AS_OVER_ANNOTATED
    reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC
      rather than a broad nucleic-acid or RNA-binding label.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0003727
    label: single-stranded RNA binding
  evidence_type: IDA
  original_reference_id: PMID:19966796
  review:
    summary: single-stranded RNA binding is too generic for AGO3.
    action: MARK_AS_OVER_ANNOTATED
    reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC
      rather than a broad nucleic-acid or RNA-binding label.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0016442
    label: RISC complex
  evidence_type: IDA
  original_reference_id: PMID:19966796
  review:
    summary: RISC complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0070578
    label: RISC-loading complex
  evidence_type: IDA
  original_reference_id: PMID:19966796
  review:
    summary: RISC-loading complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC
      effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0070922
    label: RISC complex assembly
  evidence_type: IDA
  original_reference_id: PMID:19966796
  review:
    summary: RISC complex assembly is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC
      effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0090625
    label: siRNA-mediated gene silencing by mRNA destabilization
  evidence_type: IDA
  original_reference_id: PMID:15260970
  negated: true
  review:
    summary: This NOT annotation is appropriate for the PMID:15260970 assay context; AGO3 did not
      behave as the AGO2-like siRNA-guided target-cleavage Argonaute in that study.
    action: ACCEPT
    reason: PMID:15260970 supports AGO3 association with miRNAs/RISC but reports target-cleavage
      activity only for AGO2 in the tested assays. Later work supports conditional AGO3 slicing, so
      this NOT call should be treated as scoped to the 2004 AGO2-specific siRNA-cleavage context.
    supported_by:
    - reference_id: PMID:15260970
      supporting_text: Purification of the FLAG/HA-epitope-tagged Ago containing complexes from
        different human cell lines revealed that endonuclease activity is exclusively associated
        with Ago2.
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Historically, mammalian **AGO2** was regarded as the only catalytically
        competent slicer, while AGO1/AGO3/AGO4 were categorized as non-slicer AGOs for most
        miRNA-mediated repression.
- term:
    id: GO:0005737
    label: cytoplasm
  evidence_type: IDA
  original_reference_id: PMID:15260970
  review:
    summary: cytoplasm is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0016442
    label: RISC complex
  evidence_type: IDA
  original_reference_id: PMID:15260970
  review:
    summary: RISC complex is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035198
    label: miRNA binding
  evidence_type: IDA
  original_reference_id: PMID:15260970
  review:
    summary: miRNA binding is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035279
    label: miRNA-mediated gene silencing by mRNA destabilization
  evidence_type: IDA
  original_reference_id: PMID:15260970
  negated: true
  review:
    summary: This older NOT annotation should be removed because later AGO3-specific evidence supports
      guide-dependent miRNA-paired target cleavage.
    action: REMOVE
    reason: PMID:15260970 did not detect AGO3-dependent target cleavage in the tested 2004 assay, but
      PMID:29040713 supersedes that broad negative by demonstrating guide- and target-dependent AGO3
      slicer activity.
    supported_by:
    - reference_id: PMID:15260970
      supporting_text: The specific role of Ago2 in guiding target RNA cleavage was confirmed
        independently by siRNA-based depletion of individual Ago members in combination with a
        sensitive positive-readout reporter assay.
    - reference_id: PMID:29040713
      supporting_text: recombinant AGO3 loaded with miR-20a cleaves complementary target RNAs, whereas
        AGO3 loaded with let-7a, miR-19b or miR-16 does not, indicating that AGO3 has slicer activity
        but that this activity depends on the guide RNA.
- term:
    id: GO:0016020
    label: membrane
  evidence_type: HDA
  original_reference_id: PMID:19946888
  review:
    summary: membrane is a plausible or reported location/context but is not the core functional
      site annotation for AGO3.
    action: KEEP_AS_NON_CORE
    reason: The core location is cytoplasmic RISC/RNP granules; nuclear, membrane, synaptic,
      exosomal, or other compartment annotations should remain non-core unless tied to a specific
      curated mechanism.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0003723
    label: RNA binding
  evidence_type: HDA
  original_reference_id: PMID:22658674
  review:
    summary: RNA binding is too generic for AGO3.
    action: MARK_AS_OVER_ANNOTATED
    reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC
      rather than a broad nucleic-acid or RNA-binding label.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0003723
    label: RNA binding
  evidence_type: HDA
  original_reference_id: PMID:22681889
  review:
    summary: RNA binding is too generic for AGO3.
    action: MARK_AS_OVER_ANNOTATED
    reason: The informative molecular function is Argonaute small-RNA/miRNA binding within RISC
      rather than a broad nucleic-acid or RNA-binding label.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-1606561
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-1606682
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-1912362
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-1912363
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-1912364
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-1912366
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-1912367
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-1912368
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-1912406
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-2318752
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-3209151
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-426489
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-426522
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-4518575
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-5687103
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8935766
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8937097
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8937134
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8938440
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8938487
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8938507
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8944650
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8944684
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8944706
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9011958
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9012203
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9012208
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9038545
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9618392
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9618486
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9624925
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9657791
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9858289
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9858309
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9924921
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9925095
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9925158
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9925159
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005829
    label: cytosol
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-9925324
  review:
    summary: cytosol is consistent with AGO3 as a small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0072091
    label: regulation of stem cell proliferation
  evidence_type: IMP
  original_reference_id: PMID:23064648
  review:
    summary: regulation of stem cell proliferation is downstream or context-specific for AGO3.
    action: KEEP_AS_NON_CORE
    reason: This process may be supported in a particular experiment or pathway model, but it is
      peripheral to the core Argonaute/RISC molecular role.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:22915799
  review:
    summary: Generic protein binding does not capture the specific Argonaute/RISC function.
    action: MARK_AS_OVER_ANNOTATED
    reason: The cited interactions are best interpreted as RISC/cofactor context for
      small-RNA-guided silencing, not as a standalone core function.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0006402
    label: mRNA catabolic process
  evidence_type: IDA
  original_reference_id: PMID:18771919
  review:
    summary: mRNA catabolic process is downstream or context-specific for AGO3.
    action: KEEP_AS_NON_CORE
    reason: This process may be supported in a particular experiment or pathway model, but it is
      peripheral to the core Argonaute/RISC molecular role.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
- term:
    id: GO:0035278
    label: miRNA-mediated gene silencing by inhibition of translation
  evidence_type: IDA
  original_reference_id: PMID:18771919
  review:
    summary: miRNA-mediated gene silencing by inhibition of translation is consistent with AGO3 as a
      small-RNA-loaded Argonaute/RISC effector.
    action: ACCEPT
    reason: The falcon report supports AGO3 function in miRISC-mediated small-RNA-guided repression
      and related RISC complexes or cytoplasmic RNP compartments.
    supported_by:
    - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
      supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
        Argonaute effector**
references:
- id: GO_REF:0000002
  title: Gene Ontology annotation through association of InterPro records with GO terms
  findings: []
- id: GO_REF:0000033
  title: Annotation inferences using phylogenetic trees
  findings: []
- id: GO_REF:0000044
  title: Gene Ontology annotation based on UniProtKB/Swiss-Prot Subcellular Location vocabulary
    mapping, accompanied by conservative changes to GO terms applied by UniProt
  findings: []
- id: GO_REF:0000052
  title: Gene Ontology annotation based on curation of immunofluorescence data
  findings: []
- id: GO_REF:0000107
  title: Automatic transfer of experimentally verified manual GO annotation data to orthologs using
    Ensembl Compara
  findings: []
- id: GO_REF:0000108
  title: Automatic assignment of GO terms using logical inference, based on on inter-ontology links
  findings: []
- id: GO_REF:0000117
  title: Electronic Gene Ontology annotations created by ARBA machine learning models
  findings: []
- id: GO_REF:0000120
  title: Combined Automated Annotation using Multiple IEA Methods
  findings: []
- id: PMID:15260970
  title: Human Argonaute2 mediates RNA cleavage targeted by miRNAs and siRNAs.
  findings: []
- id: PMID:18771919
  title: Importance of translation and nonnucleolytic ago proteins for on-target RNA interference.
  findings: []
- id: PMID:19167051
  title: Importin 8 is a gene silencing factor that targets argonaute proteins to distinct mRNAs.
  findings: []
- id: PMID:19324964
  title: The C-terminal half of human Ago2 binds to multiple GW-rich regions of GW182 and requires
    GW182 to mediate silencing.
  findings: []
- id: PMID:19383768
  title: The C-terminal domains of human TNRC6A, TNRC6B, and TNRC6C silence bound transcripts
    independently of Argonaute proteins.
  findings: []
- id: PMID:19946888
  title: Defining the membrane proteome of NK cells.
  findings: []
- id: PMID:19966796
  title: ATP-dependent human RISC assembly pathways.
  findings: []
- id: PMID:22658674
  title: Insights into RNA biology from an atlas of mammalian mRNA-binding proteins.
  findings: []
- id: PMID:22681889
  title: The mRNA-bound proteome and its global occupancy profile on protein-coding transcripts.
  findings: []
- id: PMID:22795694
  title: Slicing-independent RISC activation requires the argonaute PAZ domain.
  findings: []
- id: PMID:22915799
  title: HIV-1 replication and APOBEC3 antiviral activity are not regulated by P bodies.
  findings: []
- id: PMID:23064648
  title: DICER- and AGO3-dependent generation of retinoic acid-induced DR2 Alu RNAs regulates human
    stem cell proliferation.
  findings: []
- id: PMID:28330616
  title: Systematic Analysis of Human Protein Phosphatase Interactions and Dynamics.
  findings: []
- id: PMID:28683311
  title: Argonaute Utilization for miRNA Silencing Is Determined by Phosphorylation-Dependent
    Recruitment of LIM-Domain-Containing Proteins.
  findings: []
- id: PMID:28781232
  title: Multivalent Recruitment of Human Argonaute by GW182.
  findings: []
- id: PMID:29040713
  title: Human Argonaute3 has slicer activity.
  findings: []
- id: Reactome:R-HSA-1606561
  title: MIR449 microRNAs bind 3'UTR of NOTCH1 mRNA
  findings: []
- id: Reactome:R-HSA-1606682
  title: MIR34 microRNAs bind 3'UTR of NOTCH1 mRNA
  findings: []
- id: Reactome:R-HSA-1912362
  title: MIR150 microRNA binds 3'UTR of NOTCH3 mRNA
  findings: []
- id: Reactome:R-HSA-1912363
  title: MIR200B/C microRNAs bind NOTCH1 mRNA
  findings: []
- id: Reactome:R-HSA-1912364
  title: MIR181C microRNA binds 3'UTR of NOTCH4 mRNA
  findings: []
- id: Reactome:R-HSA-1912366
  title: MIR206 microRNA binds 3'UTR of NOTCH3 mRNA
  findings: []
- id: Reactome:R-HSA-1912367
  title: MIR34 microRNAs bind 3'UTR of NOTCH2 mRNA
  findings: []
- id: Reactome:R-HSA-1912368
  title: MIR302A microRNA binds 3'UTR of NOTCH4 mRNA
  findings: []
- id: Reactome:R-HSA-1912406
  title: p53 positively regulates transcription of MIR34 microRNAs
  findings: []
- id: Reactome:R-HSA-2318752
  title: miR-26A microRNAs bind PTEN mRNA
  findings: []
- id: Reactome:R-HSA-3209151
  title: miR-24 binds p16INK4A and p14ARF mRNAs
  findings: []
- id: Reactome:R-HSA-426489
  title: RISC binds inexactly matching target RNAs
  findings: []
- id: Reactome:R-HSA-426522
  title: Nonendonucleolytic RISC binds exactly matching target RNAs
  findings: []
- id: Reactome:R-HSA-4518575
  title: miR-92b binds 3'UTR of NLK mRNA
  findings: []
- id: Reactome:R-HSA-5687103
  title: FOXO3 regulates MIR34B,C expression
  findings: []
- id: Reactome:R-HSA-8935766
  title: miR-378 binds RUNX1 mRNA
  findings: []
- id: Reactome:R-HSA-8937097
  title: MIR27A gene transcription is stimulated by the complex containing RUNX1, PRMT1 and GATA1
    and inhibited by the complex of RUNX1, SIN3A and PRMT6
  findings: []
- id: Reactome:R-HSA-8937134
  title: miR-27a binds RUNX1 mRNA
  findings: []
- id: Reactome:R-HSA-8938440
  title: miR-17 binds RUNX1 mRNA
  findings: []
- id: Reactome:R-HSA-8938487
  title: miR-20a binds RUNX1 mRNA
  findings: []
- id: Reactome:R-HSA-8938507
  title: miR-106a binds RUNX1 mRNA
  findings: []
- id: Reactome:R-HSA-8944650
  title: miR-214 microRNA binds PTEN mRNA
  findings: []
- id: Reactome:R-HSA-8944684
  title: miR-205 microRNA binds PTEN mRNA
  findings: []
- id: Reactome:R-HSA-8944706
  title: miR-21 nonendonucleolytic RISC binds PTEN mRNA
  findings: []
- id: Reactome:R-HSA-9011958
  title: miR-26A and B bind to the 3'UTR of the GREB1 mRNA
  findings: []
- id: Reactome:R-HSA-9012203
  title: miR-26A and B bind to the 3'UTR of the CHD11 mRNA
  findings: []
- id: Reactome:R-HSA-9012208
  title: miR-26A and B bind to the 3'UTR of the KPNA2 mRNA
  findings: []
- id: Reactome:R-HSA-9038545
  title: miR-613 binds to the 3'UTR of the NR1H3 mRNA
  findings: []
- id: Reactome:R-HSA-9618392
  title: miR-26 binds to the 3'UTR of the ARL4C mRNA
  findings: []
- id: Reactome:R-HSA-9618486
  title: miR-26 binds to the 3'UTR of the ABCA1 mRNA
  findings: []
- id: Reactome:R-HSA-9624925
  title: miR-33 binds to the 3'UTR of the ABCA1 mRNA
  findings: []
- id: Reactome:R-HSA-9657791
  title: miR-144 binds to the 3'UTR of the ABCA1 mRNA
  findings: []
- id: Reactome:R-HSA-9858289
  title: MITF-M-dependent miR-211 expression
  findings: []
- id: Reactome:R-HSA-9858309
  title: miR-211 RISC binds POU3F2 mRNA
  findings: []
- id: Reactome:R-HSA-9924921
  title: CD274 mRNA binds miR-34 RISC
  findings: []
- id: Reactome:R-HSA-9925095
  title: CD274 mRNA binds miR-340 RISC
  findings: []
- id: Reactome:R-HSA-9925158
  title: CD274 mRNA binds miR-152 RISC
  findings: []
- id: Reactome:R-HSA-9925159
  title: CD274 mRNA binds miR-140 RISC
  findings: []
- id: Reactome:R-HSA-9925324
  title: CD274 mRNA binds miR-200B/C RISC
  findings: []
- id: file:human/AGO3/AGO3-uniprot.txt
  title: UniProt text export for AGO3
  findings:
  - statement: UniProt identifies AGO3 and its protein family/domain context.
    supporting_text: Belongs to the argonaute family. Ago subfamily.
- id: file:human/AGO3/AGO3-deep-research-falcon.md
  title: Falcon deep research report for AGO3
  findings:
  - statement: Falcon research supports the reviewed core function of AGO3.
    supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
      Argonaute effector**
core_functions:
- description: Small-RNA-guided Argonaute/RISC effector function in post-transcriptional gene
    silencing.
  molecular_function:
    id: GO:0035198
    label: miRNA binding
  directly_involved_in:
  - id: GO:0035194
    label: regulatory ncRNA-mediated post-transcriptional gene silencing
  - id: GO:0035278
    label: miRNA-mediated gene silencing by inhibition of translation
  in_complex:
    id: GO:0016442
    label: RISC complex
  locations:
  - id: GO:0005737
    label: cytoplasm
  supported_by:
  - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
    supporting_text: Human **AGO3 (EIF2C3; Q9H9G7)** is best annotated as a **small-RNA-binding
      Argonaute effector**
- description: Conditional RNA-guided endonuclease activity that depends on guide length/sequence
    and target context.
  molecular_function:
    id: GO:0090624
    label: endoribonuclease activity, cleaving miRNA-paired mRNA
  in_complex:
    id: GO:0016442
    label: RISC complex
  locations:
  - id: GO:0005829
    label: cytosol
  supported_by:
  - reference_id: file:human/AGO3/AGO3-deep-research-falcon.md
    supporting_text: AGO3 cleavage is **guide-sequence and guide-length dependent**. miR-20a
      activates AGO3, whereas let-7a, miR-19b, and miR-16 do not in the reported assays.
proposed_new_terms: []
suggested_questions:
- question: Which endogenous human tissues or stress states require AGO3-specific RISC activity
    rather than redundant activity from other AGO paralogs?
- question: Which protein cofactors determine guide loading, localization, and non-core nuclear or
    granule-associated functions of AGO3?
suggested_experiments:
- description: Endogenous tagging of AGO3 followed by small-RNA CLIP, target RNA profiling, and
    paralog-specific rescue in AGO-depleted human cells.
  experiment_type: genome editing/RNA-protein interaction profiling
  hypothesis: AGO3 has paralog-specific guide, target, and compartment preferences within human RISC
    biology.
- description: Biochemical reconstitution of AGO3-RISC with representative miRNA and siRNA guides to
    compare target repression, target cleavage, and TNRC6 recruitment.
  experiment_type: biochemical reconstitution
  hypothesis: AGO3 activity is best explained by guide-loaded RISC function rather than generic RNA
    or protein binding.