AP2M1

UniProt ID: Q96CW1
Organism: Homo sapiens
Review Status: COMPLETE
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Gene Description

AP2M1 encodes the medium ΞΌ2 subunit of the heterotetrameric AP-2 clathrin adaptor complex. Its C-terminal domain recognizes cytoplasmic sorting signals in membrane cargo, especially tyrosine-based motifs, while its N-terminal region helps assemble the adaptor core. AP-2 cycles between the cytosol and coats on the cytoplasmic face of the plasma membrane, coupling cargo selection to clathrin-dependent internalization. Membrane-dependent rearrangement and phosphorylation of ΞΌ2 Thr156 coordinate cargo capture and accessory-protein recruitment. This machinery supports receptor turnover and synaptic membrane-protein retrieval. Pathogenic AP2M1 variants can impair endocytosis and cause developmental and epileptic encephalopathy.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0002092 positive regulation of receptor internalization
IEA
GO_REF:0000107
ACCEPT
Summary: Promotes internalization by selecting cargo for AP-2 coated pits.
Reason: The rat-derived electronic assertion agrees with the conserved cargo-recognition role and human-cell depletion/rescue evidence. The retained regulatory process describes ΞΌ2-dependent uptake efficiency, not a separate kinase or an independently reconstructed rat experiment.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
PMID:17035630
Mutating the PtdIns(4,5)P2 binding site of alpha, the phosphorylation site of mu2, or the YXXPhi binding site of mu2 impairs AP-2 function, as assayed by transferrin uptake.
GO:0005048 signal sequence receptor activity
IDA
PMID:8918456
Sequence requirements for the recognition of tyrosine-based ...
ACCEPT
Summary: Recognizes tyrosine-based endocytic sorting signals.
Reason: The cited primary abstract reports ΞΌ2 recognition of Yxx-hydrophobic peptides with sequence specificity. This fits the current GO definition of a short localization signal, which is not restricted to ER signal peptides. Reagent species is not resolved by the abstract; human ΞΌ2 motif-recognition evidence corroborates the conserved activity.
Supporting Evidence:
PMID:8918456
Both mu 2 and AP-2 are shown to interact with various sequences of the form tyrosine-polar-polar-hydrophobic (YppΓΈ) found on receptors that follow the clathrin pathway.
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
GO:0005048 signal sequence receptor activity
IDA
PMID:9171339
Regulatory interactions in the recognition of endocytic sort...
ACCEPT
Summary: Recognizes cargo sorting signals through ΞΌ2.
Reason: The cited study explicitly analyzes ΞΌ2 interactions with endocytic sorting sequences. Preserve the source IDA annotation; the abstract establishes the recognition mechanism without resolving every reagent species. Human ΞΌ2 cargo-tail assays independently support the same established activity.
Supporting Evidence:
PMID:9171339
this binding is strengthened significantly when the AP-2 complex is present in clathrin coats
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
GO:0005048 signal sequence receptor activity
IEA
GO_REF:0000107
ACCEPT
Summary: Retain conserved ΞΌ2 sorting-signal recognition.
Reason: Rat-to-human transfer agrees with target-specific human ΞΌ2 cargo-tail and purified-domain experiments. The source transfer is preserved, while the exact donor experiment has not been reconstructed.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
PMID:33436498
a binding site for the clathrin adaptor AP2 ΞΌ2 in ACE2 overlaps with a phospho-dependent binding site for the SH2 domains of Src family tyrosine kinases.
GO:0005515 protein binding
IPI
PMID:12032142
Subunit H of the V-ATPase binds to the medium chain of adapt...
KEEP AS NON CORE
Summary: Retain the mapped interaction with the regulatory V-ATPase H subunit.
Reason: The source abstract explicitly maps binding between ΞΌ2 residues 1–145 and ATP6V1H residues 133–363. This is a supported accessory interaction; no sufficiently justified narrower function is assigned from the abstract alone. The regulatory H subunit is not itself the ATPase catalytic subunit, so its identity does not warrant ATPase binding or ATPase activity for ΞΌ2.
Supporting Evidence:
PMID:12032142
The interaction sites of V1H and mu2 were mapped to a central region in V1H from positions 133 to 363
GO:0005515 protein binding
IPI
PMID:16189514
Towards a proteome-scale map of the human protein-protein in...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P51116 remains unresolved.
Reason: The human interactome abstract describes a broad interaction map; the exact pair assay and target construct were not inspected.
GO:0005515 protein binding
IPI
PMID:16189514
Towards a proteome-scale map of the human protein-protein in...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P63010 remains unresolved.
Reason: The human interactome abstract describes a broad interaction map; the exact pair assay and target construct were not inspected. P63010 is the AP2B1 beta subunit; independent AP-2 architecture supports a plausible association but does not establish what this original experiment measured.
GO:0005515 protein binding
IPI
PMID:16189514
Towards a proteome-scale map of the human protein-protein in...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q59EK9 remains unresolved.
Reason: The human interactome abstract describes a broad interaction map; the exact pair assay and target construct were not inspected.
GO:0005515 protein binding
IPI
PMID:16189514
Towards a proteome-scale map of the human protein-protein in...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q92609 remains unresolved.
Reason: The human interactome abstract describes a broad interaction map; the exact pair assay and target construct were not inspected.
GO:0005515 protein binding
IPI
PMID:16189514
Towards a proteome-scale map of the human protein-protein in...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q9UHP6 remains unresolved.
Reason: The human interactome abstract describes a broad interaction map; the exact pair assay and target construct were not inspected.
GO:0005515 protein binding
IPI
PMID:19419997
The dyslexia-associated protein KIAA0319 interacts with adap...
MODIFY
Summary: Refine KIAA0319 binding to sorting-signal recognition.
Reason: Actual source Methods identify human ΞΌ2 NM_004068 constructs. Tail mapping and the KIAA0319 Y995A mutant connect ΞΌ2 association with its internalization signal. The refinement describes signal recognition; lysate pull-down and yeast interaction tests are not mislabeled as purified binary binding.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
GO:0005515 protein binding
IPI
PMID:20029029
Regulation of epidermal growth factor receptor trafficking b...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P00533 remains unresolved.
Reason: The abstract addresses HDAC6-dependent EGFR trafficking and a broader interaction survey; the AP2M1–EGFR supporting assay was not inspected.
GO:0005515 protein binding
IPI
PMID:21653829
Protein interactome reveals converging molecular pathways am...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P51114 remains unresolved.
Reason: The autism interactome abstract does not expose the exact target-pair experiment.
GO:0005515 protein binding
IPI
PMID:22916011
Identification and targeting of an interaction between a tyr...
MODIFY
Summary: Refine HCV-core binding to recognition of its sorting motif.
Reason: Actual source Methods use Human ORFeome AP2M1 constructs. Motif mutations, microfluidic binding and endogenous co-IP in infected Huh-7.5 cells support recognition of the core YxxΦ signal. Preserve the processed viral partner identifier and both recorded μ2 products; no viral-assembly BP is added.
Supporting Evidence:
PMID:22916011
Using microfluidics affinity analysis, protein-fragment complementation assays, and co-immunoprecipitations in infected cells, we show that this motif mediates core binding to AP2M1.
GO:0005515 protein binding
IPI
PMID:23676497
Endocytosis of synaptic ADAM10 in neuronal plasticity and Al...
KEEP AS NON CORE
Summary: Retain the source-specific ADAM10/AP-2 association.
Reason: The inspected human hippocampal co-IP detects ADAM10 with ΞΌ2 and the AP-2 alpha/beta subunits. The source row combines O14672, O94973, O95782 and P63010; a single new direct-binding mechanism should not be assigned to every listed pair. Preserve this positive complex association and express adaptor activity in the existing contributes_to annotation.
Supporting Evidence:
PMID:23676497
we identified the clathrin adaptor AP2 as an interacting partner of a previously uncharacterized atypical binding motif in the ADAM10 C-terminal domain.
GO:0005515 protein binding
IPI
PMID:23864651
The identification of novel proteins that interact with the ...
MODIFY
Summary: Specify binding to the GLP-1 G protein-coupled receptor.
Reason: The human receptor/human fetal-brain interaction screen and reported CHO co-IP validation directly identify AP2M1 as a GLP1R interactor. The available cache contains abstract and Discussion rather than complete Methods/Results; the refinement rests on the named partner class, not an inferred interface. Intracellular GPCR binding does not imply ligand, agonist or antagonist activity.
Supporting Evidence:
PMID:23864651
A screen of a human fetal brain cDNA prey library with an unliganded human GLP-1R as bait in yeast revealed 38 novel interactor protein candidates. These interactions were confirmed in mammalian Chinese hamster ovarian cells by coimmunoprecipitation.
GO:0005515 protein binding
IPI
PMID:24189400
Perturbation of the mutated EGFR interactome identifies vuln...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P00533 remains unresolved.
Reason: The EGFR network study is relevant, but its exact AP2M1 interaction evidence and validation table were not inspected.
GO:0005515 protein binding
IPI
PMID:24189400
Perturbation of the mutated EGFR interactome identifies vuln...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P63010 remains unresolved.
Reason: The EGFR network study is relevant, but its exact AP2M1 interaction evidence and validation table were not inspected. P63010 is the AP2B1 beta subunit; independent AP-2 architecture supports a plausible association but does not establish what this original experiment measured.
GO:0005515 protein binding
IPI
PMID:24603492
TBC1D5 and the AP2 complex regulate ATG9 trafficking and ini...
KEEP AS NON CORE
Summary: Retain ATG9A association with AP-2.
Reason: The inspected original Fig. 3 and Results show ATG9A co-precipitating the AP-2/clathrin machinery in human U2OS cells. They do not resolve a purified ΞΌ2-specific interface for this source row. Retain the positive association without borrowing the more specific motif mechanism from a different study.
Supporting Evidence:
PMID:24603492
ATG9 and TBC1D5 interact with clathrin and the AP2 complex.
GO:0005515 protein binding
IPI
PMID:24603492
TBC1D5 and the AP2 complex regulate ATG9 trafficking and ini...
KEEP AS NON CORE
Summary: Retain TBC1D5 association with ΞΌ2.
Reason: The inspected original Fig. 3 uses purified GST-AP2M1 to recover TBC1D5 from human HeLa lysate. Its additional purified-protein supplement was described but not independently inspected. This supports the pair without establishing a new molecular activity of ΞΌ2 or transferring TBC1D5's RabGAP function.
Supporting Evidence:
PMID:24603492
ATG9 and TBC1D5 interact with clathrin and the AP2 complex.
GO:0005515 protein binding
IPI
PMID:30021884
Histone Interaction Landscapes Visualized by Crosslinking Ma...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P63010 remains unresolved.
Reason: The nuclear crosslinking study is not rejected from its title or compartment; the exact AP2M1 pair and crosslink evidence were not inspected. P63010 is the AP2B1 beta subunit; independent AP-2 architecture supports a plausible association but does not establish what this original experiment measured.
GO:0005515 protein binding
IPI
PMID:31515488
Extensive disruption of protein interactions by genetic vari...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P51116 remains unresolved.
Reason: The variant-interaction survey was read at abstract scope; the exact pair and variant conditions were not inspected.
GO:0005515 protein binding
IPI
PMID:31515488
Extensive disruption of protein interactions by genetic vari...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q92609 remains unresolved.
Reason: The variant-interaction survey was read at abstract scope; the exact pair and variant conditions were not inspected.
GO:0005515 protein binding
IPI
PMID:31980649
Extensive rewiring of the EGFR network in colorectal cancer ...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P00533 remains unresolved.
Reason: The KRAS-dependent EGFR network abstract does not resolve the precise AP2M1 pair experiment.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P0DPH9 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P35268 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P55081 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P62807 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q00013 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q03393 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q16778 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q1ED39 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q68CQ4 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q6ZUT1 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q86YD7 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q8N3F0 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
KEEP AS NON CORE
Summary: HuRI records the AP2M1–MAB21L3 pair; its physiological role is unresolved.
Reason: The primary HuRI portal interaction table explicitly records MAB21L3–AP2M1 in HI-III. Together with the original human ORFeome Y2H workflow, this supports the reported pair as an experimental interaction. It does not identify an endocytic sorting motif, cargo/adaptor mechanism, endogenous colocalization, or an isoform-specific interface. Retain as non-core rather than invent a specific molecular function. The indexed primary table at https://interactome-atlas.org/search/MAB21L3 lists this pair with score 0.823; sequence-level clone details were not retrieved. The score is not used to infer a physiological mechanism.
Supporting Evidence:
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q8N9N8 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
KEEP AS NON CORE
Summary: HuRI records the AP2M1–PRPF38A pair; its physiological role is unresolved.
Reason: The primary HuRI portal interaction table explicitly records PRPF38A–AP2M1 in HI-III. Together with the original human ORFeome Y2H workflow, this supports the reported pair as an experimental interaction. It does not identify an endocytic sorting motif, cargo/adaptor mechanism, endogenous colocalization, or an isoform-specific interface. Retain as non-core rather than invent a specific molecular function. The indexed primary table at https://interactome-atlas.org/search/PRPF38A lists this pair with score 0.888; sequence-level clone details were not retrieved. The score is not used to infer a physiological mechanism.
Supporting Evidence:
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q96EY4 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
KEEP AS NON CORE
Summary: HuRI records the AP2M1–EAF1 pair; its physiological role is unresolved.
Reason: The primary HuRI portal interaction table explicitly records EAF1–AP2M1 in HI-III. Together with the original human ORFeome Y2H workflow, this supports the reported pair as an experimental interaction. It does not identify an endocytic sorting motif, cargo/adaptor mechanism, endogenous colocalization, or an isoform-specific interface. Retain as non-core rather than invent a specific molecular function. The EAF1–AP2M1 row was observed among neighboring-network interactions on the indexed MAB21L3 page (https://interactome-atlas.org/search/MAB21L3), with HI-III score 0.809. This is not a claim that an EAF1-specific portal page was inspected; sequence-level clone details were not retrieved. The score is not used to infer a physiological mechanism.
Supporting Evidence:
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q96MH2 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q99633 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q99880 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q9BSU3 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q9BU76 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q9NZ81 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q9P0T4 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q9UHP6 remains unresolved.
Reason: The recovered HuRI article describes the reference interaction map, but its body contains no AP2M1/Q96CW1 target row and does not include the pair-level supplementary tables. The original target-pair record and construct controls are therefore not available in the inspected material. A new source-specific molecular mechanism cannot be assigned from the paper-level assay description.
GO:0005515 protein binding
IPI
PMID:33436498
Cytoplasmic short linear motifs in ACE2 and integrin Ξ²(3) li...
MODIFY
Summary: Specify recognition of the ATG9A sorting sequence.
Reason: The inspected original Table 2/Fig. 3 measures binding of human Q96CW1 ΞΌ2 residues 160–435 to the ATG9A 4–14 YxxΞ¦ probe. This supports the specific signal-recognition refinement for this source, independently of the other ATG9A screen rows.
Supporting Evidence:
GO:0005515 protein binding
IPI
PMID:33436498
Cytoplasmic short linear motifs in ACE2 and integrin Ξ²(3) li...
MODIFY
Summary: Specify recognition of the ACE2 endocytic sorting sequence.
Reason: The inspected original Figs. 3–4 and Table 4 measure human ΞΌ2-domain binding to ACE2-tail peptides; Tyr781 phosphorylation inhibits it. This is a low-affinity biochemical motif interaction, not proof that the interaction is required for viral entry.
Supporting Evidence:
PMID:33436498
a binding site for the clathrin adaptor AP2 ΞΌ2 in ACE2 overlaps with a phospho-dependent binding site for the SH2 domains of Src family tyrosine kinases.
GO:0005515 protein binding
IPI
PMID:33961781
Dual proteome-scale networks reveal cell-specific remodeling...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P63010 remains unresolved.
Reason: The two-cell-line interactome combines context-dependent co-complex evidence; the exact AP2M1 pair record was not inspected. P63010 is the AP2B1 beta subunit; independent AP-2 architecture supports a plausible association but does not establish what this original experiment measured.
GO:0005515 protein binding
IPI
PMID:33961781
Dual proteome-scale networks reveal cell-specific remodeling...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q59EK9 remains unresolved.
Reason: The two-cell-line interactome combines context-dependent co-complex evidence; the exact AP2M1 pair record was not inspected.
GO:0005515 protein binding
IPI
PMID:33961781
Dual proteome-scale networks reveal cell-specific remodeling...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q68CQ4 remains unresolved.
Reason: The two-cell-line interactome combines context-dependent co-complex evidence; the exact AP2M1 pair record was not inspected.
GO:0005515 protein binding
IPI
PMID:34799561
Large scale discovery of coronavirus-host factor protein int...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q7Z3C6 remains unresolved.
Reason: The coronavirus motif-screen abstract does not establish the precise AP2M1–ATG9A construct and assay used for this row. Independent targeted ATG9A motif evidence is considered separately and is not relabeled as this source's experiment.
GO:0005515 protein binding
IPI
PMID:34964704
AP2M1 mediates autophagy-induced CLDN2 (claudin 2) degradati...
MODIFY
Summary: Specify CLDN2 sorting-sequence recognition.
Reason: The inspected Fig. 7/Methods show endogenous human AP2M1 association with CLDN2 and loss of association after mutations in CLDN2 YxxΦ motifs. Human epithelial-cell depletion links this interaction to internalization. The evidence supports motif-dependent cargo recognition without a purified binary-interface claim.
Supporting Evidence:
PMID:34964704
Site-directed mutagenesis revealed that AP2M1 binds to CLDN2 tyrosine motifs (YXXΠ€) (67-70 and 148-151).
GO:0005515 protein binding
IPI
PMID:35271311
OpenCell: Endogenous tagging for the cartography of human ce...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P63010 remains unresolved.
Reason: The OpenCell abstract describes endogenous tagging and interaction mapping; the AP2M1–AP2B1 pair record was not inspected. P63010 is the AP2B1 beta subunit; independent AP-2 architecture supports a plausible association but does not establish what this original experiment measured.
GO:0005515 protein binding
IPI
PMID:35384245
Physical and functional interactome atlas of human receptor ...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P00533 remains unresolved.
Reason: The receptor-tyrosine-kinase atlas combines several assay types; the exact AP2M1–EGFR evidence was not inspected.
GO:0005515 protein binding
IPI
PMID:37100772
Large-scale phage-based screening reveals extensive pan-vira...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P00533 remains unresolved.
Reason: The pan-viral motif screen contains many interactions, but this exact AP2M1 partner peptide and its validation were not inspected.
GO:0005515 protein binding
IPI
PMID:37100772
Large-scale phage-based screening reveals extensive pan-vira...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P18272 remains unresolved.
Reason: The pan-viral motif screen contains many interactions, but this exact AP2M1 partner peptide and its validation were not inspected.
GO:0005515 protein binding
IPI
PMID:37100772
Large-scale phage-based screening reveals extensive pan-vira...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q77IS8 remains unresolved.
Reason: The pan-viral motif screen contains many interactions, but this exact AP2M1 partner peptide and its validation were not inspected.
GO:0005515 protein binding
IPI
PMID:37100772
Large-scale phage-based screening reveals extensive pan-vira...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q7Z3C6 remains unresolved.
Reason: The pan-viral motif screen contains many interactions, but this exact AP2M1 partner peptide and its validation were not inspected. Independent targeted ATG9A motif evidence is considered separately and is not relabeled as this source's experiment.
GO:0005515 protein binding
IPI
PMID:37219487
Large-scale phosphomimetic screening identifies phospho-modu...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:Q7Z3C6 remains unresolved.
Reason: The phosphomimetic-screen abstract does not resolve the AP2M1–ATG9A peptide conditions or binding result. Independent targeted ATG9A motif evidence is considered separately and is not relabeled as this source's experiment.
GO:0005515 protein binding
IPI
PMID:39251607
Systematic identification of post-transcriptional regulatory...
UNDECIDED
Summary: The source-specific interaction with UniProtKB:P51116 remains unresolved.
Reason: The post-transcriptional regulatory-network abstract does not expose this AP2M1–FXR2 experiment or distinguish proximity from other interaction evidence.
GO:0005765 lysosomal membrane
TAS
Reactome:R-HSA-190519
UNDECIDED
Summary: Destination-membrane localization is not resolved by the available event record.
Reason: The R-HSA-190519 summary describes internalized annular gap-junction cargo and subsequent endo/lysosomal degradation. It does not expose a ΞΌ2-localization measurement or the compartmented molecular participants behind the TAS mapping. Keep that specific localization unresolved; cargo delivery is neither evidence of ΞΌ2 residence nor proof that no transient destination pool can exist.
GO:0005765 lysosomal membrane
TAS
Reactome:R-HSA-190829
UNDECIDED
Summary: Destination-membrane localization is not resolved by the available event record.
Reason: R-HSA-190829 describes lysosomal destruction of internalized gap-junction plaques. The exact AP2M1-bearing entity and primary localization evidence underlying the TAS assertion have not been recovered. Preserve uncertainty rather than infer either destination residence or its absence from a cargo summary.
GO:0005829 cytosol
IBA
GO_REF:0000033
ACCEPT
Summary: Retain the soluble cytosolic AP-2 pool.
Reason: The actual PTN000242370 cytosol IBD was inspected. AP-2 structural and recruitment evidence independently establishes a cytosolic pool that engages the plasma membrane. Full target-to-node topology was not reconstructed; the donor list is not treated as a support score.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
PANTHER:PTN000242370 UNRESOLVED
The actual PAINT node assertion was inspected in the local family export; full tree/MSA placement was not reconstructed. Target self-inclusion and donor count are not defects.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
GO:0005829 cytosol
TAS
Reactome:R-HSA-167537
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-167537 places AP-2 recruitment at the cytoplasmic Nef–CD4 complex. Retain the broad cytosolic ΞΌ2 pool in this adaptor-recruitment context.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-167537
GO:0005829 cytosol
TAS
Reactome:R-HSA-167597
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-167597 describes internalization of the Nef–CD4–AP2 complex. This is compatible with ΞΌ2 functioning on the cytoplasmic coat; it is not evidence that ΞΌ2 enters the vesicle lumen.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-167597
GO:0005829 cytosol
TAS
Reactome:R-HSA-167601
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-167601 describes CD4 degradation. Retain the existing broad cytosolic AP2 annotation using established adaptor localization, while distinguishing ΞΌ2 from the cargo delivered for proteolysis.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-167601
GO:0005829 cytosol
TAS
Reactome:R-HSA-167637
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-167637 follows Nef-driven CD28 uptake toward lysosomal degradation. ΞΌ2 participates from the cytoplasmic adaptor pool; the destination of CD28 is not automatically the location of ΞΌ2.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-167637
GO:0005829 cytosol
TAS
Reactome:R-HSA-182171
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-182171 concerns CD8 degradation after uptake. The accepted cytosolic ΞΌ2 annotation reflects its established adaptor pool, not a claim that it is the luminal lysosomal degradation substrate.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-182171
GO:0005829 cytosol
TAS
Reactome:R-HSA-182186
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-182186 describes Nef-dependent CD8 trafficking and explicitly leaves some downstream COPI interactions unresolved. The broad cytosolic AP2 location is retained without claiming those later interfaces have been reconstructed.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-182186
GO:0005829 cytosol
TAS
Reactome:R-HSA-182198
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-182198 describes CD8 endocytosis through the named AP2-containing complex. Retain ΞΌ2’s cytoplasmic coat/adaptor context.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-182198
GO:0005829 cytosol
TAS
Reactome:R-HSA-3928654
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-3928654 describes EPH/ephrin internalization with cytoplasmic partners and clathrin coats. The broad cytosolic ΞΌ2 annotation fits that adaptor role; EEA1-positive cargo arrival is a separate observation.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-3928654
GO:0005829 cytosol
TAS
Reactome:R-HSA-5138433
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-5138433 explicitly describes ΞΌ2 recognition of DVL2 DEP/YHEL elements during FZD4 uptake. This supports its cytoplasmic cargo-recognition role, with the model’s uncertainty about beta-arrestin requirement preserved.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-5138433
GO:0005829 cytosol
TAS
Reactome:R-HSA-5138459
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-5138459 describes AP2-dependent WNT5A–FZD4 uptake and possible beta-arrestin dissociation. Retain the broad cytosolic ΞΌ2 location; beta-arrestin timing is not treated as direct ΞΌ2 dissociation evidence.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-5138459
GO:0005829 cytosol
TAS
Reactome:R-HSA-8856808
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8856808 describes recruitment of soluble AP2, opening of its core and cargo capture at the plasma membrane. These transitions directly fit an adaptor cycling from the cytosol to the coat.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8856808
GO:0005829 cytosol
TAS
Reactome:R-HSA-8856813
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8856813 identifies ΞΌ2 Thr156 as the AAK1 phosphorylation substrate. Retain the cytoplasmic AP2 pool in this regulatory event; AAK1 performs the phosphorylation.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8856813
GO:0005829 cytosol
TAS
Reactome:R-HSA-8862280
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8862280 describes FCHo/SGIP-dependent stabilization and opening of AP2 at nascent pits. The cytosolic annotation fits recruitment of the adaptor into that assembly, distinct from the F-BAR membrane interaction.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8862280
GO:0005829 cytosol
TAS
Reactome:R-HSA-8866277
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8866277 distinguishes direct cargo recognition by the AP2 core from accessory-protein recruitment by its ears. Retain the cytosolic ΞΌ2 location consistent with the core’s cargo-recognition role.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8866277
GO:0005829 cytosol
TAS
Reactome:R-HSA-8866283
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8866283 recruits GPCRs through beta-arrestins and clathrin-coat components. ΞΌ2’s cytosolic AP2 location is retained without assigning it every beta-arrestin receptor interface.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8866283
GO:0005829 cytosol
TAS
Reactome:R-HSA-8867754
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8867754 concerns staged BAR-protein recruitment to the AP2/clathrin pit. Retain the broad cytosolic adaptor pool; the curvature-recognition domains belong to the recruited BAR proteins.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8867754
GO:0005829 cytosol
TAS
Reactome:R-HSA-8867756
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8867756 describes concurrent CLASP/cargo and AP2/clathrin assembly. Retain ΞΌ2’s cytosolic recruitment context while preserving the source’s caveat that these events need not be strictly sequential.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8867756
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868071
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868071 recruits PIK3C2A through clathrin during pit maturation. The retained ΞΌ2 annotation is cytosolic coat/adaptor context; direct PIK3C2A recruitment by ΞΌ2 is not inferred.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868071
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868072
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868072 describes PIK3C2A lipid phosphorylation within a maturing coat. Retain the broad ΞΌ2 cytosolic context rather than interpret the reaction title as ΞΌ2 catalytic activity.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868072
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868230
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868230 describes SNX9-linked actin machinery at late pits, particularly under membrane tension. ΞΌ2 remains a cytoplasmic adaptor component; this record does not establish its own actin-nucleation mechanism.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868230
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868236
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868236 recruits dynamin through BAR and other coat partners. Retain the cytoplasmic AP2 context without claiming a ΞΌ2-specific dynamin interface from this summary.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868236
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868648
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868648 connects phosphoinositide turnover with coat disassembly and uncoating. This is compatible with AP2 exchange between membrane-bound and cytosolic pools; the exact ΞΌ2 release kinetics are not supplied.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868648
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868651
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868651 describes synaptojanin recruitment by endophilin, clathrin and AP2-associated machinery. Retain broad cytosolic ΞΌ2 localization while distinguishing the synaptojanin isoform/timing findings from ΞΌ2 itself.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868651
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868658
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868658 concerns HSPA8-mediated clathrin disassembly. Retain ΞΌ2’s independently established cytosolic pool, but this specific clathrin–HSPA8 mechanism is not recast as an isolated ΞΌ2 assay.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868658
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868659
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868659 recruits auxilins after fission, priming vesicles for endosomal fusion. The broad ΞΌ2 cytosolic annotation is consistent with coat recycling; auxilin-mediated uncoating does not itself locate ΞΌ2 on the destination membrane.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868659
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868660
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868660 describes HSPA8 recruitment and persistence on liberated clathrin. Retain the general ΞΌ2 cytosolic pool without transferring the measured HSPA8/clathrin stoichiometry to AP2.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868660
GO:0005829 cytosol
TAS
Reactome:R-HSA-8868661
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8868661 describes dynamin-dependent fission of an AP2/clathrin-coated pit. Retain ΞΌ2’s cytoplasmic adaptor context; dynamin is the GTPase that executes scission.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8868661
GO:0005829 cytosol
TAS
Reactome:R-HSA-8869438
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8869438 explicitly states that the timing and mechanism of coat-associated protein loss remain poorly understood. Retain cytosolic AP2 recycling without converting this broad model into a universal claim about ΞΌ2 release timing.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8869438
GO:0005829 cytosol
TAS
Reactome:R-HSA-8871193
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8871193 links AAK1 displacement and ΞΌ2 dephosphorylation with weakened AP2–membrane association. This directly fits regulated return toward a cytosolic pool; exact destination exclusion is not demonstrated.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8871193
GO:0005829 cytosol
TAS
Reactome:R-HSA-8871194
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8871194 models RAB5/GAPVD1-dependent AP2 uncoating and membrane-lipid turnover. Retain the broad cytosolic adaptor location while preserving the predictive nature of the timing/model.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8871194
GO:0005829 cytosol
TAS
Reactome:R-HSA-8871196
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-8871196 describes transient AP2/clathrin recruitment from soluble pools to PI(4,5)P2-rich plasma membrane. The source explicitly notes variable temporal order and abortive pits, supporting a dynamic cytosolic/membrane cycle.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-8871196
GO:0005829 cytosol
TAS
Reactome:R-HSA-9681616
ACCEPT
Summary: Retain the cytosolic coat/adaptor pool in this pathway event.
Reason: R-HSA-9681616 describes chlorpromazine perturbing AP2/clathrin relocation. Retain the broad cytosolic adaptor pool; pharmacological disruption is not a newly assigned physiological ΞΌ2 ligand function.
Supporting Evidence:
PMID:20603002
In the "locked" cytosolic form, AP2's binding sites for the two endocytic motifs, YxxPhi on the C-terminal domain of mu2 (C-mu2) and [ED]xxxL[LI] on sigma2, are blocked by parts of beta2.
Reactome:R-HSA-9681616
GO:0005886 plasma membrane
EXP
PMID:31104773
A Recurrent Missense Variant in AP2M1 Impairs Clathrin-Media...
ACCEPT
Summary: ΞΌ2 localizes to the plasma membrane during coat formation.
Reason: Actual source imaging shows ΞΌ2 recruitment in human HeLa cells; Arg170Trp preserves membrane/pit localization despite impaired uptake. The named mCherry rescue reagent is rat, distinct from human host-cell depletion and patient genetics. Preserve the source annotation without claiming all constructs are human.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
GO:0005886 plasma membrane
IDA
GO_REF:0000052
ACCEPT
Summary: Retain plasma-membrane localization.
Reason: The HPA-derived cellular localization agrees with independently inspected ΞΌ2 recruitment and uptake experiments. The HPA image-level evidence was not separately reanalyzed, and this acceptance does not infer an integral membrane topology.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
GO:0005886 plasma membrane
IDA
PMID:14530274
The AP-2 complex is excluded from the dynamic population of ...
ACCEPT
Summary: Retain activity at plasma-membrane coated sites.
Reason: The cited study images alpha/beta2 adaptors at the plasma membrane. Independent ΞΌ2 imaging confirms the location, while later work documents limitations of tagged adaptor dynamics. Preserve the location assertion without adopting the older kinetic interpretation as a universal exclusion of AP-2 from vesicles.
Supporting Evidence:
PMID:17035630
Mutating the PtdIns(4,5)P2 binding site of alpha, the phosphorylation site of mu2, or the YXXPhi binding site of mu2 impairs AP-2 function, as assayed by transferrin uptake.
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
GO:0005886 plasma membrane
IEA
GO_REF:0000044
ACCEPT
Summary: Retain peripheral plasma-membrane localization.
Reason: The UniProt subcellular-location mapping matches independently inspected ΞΌ2 localization. AP2M1 acts on the cytoplasmic side as an adaptor subunit, rather than as a transmembrane receptor.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
UniProtKB-SubCell:SL-0039 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-167633
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-167633 event concerns Formation of Nef:Cd28:Clathrin-coated Pit Adapter Protein complex. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-167633
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-167637
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-167637 event concerns Internalization of Nef:CD28:Clathrin-Coated Pit Adapter Protein Complex. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-167637
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-177479
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-177479 event concerns Axonal transport of NGF:Trk complexes. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-177479
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-190519
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-190519 event concerns Internalization of gap junction plaques. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-190519
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-196017
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-196017 event concerns Dynamin is recruited to the gap junction plaque. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-196017
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-196026
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-196026 event concerns Dab2 is recruited to the junctional plaques. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-196026
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2130640
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-2130640 event concerns Recruitment of clathrin coated vesicle by Ii. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-2130640
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2130725
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-2130725 event concerns Internalization of MHC II:Ii clathrin coated vesicle. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-2130725
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-392748
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-392748 event concerns L1 binds to AP-2 Clathrin complex. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-392748
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-392749
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-392749 event concerns Transport of L1 into endosomes. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-392749
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-445071
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-445071 event concerns Reinsertion of L1 into the plasma membrane. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-445071
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-445079
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-445079 event concerns Phosphorylation of L1 by ERK. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-445079
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-555065
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-555065 event concerns Formation of clathrin coated vesicle. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-555065
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-6784729
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-6784729 event concerns PCSK9:LDLR:Clathrin-coated vesicle transport from plasma membrane to endolysosome. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-6784729
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-6784735
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-6784735 event concerns PCSK9:LDLR bind to Clathrin. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-6784735
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8855130
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8855130 event concerns VLDLR:PCSK9:Clathrin-coated vesicle translocates from the plasma membrane to lysosomal membrane. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8855130
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8855131
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8855131 event concerns VLDLR:PCSK9 binds Clathrin-coated vesicles. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8855131
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8866279
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8866279 event concerns Epsin family proteins bind ubiquitinated cargo. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8866279
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8867754
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8867754 event concerns F- and N- BAR domain proteins bind the clathrin-coated pit. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8867754
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8867756
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8867756 event concerns CLASP proteins and cargo are recruited to the nascent clathrin-coated pit. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8867756
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8868071
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8868071 event concerns Clathrin recruits PIK3C2A. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8868071
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8868072
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8868072 event concerns Clathrin-associated PIK3C2A phosphorylates PI(4)P to PI(3,4)P2. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8868072
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8868230
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8868230 event concerns SNX9 recruits components of the actin polymerizing machinery. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8868230
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8868236
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8868236 event concerns BAR domain proteins recruit dynamin. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8868236
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8868648
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8868648 event concerns SYNJ hydrolyze PI(4,5)P2 to PI(4)P. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8868648
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8868651
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8868651 event concerns Endophilins recruit synaptojanins to the clathrin-coated pit. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8868651
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8868661
ACCEPT
Summary: Retain the plasma-membrane-associated AP-2 pool.
Reason: The inspected Reactome:R-HSA-8868661 event concerns Dynamin-mediated GTP hydrolysis promotes vesicle scission. Its component-level plasma-membrane assignment agrees with independently observed ΞΌ2 recruitment to endocytic sites. This retains location, not a claim that ΞΌ2 performs every molecular reaction or accompanies cargo throughout later degradation.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
Reactome:R-HSA-8868661
GO:0005905 clathrin-coated pit
IDA
PMID:31104773
A Recurrent Missense Variant in AP2M1 Impairs Clathrin-Media...
ACCEPT
Summary: ΞΌ2 occupies clathrin-coated pits.
Reason: Actual source imaging and quantitative coat analysis retain ΞΌ2 at clathrin-positive sites, including the disease-associated mutant that impairs uptake. Localization and functional impairment are distinct observations.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
GO:0006886 intracellular protein transport
IEA
GO_REF:0000002
ACCEPT
Summary: Retain intracellular protein transport through the AP-2 adaptor.
Reason: The InterPro family mapping is broad, but human cargo-recognition and uptake evidence supports this existing transport assertion. It does not give ΞΌ2 every trafficking route of the wider adaptor family.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
InterPro:IPR001392 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
InterPro:IPR018240 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
GO:0006896 Golgi to vacuole transport
IBA
GO_REF:0000033
UNDECIDED
Summary: The inherited Golgi-to-vacuole route remains unresolved.
Reason: The exact PTN000055849 GO:0006896 IBD was inspected, together with a same-term IRD at PTN002575694. Full family topology/MSA are unavailable, so that loss cannot be assigned to AP2M1 from node order or taxon alone. The current term includes Golgi-to-lysosome transport; plasma-membrane cargo entry is not equivalent evidence. No donor-count, self-source or yeast-only objection is made.
Propagation Review
Root cause: UNRESOLVED
Sources checked:
PANTHER:PTN000055849 UNRESOLVED
Actual PTN000055849 GO:0006896 IBD and same-term IRD at PTN002575694 were read, but topology linking the target to the loss branch is unresolved. Node order and taxon alone cannot assign a loss to AP2M1.
GO:0006900 vesicle budding from membrane
IMP
PMID:25898166
CALM regulates clathrin-coated vesicle size and maturation b...
ACCEPT
Summary: AP-2 contributes structurally to endocytic budding.
Reason: The actual source includes ΞΌ2 depletion and rescue experiments: loss of ΞΌ2 strongly reduces coated-pit abundance and changes coat size. AP2M1 supplies cargo/coat structure; CALM supplies the directly characterized curvature-sensing amphipathic helix. This existing process assertion does not assign CALM's membrane-bending chemistry to ΞΌ2.
Supporting Evidence:
PMID:25898166
AP2 depletion (of μ2) resulted in an ∼12-fold decrease in the number of endocytic CCPs/CCVs
GO:0008021 synaptic vesicle
IBA
GO_REF:0000033
ACCEPT
Summary: Retain the synaptic-vesicle retrieval context.
Reason: The conserved AP-2 coat participates in reformation and sorting of synaptic-vesicle membrane. This is a transient active retrieval pool, not a claim that AP2M1 is an integral resident of every mature synaptic vesicle. The actual synaptic-vesicle IBD is available for row 134; the full topology and exact rat transfer experiment remain uninspected.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
PANTHER:PTN008307226 UNRESOLVED
The actual PAINT node assertion was inspected in the local family export; full tree/MSA placement was not reconstructed. Target self-inclusion and donor count are not defects.
Supporting Evidence:
PMID:11102472
Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.
PMID:31671891
ΞΌ2-subunit also interacts with C2 domains as present in Synaptotagmin1 [11] or Otoferlin [12].
GO:0008021 synaptic vesicle
IEA
GO_REF:0000107
ACCEPT
Summary: Retain the synaptic-vesicle retrieval context.
Reason: The conserved AP-2 coat participates in reformation and sorting of synaptic-vesicle membrane. This is a transient active retrieval pool, not a claim that AP2M1 is an integral resident of every mature synaptic vesicle. The actual synaptic-vesicle IBD is available for row 134; the full topology and exact rat transfer experiment remain uninspected.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:11102472
Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.
PMID:31671891
ΞΌ2-subunit also interacts with C2 domains as present in Synaptotagmin1 [11] or Otoferlin [12].
GO:0009898 cytoplasmic side of plasma membrane
NAS
PMID:29184887
Weak Molecular Interactions in Clathrin-Mediated Endocytosis...
ACCEPT
Summary: AP2M1 acts at the cytoplasmic plasma-membrane surface.
Reason: The inspected adaptor review places the AP-2 cargo-recognition machinery on the cytoplasmic side, consistent with ΞΌ2 cargo-tail recognition and peripherally recruited coat architecture.
Supporting Evidence:
PMID:29184887
It interacts directly with two types of internalization motifs (LL and (Y-X-X-Ξ¦) (Ξ¦ = hydrophobic residue) found within the cytoplasmic domains of integral membrane protein receptors via its Οƒ (LL) and ΞΌ2 (Y-X-X-Ξ¦) subunits
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
GO:0016020 membrane
IEA
GO_REF:0000044
ACCEPT
Summary: Retain membrane association of the peripheral adaptor.
Reason: The broad electronic membrane assignment agrees with direct ΞΌ2 imaging and membrane-dependent AP-2 opening. It does not imply a transmembrane segment or indiscriminate residence at all organelles.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
UniProtKB-SubCell:SL-0162 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:31104773
However, we found that AP-2ΞΌ WT and p.Arg170Trp variants colocalize with clathrin equally well in HeLa cells, suggesting that localization of the pathogenic AP-2ΞΌ variant in clathrin-coated pits is not altered.
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
GO:0016192 vesicle-mediated transport
IEA
GO_REF:0000002
ACCEPT
Summary: AP2M1 participates in vesicle-mediated cargo transport.
Reason: The broad family mapping is independently grounded by ΞΌ2 sorting-signal recognition and AP-2-dependent uptake. It is restricted here to the established adaptor role.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
InterPro:IPR001392 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
InterPro:IPR018240 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
GO:0016192 vesicle-mediated transport
NAS
PMID:15941406
Analysis of the AP-2 adaptor complex and cargo during clathr...
ACCEPT
Summary: Retain vesicle-mediated transport as an adaptor function.
Reason: The cited paper discusses tagged AP-2 and cargo dynamics. Its early interpretation of AP-2 exclusion from internalizing carriers has construct-dependent limitations documented by later work. The broad transport function is independently well established, so the retained annotation does not require endorsing every kinetic conclusion.
Supporting Evidence:
PMID:17035630
Mutating the PtdIns(4,5)P2 binding site of alpha, the phosphorylation site of mu2, or the YXXPhi binding site of mu2 impairs AP-2 function, as assayed by transferrin uptake.
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
GO:0030122 AP-2 adaptor complex
IBA
GO_REF:0000033
ACCEPT
Summary: AP2M1 is the medium subunit of the AP-2 complex.
Reason: Sequence, AP-2 structural studies and human tissue co-immunoprecipitation agree on the medium subunit's complex membership. The original reference and qualifier are retained; PAINT target self-inclusion is legitimate experimental grounding, not circularity.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
PANTHER:PTN000242370 UNRESOLVED
The actual PAINT node assertion was inspected in the local family export; full tree/MSA placement was not reconstructed. Target self-inclusion and donor count are not defects.
Supporting Evidence:
PMID:12086608
We describe the structure of the 200 kDa AP2 "core" (alpha trunk, beta2 trunk, mu2, and sigma2)
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:23676497
we identified the clathrin adaptor AP2 as an interacting partner of a previously uncharacterized atypical binding motif in the ADAM10 C-terminal domain.
GO:0030122 AP-2 adaptor complex
IDA
PMID:23676497
Endocytosis of synaptic ADAM10 in neuronal plasticity and Al...
ACCEPT
Summary: AP2M1 is the medium subunit of the AP-2 complex.
Reason: Sequence, AP-2 structural studies and human tissue co-immunoprecipitation agree on the medium subunit's complex membership. The original reference and qualifier are retained; PAINT target self-inclusion is legitimate experimental grounding, not circularity.
Supporting Evidence:
PMID:12086608
We describe the structure of the 200 kDa AP2 "core" (alpha trunk, beta2 trunk, mu2, and sigma2)
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:23676497
we identified the clathrin adaptor AP2 as an interacting partner of a previously uncharacterized atypical binding motif in the ADAM10 C-terminal domain.
GO:0030122 AP-2 adaptor complex
IEA
GO_REF:0000120
ACCEPT
Summary: AP2M1 is the medium subunit of the AP-2 complex.
Reason: Sequence, AP-2 structural studies and human tissue co-immunoprecipitation agree on the medium subunit's complex membership. The original reference and qualifier are retained; PAINT target self-inclusion is legitimate experimental grounding, not circularity.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
ARBA:ARBA00027293 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:12086608
We describe the structure of the 200 kDa AP2 "core" (alpha trunk, beta2 trunk, mu2, and sigma2)
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:23676497
we identified the clathrin adaptor AP2 as an interacting partner of a previously uncharacterized atypical binding motif in the ADAM10 C-terminal domain.
GO:0030122 AP-2 adaptor complex
NAS
PMID:12086608
Molecular architecture and functional model of the endocytic...
ACCEPT
Summary: AP2M1 is the medium subunit of the AP-2 complex.
Reason: Sequence, AP-2 structural studies and human tissue co-immunoprecipitation agree on the medium subunit's complex membership. The original reference and qualifier are retained; PAINT target self-inclusion is legitimate experimental grounding, not circularity.
Supporting Evidence:
PMID:12086608
We describe the structure of the 200 kDa AP2 "core" (alpha trunk, beta2 trunk, mu2, and sigma2)
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:23676497
we identified the clathrin adaptor AP2 as an interacting partner of a previously uncharacterized atypical binding motif in the ADAM10 C-terminal domain.
GO:0030122 AP-2 adaptor complex
TAS
PMID:12121421
The mu2 subunit of the clathrin adaptor AP-2 binds to FDNPVY...
ACCEPT
Summary: AP2M1 is the medium subunit of the AP-2 complex.
Reason: Sequence, AP-2 structural studies and human tissue co-immunoprecipitation agree on the medium subunit's complex membership. The original reference and qualifier are retained; PAINT target self-inclusion is legitimate experimental grounding, not circularity.
Supporting Evidence:
PMID:12086608
We describe the structure of the 200 kDa AP2 "core" (alpha trunk, beta2 trunk, mu2, and sigma2)
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:23676497
we identified the clathrin adaptor AP2 as an interacting partner of a previously uncharacterized atypical binding motif in the ADAM10 C-terminal domain.
GO:0030131 clathrin adaptor complex
IEA
GO_REF:0000002
ACCEPT
Summary: Retain membership in the clathrin adaptor machinery.
Reason: The family-level complex term is broader than AP-2 membership but correct. ΞΌ2 is a constituent of the heterotetramer, not an independently complete clathrin coat.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
InterPro:IPR001392 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
InterPro:IPR018240 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:12086608
We describe the structure of the 200 kDa AP2 "core" (alpha trunk, beta2 trunk, mu2, and sigma2)
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
GO:0030666 endocytic vesicle membrane
TAS
Reactome:R-HSA-416639
ACCEPT
Summary: Retain the transient endocytic carrier membrane pool.
Reason: The Reactome record describes constitutive AMPA-receptor recycling and does not itself resolve ΞΌ2 residence at every trafficking stage. Independent AP-2 structural/endocytic evidence supports transient peripheral association with forming endocytic carriers; no permanent recycling-endosome residence is inferred.
Supporting Evidence:
Reactome:R-HSA-416639
PMID:29184887
We have so far discussed how these interactions occur at the membrane, but in order for internalization to occur, the coated vesicle itself must form. This process of coat formation is driven by interactions between AP2 and its network of binding partners which bind to the appendage (or β€œear”) domains of Ξ±2-adaptin and Ξ²2-adaptin.
GO:0030669 clathrin-coated endocytic vesicle membrane
TAS
Reactome:R-HSA-5138459
ACCEPT
Summary: Retain association with forming clathrin-coated endocytic carriers.
Reason: The inspected WNT5A/FZD4 event explicitly depends on AP-2 during uptake. ΞΌ2 contributes to the coat-associated adaptor pool; this does not assert that the receptor, arrestin and AP-2 remain bound throughout all later traffic.
Supporting Evidence:
Reactome:R-HSA-5138459
PMID:29184887
We have so far discussed how these interactions occur at the membrane, but in order for internalization to occur, the coated vesicle itself must form. This process of coat formation is driven by interactions between AP2 and its network of binding partners which bind to the appendage (or β€œear”) domains of Ξ±2-adaptin and Ξ²2-adaptin.
GO:0031623 receptor internalization
IEA
GO_REF:0000117
ACCEPT
Summary: Retain receptor internalization.
Reason: Independent human-cell uptake and cargo-tail experiments support the automated process assignment, although the exact ARBA rule was not reconstructed. ΞΌ2 recognizes and organizes cargo within AP-2 rather than merely being consumed by internalization.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
ARBA:ARBA00087457 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
GO:0031623 receptor internalization
IMP
PMID:25898166
CALM regulates clathrin-coated vesicle size and maturation b...
ACCEPT
Summary: ΞΌ2 contributes to receptor uptake.
Reason: The source's ΞΌ2-depletion and rescue controls directly establish AP-2-dependent transferrin uptake. CALM is the focal protein, but actual ΞΌ2 experiments were inspected, so the paper title is not a misattribution argument.
Supporting Evidence:
PMID:25898166
AP2 depletion (of μ2) resulted in an ∼12-fold decrease in the number of endocytic CCPs/CCVs
GO:0035615 clathrin-cargo adaptor activity
IBA
GO_REF:0000033
ACCEPT
Summary: Retain ΞΌ2 contribution to clathrin-cargo adaptor activity.
Reason: ΞΌ2 supplies cargo-signal recognition within the assembled AP-2 adaptor; other subunits supply additional membrane and clathrin contacts. The authored core explicitly uses contributes_to_molecular_function. Original qualifiers are preserved, including the two existing contributes_to rows; acceptance does not claim autonomous isolated-ΞΌ2 bridging.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
PANTHER:PTN000055849 UNRESOLVED
The actual PAINT node assertion was inspected in the local family export; full tree/MSA placement was not reconstructed. Target self-inclusion and donor count are not defects.
Supporting Evidence:
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:23676497
we identified the clathrin adaptor AP2 as an interacting partner of a previously uncharacterized atypical binding motif in the ADAM10 C-terminal domain.
PMID:12121421
recognition of the FDNPVY signal is mediated by a binding site in the mu2-subunit that is distinct from the site for the more general YppØ sorting signal
GO:0035615 clathrin-cargo adaptor activity
IDA
PMID:23676497
Endocytosis of synaptic ADAM10 in neuronal plasticity and Al...
ACCEPT
Summary: Retain ΞΌ2 contribution to clathrin-cargo adaptor activity.
Reason: ΞΌ2 supplies cargo-signal recognition within the assembled AP-2 adaptor; other subunits supply additional membrane and clathrin contacts. The authored core explicitly uses contributes_to_molecular_function. Original qualifiers are preserved, including the two existing contributes_to rows; acceptance does not claim autonomous isolated-ΞΌ2 bridging.
Supporting Evidence:
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:23676497
we identified the clathrin adaptor AP2 as an interacting partner of a previously uncharacterized atypical binding motif in the ADAM10 C-terminal domain.
PMID:12121421
recognition of the FDNPVY signal is mediated by a binding site in the mu2-subunit that is distinct from the site for the more general YppØ sorting signal
GO:0035615 clathrin-cargo adaptor activity
TAS
PMID:12121421
The mu2 subunit of the clathrin adaptor AP-2 binds to FDNPVY...
ACCEPT
Summary: Retain ΞΌ2 contribution to clathrin-cargo adaptor activity.
Reason: ΞΌ2 supplies cargo-signal recognition within the assembled AP-2 adaptor; other subunits supply additional membrane and clathrin contacts. The authored core explicitly uses contributes_to_molecular_function. Original qualifiers are preserved, including the two existing contributes_to rows; acceptance does not claim autonomous isolated-ΞΌ2 bridging.
Supporting Evidence:
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:23676497
we identified the clathrin adaptor AP2 as an interacting partner of a previously uncharacterized atypical binding motif in the ADAM10 C-terminal domain.
PMID:12121421
recognition of the FDNPVY signal is mediated by a binding site in the mu2-subunit that is distinct from the site for the more general YppØ sorting signal
GO:0036020 endolysosome membrane
TAS
Reactome:R-HSA-2130486
UNDECIDED
Summary: Destination-membrane localization is not resolved by the available event record.
Reason: The R-HSA-2130486 event is named uncoating and fusion, while its short summary states that MHC-II/Ii vesicles fuse with endosomes. This suggests a coat-to-destination mapping question, but does not establish when ΞΌ2 leaves in that exact model or exclude a transient destination pool. The source-specific endolysosome-membrane claim remains unresolved.
GO:0036020 endolysosome membrane
TAS
Reactome:R-HSA-2130725
UNDECIDED
Summary: Destination-membrane localization is not resolved by the available event record.
Reason: R-HSA-2130725 describes MHC-II/Ii entry from the plasma membrane and conflicting dynamin-perturbation findings. Those experiments concern cargo uptake, not a ΞΌ2-specific measurement at the endolysosome. The original TAS localization remains unresolved without declaring the curated event incorrect.
GO:0036020 endolysosome membrane
TAS
Reactome:R-HSA-6784729
UNDECIDED
Summary: Destination-membrane localization is not resolved by the available event record.
Reason: R-HSA-6784729 follows PCSK9–LDLR cargo through endosomes to lysosomal degradation. The accessible summary does not expose the AP2M1-containing entity or a destination-localization assay. Retain uncertainty; well-established plasma-membrane coat function does not by itself exclude other membrane pools.
GO:0036020 endolysosome membrane
TAS
Reactome:R-HSA-6784738
UNDECIDED
Summary: Destination-membrane localization is not resolved by the available event record.
Reason: R-HSA-6784738 separates clathrin-dependent entry of PCSK9–LDLR from later lysosomal degradation. It does not establish ΞΌ2 persistence or absence at the destination; the exact compartment-mapping evidence needs inspection before a confident overannotation judgment.
GO:0036020 endolysosome membrane
TAS
Reactome:R-HSA-8855130
UNDECIDED
Summary: Destination-membrane localization is not resolved by the available event record.
Reason: R-HSA-8855130 names a VLDLR/PCSK9/clathrin-coated vesicle moving toward lysosomal membrane. The summary retains a coat label but does not resolve the timing or identity of AP2M1-containing participants. Keep the specific TAS localization unresolved rather than turn a plausible general uncoating model into an inspected source result.
GO:0044325 transmembrane transporter binding
IPI
PMID:23529131
Oestrogen promotes KCNQ1 potassium channel endocytosis and p...
ACCEPT
Summary: Retain binding to the membrane transporter KCNQ1.
Reason: The inspected original Fig. 3 shows endogenous ΞΌ2 co-IP with KCNQ1 in human HT29cl.19A cells, enhanced by estrogen during uptake. Co-IP supports the existing partner-class term; it does not establish a purified binary interface or experimentally map a particular KCNQ1 sorting motif.
Supporting Evidence:
PMID:23529131
KCNQ1 was retrieved from the plasma membrane by a clathrin-mediated endocytosis, which involved the association between KCNQ1 and the clathrin adaptor, AP-2.
GO:0045334 clathrin-coated endocytic vesicle
NAS
PMID:25898166
CALM regulates clathrin-coated vesicle size and maturation b...
ACCEPT
Summary: Retain association with the clathrin-coated endocytic carrier.
Reason: The actual study characterizes ΞΌ2-dependent coated-pit/vesicle production and uses ΞΌ2 depletion/rescue controls. The adaptor is a transient coat component, not a permanent luminal or integral membrane protein.
Supporting Evidence:
PMID:25898166
AP2 depletion (of μ2) resulted in an ∼12-fold decrease in the number of endocytic CCPs/CCVs
PMID:17035630
Mutating the PtdIns(4,5)P2 binding site of alpha, the phosphorylation site of mu2, or the YXXPhi binding site of mu2 impairs AP-2 function, as assayed by transferrin uptake.
GO:0048488 synaptic vesicle endocytosis
IDA
PMID:11102472
A conserved clathrin assembly motif essential for synaptic v...
ACCEPT
Summary: Retain participation in synaptic-vesicle retrieval.
Reason: The actual cited main paper uses bovine brain AP-2/clathrin assembly and squid presynaptic intervention to establish conserved complex-level coat function during retrieval. Preserve the curated human process assertion with this cross-species scope explicit; the experiments do not isolate a human ΞΌ2 catalytic step or transfer beta2 clathrin-binding chemistry to ΞΌ2.
Supporting Evidence:
PMID:11102472
Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.
PMID:31671891
ΞΌ2-subunit also interacts with C2 domains as present in Synaptotagmin1 [11] or Otoferlin [12].
GO:0048488 synaptic vesicle endocytosis
IEA
GO_REF:0000107
ACCEPT
Summary: Retain the conserved synaptic-vesicle endocytic role.
Reason: The rat transfer agrees with the conserved AP-2 cargo-sorting/coat role and independent neuronal adaptor evidence. Exact donor experiments were not reconstructed; this does not imply that ΞΌ2 acts as dynamin or clathrin.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:11102472
Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.
PMID:31671891
ΞΌ2-subunit also interacts with C2 domains as present in Synaptotagmin1 [11] or Otoferlin [12].
GO:0048488 synaptic vesicle endocytosis
IMP
PMID:11102472
A conserved clathrin assembly motif essential for synaptic v...
ACCEPT
Summary: Retain participation in synaptic-vesicle retrieval.
Reason: The actual cited main paper uses bovine brain AP-2/clathrin assembly and squid presynaptic intervention to establish conserved complex-level coat function during retrieval. Preserve the curated human process assertion with this cross-species scope explicit; the experiments do not isolate a human ΞΌ2 catalytic step or transfer beta2 clathrin-binding chemistry to ΞΌ2.
Supporting Evidence:
PMID:11102472
Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.
PMID:31671891
ΞΌ2-subunit also interacts with C2 domains as present in Synaptotagmin1 [11] or Otoferlin [12].
GO:0048488 synaptic vesicle endocytosis
NAS
PMID:11102472
A conserved clathrin assembly motif essential for synaptic v...
ACCEPT
Summary: Retain participation in synaptic-vesicle retrieval.
Reason: The actual cited main paper uses bovine brain AP-2/clathrin assembly and squid presynaptic intervention to establish conserved complex-level coat function during retrieval. Preserve the curated human process assertion with this cross-species scope explicit; the experiments do not isolate a human ΞΌ2 catalytic step or transfer beta2 clathrin-binding chemistry to ΞΌ2.
Supporting Evidence:
PMID:11102472
Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.
PMID:31671891
ΞΌ2-subunit also interacts with C2 domains as present in Synaptotagmin1 [11] or Otoferlin [12].
GO:0050750 low-density lipoprotein particle receptor binding
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Retain LDL-receptor-tail binding as a secondary interaction context.
Reason: PMID:12121421 directly reports μ2 binding to FDNPVY and YppΦ peptides at distinct sites. Preserve the rat-derived assertion without claiming that the human endogenous LDLR route depends solely on direct μ2 recognition rather than accessory adaptors. Abstract-level reagent species and exact donor provenance remain unresolved.
Propagation Review
Root cause: NO FAILURE NON CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:12121421
recognition of the FDNPVY signal is mediated by a binding site in the mu2-subunit that is distinct from the site for the more general YppØ sorting signal
GO:0050750 low-density lipoprotein particle receptor binding
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: Retain LDL-receptor-tail binding as a secondary interaction context.
Reason: PMID:12121421 directly reports μ2 binding to FDNPVY and YppΦ peptides at distinct sites. Preserve the rat-derived assertion without claiming that the human endogenous LDLR route depends solely on direct μ2 recognition rather than accessory adaptors. Abstract-level reagent species and exact donor provenance remain unresolved.
Propagation Review
Root cause: NO FAILURE NON CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:12121421
recognition of the FDNPVY signal is mediated by a binding site in the mu2-subunit that is distinct from the site for the more general YppØ sorting signal
GO:0065003 protein-containing complex assembly
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Retain the reported accessory contribution to complex assembly.
Reason: The cited reconstitution study reports that removal of AP50 impairs V-ATPase reassembly and that AP-2 restores it. This supports the existing broad assembly context, with in-vitro and reagent-species limits explicit. It does not establish ATP hydrolysis by ΞΌ2 or a second principal enzyme function.
Propagation Review
Root cause: NO FAILURE NON CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:7989329
The addition of purified AP-2 containing the AP50 polypeptide restores the ability of the V1 subunits to assemble with the V0 sector
GO:0065003 protein-containing complex assembly
TAS
PMID:7989329
Activity and in vitro reassembly of the coated vesicle (H+)-...
KEEP AS NON CORE
Summary: Retain the reported accessory contribution to complex assembly.
Reason: The cited reconstitution study reports that removal of AP50 impairs V-ATPase reassembly and that AP-2 restores it. This supports the existing broad assembly context, with in-vitro and reagent-species limits explicit. It does not establish ATP hydrolysis by ΞΌ2 or a second principal enzyme function.
Supporting Evidence:
PMID:7989329
The addition of purified AP-2 containing the AP50 polypeptide restores the ability of the V1 subunits to assemble with the V0 sector
GO:0070062 extracellular exosome
HDA
PMID:19056867
Large-scale proteomics and phosphoproteomics of urinary exos...
UNDECIDED
Summary: The extracellular-vesicle detection remains unresolved.
Reason: The urinary-exosome proteomics abstract does not expose the target-specific peptide/detection table. Intracellular coat function does not exclude extracellular-vesicle cargo, but it cannot establish this source-specific observation either. Preserve the HDA annotation without inferring a false localization from the paper's general scope.
GO:0072583 clathrin-dependent endocytosis
IBA
GO_REF:0000033
ACCEPT
Summary: AP2M1 contributes to clathrin-dependent endocytosis.
Reason: Cargo recognition, membrane-recruited AP-2 architecture and ΞΌ2-dependent uptake directly establish the conserved process. Human-cell evidence and species-limited recombinant/animal assays are distinguished in the notes. For the IBA, the actual endocytosis IBD is present and target self-support is valid; full topology was not reconstructed.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
PANTHER:PTN000242371 UNRESOLVED
The actual PAINT node assertion was inspected in the local family export; full tree/MSA placement was not reconstructed. Target self-inclusion and donor count are not defects.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
GO:0072583 clathrin-dependent endocytosis
IDA
PMID:23676497
Endocytosis of synaptic ADAM10 in neuronal plasticity and Al...
ACCEPT
Summary: AP2M1 contributes to clathrin-dependent endocytosis.
Reason: Cargo recognition, membrane-recruited AP-2 architecture and ΞΌ2-dependent uptake directly establish the conserved process. Human-cell evidence and species-limited recombinant/animal assays are distinguished in the notes. For the IBA, the actual endocytosis IBD is present and target self-support is valid; full topology was not reconstructed.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
GO:0072583 clathrin-dependent endocytosis
IDA
PMID:31104773
A Recurrent Missense Variant in AP2M1 Impairs Clathrin-Media...
ACCEPT
Summary: AP2M1 contributes to clathrin-dependent endocytosis.
Reason: Cargo recognition, membrane-recruited AP-2 architecture and ΞΌ2-dependent uptake directly establish the conserved process. Human-cell evidence and species-limited recombinant/animal assays are distinguished in the notes. For the IBA, the actual endocytosis IBD is present and target self-support is valid; full topology was not reconstructed.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
GO:0072583 clathrin-dependent endocytosis
NAS
PMID:29184887
Weak Molecular Interactions in Clathrin-Mediated Endocytosis...
ACCEPT
Summary: AP2M1 contributes to clathrin-dependent endocytosis.
Reason: Cargo recognition, membrane-recruited AP-2 architecture and ΞΌ2-dependent uptake directly establish the conserved process. Human-cell evidence and species-limited recombinant/animal assays are distinguished in the notes. For the IBA, the actual endocytosis IBD is present and target self-support is valid; full topology was not reconstructed.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
GO:0072583 clathrin-dependent endocytosis
TAS
PMID:12121421
The mu2 subunit of the clathrin adaptor AP-2 binds to FDNPVY...
ACCEPT
Summary: AP2M1 contributes to clathrin-dependent endocytosis.
Reason: Cargo recognition, membrane-recruited AP-2 architecture and ΞΌ2-dependent uptake directly establish the conserved process. Human-cell evidence and species-limited recombinant/animal assays are distinguished in the notes. For the IBA, the actual endocytosis IBD is present and target self-support is valid; full topology was not reconstructed.
Supporting Evidence:
PMID:19419997
We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
PMID:20603002
The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
PMID:31104773
Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
GO:0097494 regulation of vesicle size
IMP
PMID:25898166
CALM regulates clathrin-coated vesicle size and maturation b...
ACCEPT
Summary: AP-2 organization contributes to vesicle-size regulation.
Reason: The actual source shows that ΞΌ2 depletion changes coated-structure abundance and size, with rescue controls. The existing process annotation is consistent with coat organization; the directly demonstrated curvature sensor in this paper is CALM, not ΞΌ2.
Supporting Evidence:
PMID:25898166
AP2 depletion (of μ2) resulted in an ∼12-fold decrease in the number of endocytic CCPs/CCVs
GO:0098794 postsynapse
IEA
GO_REF:0000108
ACCEPT
Summary: Retain the postsynaptic cargo-retrieval compartment.
Reason: The inferred location follows the established postsynaptic receptor-internalization role, supported by the inspected adaptor review. This is an endocytic machinery pool rather than a claim that ΞΌ2 is a postsynaptic signaling receptor or scaffold.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
GO:0098884 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:31671891
Since AP-2 is not only involved in the sorting of synaptic vesicle proteins, but also regulates the surface expression of postsynaptic neurotransmitter receptors
GO:0098884 postsynaptic neurotransmitter receptor internalization
IBA
GO_REF:0000033
ACCEPT
Summary: Retain postsynaptic neurotransmitter-receptor internalization.
Reason: The inspected AP-2 sections of the cited review explicitly discuss postsynaptic receptor surface regulation as a cargo-sorting role. This is direct adaptor participation, not an annotation inferred solely from seizures. The actual PAINT IBD for this process is present; the exact rat experiments and full family topology were not independently reconstructed.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
PANTHER:PTN000242374 UNRESOLVED
The actual PAINT node assertion was inspected in the local family export; full tree/MSA placement was not reconstructed. Target self-inclusion and donor count are not defects.
Supporting Evidence:
PMID:31671891
Since AP-2 is not only involved in the sorting of synaptic vesicle proteins, but also regulates the surface expression of postsynaptic neurotransmitter receptors
GO:0098884 postsynaptic neurotransmitter receptor internalization
IEA
GO_REF:0000107
ACCEPT
Summary: Retain postsynaptic neurotransmitter-receptor internalization.
Reason: The inspected AP-2 sections of the cited review explicitly discuss postsynaptic receptor surface regulation as a cargo-sorting role. This is direct adaptor participation, not an annotation inferred solely from seizures. The actual PAINT IBD for this process is present; the exact rat experiments and full family topology were not independently reconstructed.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:31671891
Since AP-2 is not only involved in the sorting of synaptic vesicle proteins, but also regulates the surface expression of postsynaptic neurotransmitter receptors
GO:0098884 postsynaptic neurotransmitter receptor internalization
NAS
PMID:31671891
Endocytic Adaptor Proteins in Health and Disease: Lessons fr...
ACCEPT
Summary: Retain postsynaptic neurotransmitter-receptor internalization.
Reason: The inspected AP-2 sections of the cited review explicitly discuss postsynaptic receptor surface regulation as a cargo-sorting role. This is direct adaptor participation, not an annotation inferred solely from seizures. The actual PAINT IBD for this process is present; the exact rat experiments and full family topology were not independently reconstructed.
Supporting Evidence:
PMID:31671891
Since AP-2 is not only involved in the sorting of synaptic vesicle proteins, but also regulates the surface expression of postsynaptic neurotransmitter receptors
GO:0098894 extrinsic component of presynaptic endocytic zone membrane
IEA
GO_REF:0000107
ACCEPT
Summary: Retain the peripheral presynaptic endocytic coat pool.
Reason: The conserved AP-2 complex retrieves synaptic membrane cargo from the cytoplasmic surface. The existing rat transfer fits this peripheral endocytic role; it does not imply an integral membrane or neurotransmitter-release activity.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:11102472
Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.
PMID:31671891
ΞΌ2-subunit also interacts with C2 domains as present in Synaptotagmin1 [11] or Otoferlin [12].
GO:0098978 glutamatergic synapse
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Retain glutamatergic-synapse localization as a neuronal context.
Reason: The electronic rat transfer is biologically compatible with postsynaptic receptor trafficking, but this named synapse type is a context of a broadly expressed cargo adaptor rather than its defining location. Exact donor localization experiments were not independently reconstructed; no contradictory evidence or taxon restriction is claimed.
Propagation Review
Root cause: NO FAILURE NON CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:31671891
the internalization of AMPA-type glutamate receptors by Synaptotagmin3 in addition to AP-2 [76].
GO:0099003 vesicle-mediated transport in synapse
IEA
GO_REF:0000117
ACCEPT
Summary: Retain synaptic vesicle-mediated transport.
Reason: The broad automated process fits AP-2-mediated retrieval and sorting at synapses. The specific ARBA rule is not reconstructed, and acceptance does not give ΞΌ2 the fusion or neurotransmitter-release chemistry of its cargo.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
ARBA:ARBA00027531 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:11102472
Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.
PMID:31671891
ΞΌ2-subunit also interacts with C2 domains as present in Synaptotagmin1 [11] or Otoferlin [12].
GO:1900244 positive regulation of synaptic vesicle endocytosis
IEA
GO_REF:0000107
ACCEPT
Summary: Retain promotion of synaptic-vesicle endocytosis.
Reason: The rat-derived regulatory annotation is consistent with the conserved coat-assembly/cargo-sorting contribution to retrieval. It does not assign ΞΌ2 an enzymatic regulatory step or infer a new disease process from perturbation alone.
Propagation Review
Root cause: NO FAILURE CORE
Sources checked:
UniProtKB:P84092 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
ensembl:ENSRNOP00000054900 UNRESOLVED
The exact donor experiment or electronic mapping was not reconstructed. Separately inspected target evidence is described in the rationale; no failure is inferred from donor count.
Supporting Evidence:
PMID:11102472
Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.
PMID:31671891
ΞΌ2-subunit also interacts with C2 domains as present in Synaptotagmin1 [11] or Otoferlin [12].
GO:1903077 negative regulation of protein localization to plasma membrane
IMP
PMID:19581412
Quantitative proteomics identifies a Dab2/integrin module re...
ACCEPT
Summary: AP-2 reduces cargo residence at the plasma membrane by internalization.
Reason: The actual source includes ΞΌ2 depletion and reports increased surface integrins. ΞΌ2 supplies the internalizing adaptor machinery; the study's focus on DAB2 does not negate its explicitly assayed AP-2 controls. This is the existing removal/localization process, not an inferred transcriptional repression mechanism.
Supporting Evidence:
PMID:19581412
AP2 or clathrin depletion significantly increases surface Ξ±1, Ξ±5, and Ξ²1.

Core Functions

AP2M1 recognizes cytoplasmic sorting sequences in cargo proteins and, as the medium subunit of AP-2, contributes to their coupling to the clathrin coat. Membrane-dependent rearrangement and regulated Thr156 phosphorylation coordinate cargo capture and coat maturation. This supports receptor uptake, endocytic budding and synaptic cargo retrieval; it does not make isolated ΞΌ2 the complete clathrin adaptor or a kinase.

Supporting Evidence:
  • PMID:19419997
    We also identify tyrosine-995 of KIAA0319 as a critical amino acid required for the interaction with AP-2 and subsequent internalization.
  • PMID:20603002
    The AP2 adaptor complex (alpha, beta2, sigma2, and mu2 subunits) crosslinks the endocytic clathrin scaffold to PtdIns4,5P(2)-containing membranes and transmembrane protein cargo.
  • PMID:31104773
    Functional complementation of both the ΞΌ-subunit carrying the p.Arg170Trp variant in human cells and astrocytes derived from AP-2ΞΌ conditional knockout mice revealed a significant impairment of CME of transferrin.
  • PMID:31430451
    ΞΌ2Thr156 phosphorylation favors a new, cargo-bound conformation of AP2 and simultaneously creates a binding platform for the endocytic NECAP proteins
  • PMID:11102472
    Microinjection of these peptides into squid giant presynaptic terminals reversibly blocked synaptic transmission and inhibited synaptic vesicle endocytosis by preventing coated pit formation at the plasma membrane.

References

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Suggested Questions for Experts

Q: Which target-specific assay establishes AP2M1 residence on mature lysosomal or endolysosomal membranes, as distinct from its coat on carriers carrying cargo to those destinations?

Q: Where does human AP2M1 lie relative to the GO:0006896 IBD and same-term IRD in the complete PAINT tree, and is there target-specific retention or loss evidence?

Q: Which exact interaction experiments support the unresolved generic screen pairs, and which distinguish direct sorting-signal recognition from coassembly or indirect association?

Q: How do the two human AP2M1 isoforms differ in cargo recognition and regulation under endogenous expression, beyond both tested isoforms binding HCV core?

Q: Would a dedicated endocytic sorting-motif binding term better distinguish μ2 recognition of cytoplasmic YxxΦ cargo tails from other localization-signal recognition? The current GO:0005048 definition already covers short protein-localization signals, so the present source-specific term remains applicable.

Suggested Experiments

Experiment: Compare matched human AP2M1 isoforms and cargo-site mutants in AP-2 reconstitution and endogenous rescue, measuring motif binding, coat recruitment and receptor uptake separately.

πŸ“š Additional Documentation

Notes

(AP2M1-notes.md)

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