CACNB2

UniProt ID: Q08289
Organism: Homo sapiens
Review Status: COMPLETE
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Gene Description

CACNB2 encodes the voltage-dependent L-type calcium channel subunit beta-2 (CaVbeta2), an essential cytosolic auxiliary subunit of high-voltage-activated Ca2+ channels. The protein contains a conserved SH3-HOOK-guanylate kinase (GK) module that binds the alpha-interaction domain (AID) in the I-II linker of the pore-forming alpha1 subunit with nanomolar affinity. CaVbeta2 promotes proper channel trafficking to the plasma membrane and modulates voltage-dependent activation/inactivation kinetics of CaV1.x/CaV2.x channels. Multiple splice variants (beta2a-beta2e) differ in their N-termini, with beta2a being palmitoylated for membrane anchoring and beta2e capable of nuclear translocation. CaVbeta2 also mediates beta-adrenergic regulation of cardiac L-type channels through interaction with the RGK GTPase Rad.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0006816 calcium ion transport
IBA
GO_REF:0000033
ACCEPT
Summary: CaVbeta2 is an auxiliary subunit that promotes calcium channel trafficking and modulates channel gating. PMID:1309651 demonstrated that beta2 subunit coexpression is obligatory for functional calcium channel activity, and that it increases peak calcium current. The protein enables calcium ion transport by regulating the alpha1 pore-forming subunit.
Reason: IBA annotations for calcium ion transport are appropriate. While CaVbeta2 is not itself the ion pore, it is essential for functional calcium channel expression and directly contributes to calcium transport by enabling channel trafficking to the membrane and modulating channel gating properties [PMID:1309651].
Supporting Evidence:
PMID:1309651
the beta 2 subunit appears to serve an obligatory function
GO:0005891 voltage-gated calcium channel complex
IBA
GO_REF:0000033
ACCEPT
Summary: CaVbeta2 is a core component of the voltage-gated calcium channel complex, binding the alpha1 subunit AID with nanomolar affinity (6-20 nM) via its GK domain. Structural studies confirm the SH3-GK architecture positions CaVbeta between the AID and VSD-II of the alpha1 subunit.
Reason: This is a well-established core function. CaVbeta2 is part of the calcium channel complex in 1:1:1:1 stoichiometry with alpha1, alpha2delta, and gamma subunits [UniProt CC, PMID:1309651]. ComplexPortal entries (CPX-3195 and others) document CACNB2 in multiple CaV1.x channel complexes.
Supporting Evidence:
PMID:1309651
coexpressed with the beta (beta 2) and the alpha 2 (alpha 2b) subunits
GO:0007268 chemical synaptic transmission
IBA
GO_REF:0000033
KEEP AS NON CORE
Summary: Calcium channels play essential roles in neurotransmitter release at presynaptic terminals. CaVbeta2 is expressed in neurons and contributes to calcium channel function that underlies synaptic transmission.
Reason: While L-type calcium channels contribute to synaptic transmission, this is more of a downstream physiological consequence rather than a direct molecular function of CaVbeta2. The primary role of CaVbeta2 is channel assembly and modulation; synaptic transmission is a higher-level process. This annotation is acceptable but represents a non-core function.
Supporting Evidence:
PMID:8494331
The predicted amino acid sequence of this clone shows a high degree of homology to the beta subunit of calcium channel complexes
GO:0098793 presynapse
IEA
GO_REF:0000108
KEEP AS NON CORE
Summary: Automated inference based on inter-ontology links. CaVbeta2 would be present at presynaptic terminals as part of calcium channel complexes that regulate neurotransmitter release.
Reason: Presynaptic localization is consistent with calcium channel function in neurons, but this is based on logical inference rather than direct experimental evidence for CACNB2 specifically. The broader cellular component annotation (voltage-gated calcium channel complex) is more directly supported.
Supporting Evidence:
file:human/CACNB2/CACNB2-uniprot.txt
Expressed in all tissues
GO:0099509 regulation of presynaptic cytosolic calcium ion concentration
IEA
GO_REF:0000108
KEEP AS NON CORE
Summary: Automated inference suggesting CaVbeta2 regulates presynaptic calcium levels through its role in calcium channel function.
Reason: This is a reasonable inference but represents a tissue-specific downstream effect rather than a core function. The annotation to GO:0051928 (positive regulation of calcium ion transport) from PMID:1309651 is more directly supported experimentally.
Supporting Evidence:
GO_REF:0000108
logical inference based on inter-ontology links
GO:0005245 voltage-gated calcium channel activity
IEA
GO_REF:0000002
MODIFY
Summary: InterPro-based inference. CaVbeta2 does not itself have ion channel activity but is essential for functional channel activity when coexpressed with alpha1.
Reason: CaVbeta2 is an auxiliary subunit, not the pore-forming subunit. It does not have intrinsic channel activity. The annotation should reflect its role as a regulator/auxiliary rather than having the activity itself. However, there is an IDA annotation to the same term from PMID:1309651 which is more nuanced.
GO:0005262 calcium channel activity
IEA
GO_REF:0000043
MODIFY
Summary: UniProt keyword-based inference. Similar issue as GO:0005245 - CaVbeta2 enables but does not itself possess calcium channel activity.
Reason: This annotation conflates the auxiliary subunit with the pore. CaVbeta2 is essential for channel function but the channel activity resides in the alpha1 subunit. A more appropriate annotation would be to the channel complex or a regulatory function.
GO:0005891 voltage-gated calcium channel complex
IEA
GO_REF:0000120
ACCEPT
Summary: Combined automated annotation confirming CaVbeta2 is part of the voltage-gated calcium channel complex.
Reason: This correctly annotates CaVbeta2 as a component of the channel complex, which is well-supported by multiple lines of evidence including structural data and biochemical studies [PMID:1309651, UniProt].
Supporting Evidence:
PMID:1309651
coexpression with the alpha 2b subunit
GO:0006811 monoatomic ion transport
IEA
GO_REF:0000043
ACCEPT
Summary: General ion transport term from UniProt keyword mapping.
Reason: This is a parent term of calcium ion transport (GO:0006816) and is acceptable as a broader annotation. It correctly reflects CaVbeta2's role in enabling calcium channel function.
Supporting Evidence:
file:human/CACNB2/CACNB2-uniprot.txt
Calcium transport keyword
GO:0006816 calcium ion transport
IEA
GO_REF:0000043
ACCEPT
Summary: UniProt keyword-based inference for calcium ion transport, consistent with the IBA annotation to the same term.
Reason: Consistent with the IBA annotation and well-supported by the role of CaVbeta2 in enabling functional calcium channel expression [PMID:1309651].
Supporting Evidence:
file:human/CACNB2/CACNB2-uniprot.txt
Calcium channel; Calcium transport keywords
GO:0034220 monoatomic ion transmembrane transport
IEA
GO_REF:0000043
ACCEPT
Summary: UniProt keyword-based inference for ion transmembrane transport.
Reason: Appropriate parent term reflecting CaVbeta2's role in enabling calcium flux across the plasma membrane through the channel complex.
Supporting Evidence:
file:human/CACNB2/CACNB2-uniprot.txt
Ion transport keyword
GO:0034702 monoatomic ion channel complex
IEA
GO_REF:0000043
ACCEPT
Summary: UniProt keyword-based inference for ion channel complex membership.
Reason: Correct parent term of GO:0005891 (voltage-gated calcium channel complex). CaVbeta2 is indeed part of an ion channel complex.
Supporting Evidence:
file:human/CACNB2/CACNB2-uniprot.txt
Ion channel keyword
GO:0042383 sarcolemma
IEA
GO_REF:0000044
ACCEPT
Summary: UniProt subcellular location-based inference for sarcolemma localization. UniProt explicitly states CaVbeta2 localizes to sarcolemma in cardiac muscle.
Reason: This is directly supported by UniProt subcellular location annotation stating "Cell membrane, sarcolemma; Peripheral membrane protein; Cytoplasmic side." CaVbeta2 associates with L-type channels in cardiac T-tubule/sarcolemma.
Supporting Evidence:
file:human/CACNB2/CACNB2-uniprot.txt
Cell membrane, sarcolemma; Peripheral membrane protein; Cytoplasmic side
GO:0070588 calcium ion transmembrane transport
IEA
GO_REF:0000120
ACCEPT
Summary: Combined automated inference for calcium ion transmembrane transport.
Reason: Consistent with CaVbeta2's essential role in enabling functional calcium channel activity that mediates transmembrane calcium flux.
Supporting Evidence:
PMID:1309651
directs the recombinant expression of a dihydropyridine-sensitive L-type Ca2+ channel
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
MARK AS OVER ANNOTATED
Summary: High-throughput protein interactome study (HuRI) identified binary protein interactions involving CACNB2.
Reason: While technically correct, "protein binding" is uninformative. CaVbeta2 binds specific proteins including the alpha1 subunit AID, RRAD, CAMK2A, and others. More specific binding annotations would be more informative.
Supporting Evidence:
PMID:32296183
With approximately 53,000 protein-protein interactions, HuRI has approximately four times as many such interactions as there are high-quality curated interactions from small-scale studies
GO:0051015 actin filament binding
IEA
GO_REF:0000107
ACCEPT
Summary: Ensembl Compara ortholog transfer suggesting actin filament binding. PMID:25533460 demonstrated direct CaVbeta-actin interaction.
Reason: PMID:25533460 provides experimental evidence for direct CaVbeta-actin interaction using co-sedimentation and FRET experiments. The study showed CaVbeta2 distributes along actin filaments in cardiomyocytes.
Supporting Evidence:
PMID:25533460
we uncover a direct interaction between CaVbeta and actin filaments
GO:0098684 photoreceptor ribbon synapse
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Ensembl Compara ortholog transfer suggesting localization to photoreceptor ribbon synapse. The beta2e isoform is known to predominate in retina.
Reason: The beta2e isoform is specifically expressed in retina, making this a plausible but tissue-specific localization. The deep research file notes "beta2e predominates in retina." However, this is based on ortholog transfer rather than direct experimental evidence in human.
Supporting Evidence:
file:human/CACNB2/CACNB2-deep-research-falcon.md
beta2e predominates in retina
GO:0099626 voltage-gated calcium channel activity involved in regulation of presynaptic cytosolic calcium levels
IEA
GO_REF:0000107
MODIFY
Summary: Ensembl Compara ortholog transfer for presynaptic calcium channel activity.
Reason: CaVbeta2 is not the pore and does not have intrinsic channel activity. This annotation conflates the auxiliary subunit with the functional activity. The annotation should be to the complex or a regulatory function.
GO:0070588 calcium ion transmembrane transport
NAS
PMID:22982493
Ca(V)1.1: The atypical prototypical voltage-gated Ca²⁺ chann...
ACCEPT
Summary: Non-traceable author statement from a review about CaV1.1 channels noting the general role of beta subunits in calcium channel function.
Reason: While NAS is a weaker evidence code, the annotation is consistent with the established role of CaVbeta2. PMID:22982493 is a review discussing calcium channel function.
Supporting Evidence:
PMID:22982493
Ca(V)1.1 is the prototype for the other nine known Ca(V) channel isoforms
GO:0030315 T-tubule
NAS
PMID:27273705
Voltage-gated calcium channels and their auxiliary subunits:...
ACCEPT
Summary: Non-traceable author statement from Dolphin review on voltage-gated calcium channels and their auxiliary subunits, indicating T-tubule localization.
Reason: T-tubule localization is consistent with the role of L-type calcium channels in excitation-contraction coupling in cardiac and skeletal muscle. The review discusses calcium channel physiology and pathophysiology.
Supporting Evidence:
PMID:27273705
Voltage-gated calcium channels and their auxiliary subunits
GO:0045933 positive regulation of muscle contraction
NAS
PMID:22982493
Ca(V)1.1: The atypical prototypical voltage-gated Ca²⁺ chann...
KEEP AS NON CORE
Summary: CaVbeta2 contributes to calcium influx that triggers muscle contraction.
Reason: This is a downstream physiological effect. CaVbeta2 enables calcium channel function, and calcium influx is required for muscle contraction, but this is an indirect effect rather than a direct molecular function.
Supporting Evidence:
PMID:22982493
Ca(V)1.1 is the prototype for the other nine known Ca(V) channel isoforms
GO:0045933 positive regulation of muscle contraction
NAS
PMID:34072954
Mechanisms and Regulation of Cardiac Ca(V)1.2 Trafficking.
KEEP AS NON CORE
Summary: Review on CaV1.2 trafficking mechanisms discussing the role of beta subunits in cardiac calcium channel function.
Reason: Consistent with the other NAS annotation to this term. CaVbeta2 contributes to cardiac calcium channel function which underlies muscle contraction, but this is an indirect downstream effect.
Supporting Evidence:
PMID:34072954
The number and activity of functional CaV1.2 channels at the t-tubule dyads dictates the amplitude of the Ca2+ influx
GO:0070588 calcium ion transmembrane transport
NAS
PMID:27273705
Voltage-gated calcium channels and their auxiliary subunits:...
ACCEPT
Summary: Non-traceable author statement from Dolphin review on voltage-gated calcium channels.
Reason: Consistent with other annotations and the established role of CaVbeta2 in enabling functional calcium channel activity.
Supporting Evidence:
PMID:27273705
For the CaV 1 and CaV 2 channels, their ability to reach their required destinations in the cell membrane
GO:0005515 protein binding
IPI
PMID:28130356
A Novel Human CAMK2A Mutation Disrupts Dendritic Morphology ...
MARK AS OVER ANNOTATED
Summary: Study on CAMK2A mutations and ASD-related behaviors identified interaction between CaMKIIalpha and CACNB2. The E183V mutation reduced CaMKIIalpha binding to calcium channel subunits.
Reason: The study identifies specific interaction with CAMK2A but the annotation is to generic "protein binding." UniProt confirms CACNB2 interacts with CAMK2A. A more specific annotation would be preferable.
Supporting Evidence:
PMID:28130356
The E183V mutation also reduces CaMKIIalpha binding to established ASD-linked proteins, such as Shank3 and subunits of l-type calcium channels
GO:0005891 voltage-gated calcium channel complex
IDA
PMID:17224476
Loss-of-function mutations in the cardiac calcium channel un...
ACCEPT
Summary: Antzelevitch et al. studied loss-of-function mutations in cardiac calcium channels including CACNB2 (S481L mutation). CHO-K1 cells were cotransfected with CACNB2b and other channel subunits for electrophysiology studies.
Reason: The study directly demonstrates CaVbeta2b is part of the cardiac L-type calcium channel complex through heterologous expression and electrophysiology. Mutations in CACNB2 affect channel function.
Supporting Evidence:
PMID:17224476
CHO-K1 cells were cotransfected with cDNAs encoding wild-type or mutant CACNB2b
GO:0051015 actin filament binding
ISS
PMID:25533460
Direct interaction of CaVΞ² with actin up-regulates L-type ca...
ACCEPT
Summary: Study demonstrated direct CaVbeta-actin interaction using co-sedimentation assays and FRET experiments in HL-1 cardiomyocytes. Single-molecule localization showed CaVbeta2 distributing along actin filaments.
Reason: This is strong experimental evidence for actin filament binding. The study used multiple methods to demonstrate the interaction and showed functional relevance for channel trafficking.
Supporting Evidence:
PMID:25533460
using co-sedimentation assays and FRET experiments, we uncover a direct interaction between CaVbeta and actin filaments
GO:0072659 protein localization to plasma membrane
ISS
PMID:25533460
Direct interaction of CaVΞ² with actin up-regulates L-type ca...
ACCEPT
Summary: The study demonstrated that CaVbeta promotes anterograde trafficking of L-type channels by anchoring them to actin filaments en route to the plasma membrane.
Reason: The study provides a model where CaVbeta promotes channel trafficking to the membrane via actin interaction. This is consistent with the known role of CaVbeta2 in channel trafficking.
Supporting Evidence:
PMID:25533460
CaVbeta promotes anterograde trafficking of the L-type channels by anchoring them to actin filaments in their itinerary to the plasma membrane
GO:0086007 voltage-gated calcium channel activity involved in cardiac muscle cell action potential
IMP
PMID:17224476
Loss-of-function mutations in the cardiac calcium channel un...
MODIFY
Summary: Study identified CACNB2 mutations in Brugada syndrome patients with cardiac arrhythmias. The S481L mutation caused loss-of-function affecting cardiac action potentials.
Reason: CaVbeta2 does not have intrinsic channel activity - it is an auxiliary subunit. The annotation should reflect its role in modulating channel function. The correct annotation would be to a regulatory function or the channel complex.
Supporting Evidence:
PMID:17224476
loss-of-function missense mutations in CACNA1C (A39V and G490R) and CACNB2 (S481L) encoding the alpha1- and beta2b-subunits
GO:0086045 membrane depolarization during AV node cell action potential
IMP
PMID:17224476
Loss-of-function mutations in the cardiac calcium channel un...
KEEP AS NON CORE
Summary: Study on Brugada syndrome with CACNB2 mutations affecting cardiac conduction.
Reason: This is a specific cardiac electrophysiology phenotype associated with CACNB2 mutations. While the annotation captures the disease phenotype, membrane depolarization is a downstream effect of calcium channel function rather than a direct function of CaVbeta2.
Supporting Evidence:
PMID:17224476
Loss-of-function mutations in the cardiac calcium channel underlie a new clinical entity characterized by ST-segment elevation
GO:0086056 voltage-gated calcium channel activity involved in AV node cell action potential
IMP
PMID:17224476
Loss-of-function mutations in the cardiac calcium channel un...
MODIFY
Summary: Study linking CACNB2 mutations to cardiac arrhythmias affecting AV node function.
Reason: Same issue as GO:0086007 - CaVbeta2 is not the channel pore and does not have intrinsic channel activity. It modulates channel function as an auxiliary subunit.
Supporting Evidence:
PMID:17224476
ST-segment elevation, short QT intervals, and sudden cardiac death
GO:0086091 regulation of heart rate by cardiac conduction
IMP
PMID:17224476
Loss-of-function mutations in the cardiac calcium channel un...
KEEP AS NON CORE
Summary: CACNB2 mutations cause cardiac arrhythmias affecting heart rate and conduction.
Reason: This is a downstream physiological effect of calcium channel dysfunction. CaVbeta2 contributes to cardiac calcium channel function which is essential for proper conduction and heart rate, but this is an indirect effect.
Supporting Evidence:
PMID:17224476
Brugada syndrome phenotype is combined with shorter-than-normal QT intervals
GO:0098912 membrane depolarization during atrial cardiac muscle cell action potential
IMP
PMID:17224476
Loss-of-function mutations in the cardiac calcium channel un...
KEEP AS NON CORE
Summary: Brugada syndrome study showing effects of CACNB2 mutations on atrial action potentials.
Reason: Similar to other cardiac electrophysiology annotations - this is a downstream consequence of calcium channel function rather than a direct molecular function.
Supporting Evidence:
PMID:17224476
cardiac calcium channel underlie a new clinical entity
GO:1904879 positive regulation of calcium ion transmembrane transport via high voltage-gated calcium channel
ISS
PMID:25533460
Direct interaction of CaVΞ² with actin up-regulates L-type ca...
ACCEPT
Summary: Study demonstrated that CaVbeta2 overexpression increases L-type current by increasing channels in the plasma membrane.
Reason: This accurately reflects the role of CaVbeta2 in promoting calcium channel trafficking and increasing calcium current. The study showed CaVbeta2-N3 overexpression induced an increase in L-type current.
Supporting Evidence:
PMID:25533460
Overexpression of CaVbeta2-N3 in HL-1 cells induces an increase in L-type current without altering voltage-dependent activation, thus reflecting an increased number of channels in the plasma membrane
GO:1990454 L-type voltage-gated calcium channel complex
IDA
PMID:1309651
Structure and functional expression of alpha 1, alpha 2, and...
ACCEPT
Summary: Williams et al. demonstrated functional expression of L-type calcium channels requiring coexpression of beta2 with alpha1D and alpha2b subunits in Xenopus oocytes.
Reason: This is a core annotation strongly supported by the paper. The study showed beta2 is required for functional L-type channel expression and is part of the channel complex.
Supporting Evidence:
PMID:1309651
directs the recombinant expression of a dihydropyridine-sensitive L-type Ca2+ channel when coexpressed with the beta (beta 2) and the alpha 2 (alpha 2b) subunits
GO:0008331 high voltage-gated calcium channel activity
IDA
PMID:1309651
Structure and functional expression of alpha 1, alpha 2, and...
MODIFY
Summary: The study expressed a human neuronal high-voltage-activated calcium channel requiring beta2 for function.
Reason: CaVbeta2 enables high-voltage-gated calcium channel function but does not itself have channel activity. The alpha1 subunit forms the pore. The annotation should reflect the auxiliary/regulatory role.
Supporting Evidence:
PMID:1309651
the beta 2 subunit appears to serve an obligatory function
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-5577213
ACCEPT
Summary: Reactome pathway entry for LTCC multimer transporting calcium from extracellular region to cytosol places CaVbeta2 at the plasma membrane.
Reason: CaVbeta2 localizes to the plasma membrane as part of the L-type calcium channel complex. UniProt also states sarcolemma/plasma membrane localization.
Supporting Evidence:
file:human/CACNB2/CACNB2-uniprot.txt
Cell membrane, sarcolemma; Peripheral membrane protein; Cytoplasmic side
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-9614031
ACCEPT
Summary: Reactome pathway for Class IV antihypertensives binding LTCC multimer.
Reason: Same as above - plasma membrane localization is well-established for CaVbeta2.
Supporting Evidence:
Reactome:R-HSA-9614031
Class IV antihypertensives bind LTCC multimer
GO:0051928 positive regulation of calcium ion transport
IDA
PMID:1309651
Structure and functional expression of alpha 1, alpha 2, and...
ACCEPT
Summary: Williams et al. showed that beta2 coexpression is required for functional calcium channel expression and increases peak calcium current.
Reason: This is a core function. The study demonstrated that "the beta 2 subunit appears to serve an obligatory function" and that channels with beta2 have enhanced calcium current compared to alpha1 alone (which showed no current).
Supporting Evidence:
PMID:1309651
Expression of the alpha 1D subunit alone, or coexpression with the alpha 2b subunit, did not elicit functional Ca2+ channel activity. Thus, the beta 2 subunit appears to serve an obligatory function
GO:0005245 voltage-gated calcium channel activity
IDA
PMID:1309651
Structure and functional expression of alpha 1, alpha 2, and...
MODIFY
Summary: Study demonstrated voltage-gated calcium channel activity requires beta2.
Reason: While the study demonstrates CaVbeta2 is essential for channel function, CaVbeta2 itself does not have ion channel activity - the alpha1 subunit forms the pore. The annotation should reflect the regulatory role.
Supporting Evidence:
PMID:1309651
the beta 2 subunit appears to serve an obligatory function
GO:0070509 calcium ion import
IDA
PMID:1309651
Structure and functional expression of alpha 1, alpha 2, and...
ACCEPT
Summary: Study demonstrated functional calcium influx through channels containing beta2.
Reason: CaVbeta2 enables calcium ion import by being essential for functional channel expression. The term correctly reflects the biological process.
Supporting Evidence:
PMID:1309651
dihydropyridine-sensitive L-type Ca2+ channel
GO:0005515 protein binding
IPI
PMID:17525370
Dominant negative suppression of Rad leads to QT prolongatio...
MODIFY
Summary: Study demonstrated CACNB2 interaction with RRAD (Ras associated with diabetes), an RGK GTPase that regulates L-type calcium channel function.
Reason: The study identifies a specific interaction with RRAD that is functionally important for channel trafficking. "Protein binding" is too generic; a more specific annotation would be preferable.
Proposed replacements: protein binding
Supporting Evidence:
PMID:17525370
upregulation of L-type Ca2+ channel expression in the plasma membrane
file:human/CACNB2/CACNB2-uniprot.txt
Interacts with RRAD; interaction may be involved in beta-adrenergic regulation of heart rate and contractile force
GO:0007528 neuromuscular junction development
TAS
PMID:8494331
Cloning and characterization of a Lambert-Eaton myasthenic s...
KEEP AS NON CORE
Summary: Rosenfeld et al. identified CACNB2 (MysB) as a Lambert-Eaton myasthenic syndrome antigen with homology to calcium channel beta subunits. The syndrome affects the neuromuscular junction.
Reason: While CaVbeta2 is implicated in Lambert-Eaton syndrome affecting the NMJ, this annotation represents disease involvement rather than a normal developmental function. The study cloned the antigen but did not directly demonstrate a role in NMJ development.
Supporting Evidence:
PMID:8494331
Lambert-Eaton myasthenic syndrome is a paraneoplastic neuromuscular disorder in which an immune response directed against a small-cell lung tumor crossreacts with antigens in the neuromuscular junction
GO:0005262 calcium channel activity
NAS
PMID:9594024
Effects of temperature on human L-type cardiac Ca2+ channels...
MODIFY
Summary: Allen and Mikala studied temperature effects on L-type cardiac calcium channels expressed in Xenopus oocytes using different beta subunit isoforms.
Reason: CaVbeta2 is an auxiliary subunit that modulates channel properties but does not itself have channel activity. The alpha1 subunit forms the pore.
Supporting Evidence:
PMID:9594024
Using instead a different CaCh beta subunit isoform, beta2c
GO:0005886 plasma membrane
NAS
PMID:9594024
Effects of temperature on human L-type cardiac Ca2+ channels...
ACCEPT
Summary: Study expressed calcium channels at the plasma membrane of Xenopus oocytes using various subunit combinations.
Reason: Plasma membrane localization is well-established and consistent with the function of CaVbeta2 as part of the calcium channel complex at the cell surface.
Supporting Evidence:
PMID:9594024
cloned human cardiac CaChs expressed in Xenopus oocytes

Core Functions

CaVbeta2 is an essential auxiliary subunit of voltage-gated calcium channel complexes. It binds the alpha1 subunit AID with nanomolar affinity (6-20 nM) via its GK domain. PMID:1309651 demonstrated that beta2 is obligatory for functional channel expression and promotes calcium ion transport by enabling channel trafficking to the plasma membrane and modulating channel gating.

Supporting Evidence:
  • PMID:1309651
    the beta 2 subunit appears to serve an obligatory function
  • file:human/CACNB2/CACNB2-deep-research-falcon.md
    CaVbeta2 is a non-enzymatic, cytosolic auxiliary subunit that binds the alpha1-subunit AID to promote proper channel trafficking to the plasma membrane and to tune voltage-dependent activation/inactivation

PMID:25533460 demonstrated direct CaVbeta-actin interaction using co-sedimentation and FRET experiments. This interaction promotes channel trafficking to the plasma membrane, representing a novel mechanism for beta subunit function.

Molecular Function:
actin filament binding

References

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Suggested Questions for Experts

Q: What is the relative contribution of different CaVbeta2 isoforms (beta2a-e) to channel function in different tissues? The isoforms differ in N-terminal sequences affecting membrane association (palmitoylation for beta2a, nuclear translocation for beta2e). Understanding tissue-specific isoform functions could clarify disease mechanisms.

Q: How does the CaVbeta2-RRAD interaction modulate beta-adrenergic regulation of cardiac calcium channels at the molecular level? Recent work (PMID:36424916) showed Rad phosphorylation releases CaVbeta-mediated inhibition. The structural basis of this regulation is not fully understood.

Suggested Experiments

Experiment: Cryo-EM structural determination of full L-type calcium channel complex with different CaVbeta2 isoforms to understand isoform-specific contributions. While structures exist, comparing different beta2 isoforms could reveal how N-terminal differences affect channel architecture and gating.

Experiment: Live-cell imaging of CaVbeta2e nuclear translocation dynamics and identification of nuclear target genes. The beta2e isoform can translocate to the nucleus and regulate gene expression (Frontiers in Physiology 2025), but the target genes are not well characterized.

Deep Research

Falcon

(CACNB2-deep-research-falcon.md)

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