CR1

UniProt ID: P17927
Organism: Homo sapiens
Review Status: COMPLETE
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Gene Description

CR1 encodes complement receptor type 1/CD35, a large single-pass membrane receptor composed of complement-control protein repeat domains and expressed on erythrocytes, monocytes/macrophages, granulocytes, B cells, subsets of T cells, follicular dendritic cells, and other immune-associated cells. CR1 binds complement-opsonized ligands, especially C3b and C4b, allowing immune adherence and clearance of immune complexes and particles from the circulation. CR1 also regulates complement by accelerating decay of classical and alternative pathway convertases and acting as a factor I cofactor for cleavage of C3b and C4b. In specific immune-cell contexts, CR1 engagement can modulate T-cell, B-cell, and regulatory T-cell responses, while complement-pathway interactions with C1q, MBL, ficolins, and microbial ligands broaden its immune-recognition roles.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0005576 extracellular region
IBA
GO_REF:0000033
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0005886 plasma membrane
IBA
GO_REF:0000033
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0045959 negative regulation of complement activation, classical pathway
IBA
GO_REF:0000033
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0002456 T cell mediated immunity
IBA
GO_REF:0000033
KEEP AS NON CORE
Summary: This adaptive immune-cell annotation is supported or plausible but represents T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b receptor/cofactor role.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0001848 complement binding
IEA
GO_REF:0000117
ACCEPT
Summary: CR1 directly binds complement ligands, especially C3b and C4b, and functions as their cell-surface receptor.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0004875 complement receptor activity
IEA
GO_REF:0000117
ACCEPT
Summary: CR1 directly binds complement ligands, especially C3b and C4b, and functions as their cell-surface receptor.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0005886 plasma membrane
IEA
GO_REF:0000117
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0006957 complement activation, alternative pathway
IEA
GO_REF:0000117
ACCEPT
Summary: This complement activation/MAC annotation is supported in specific B-cell complement deposition contexts while remaining part of CR1 complement receptor biology.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0016020 membrane
IEA
GO_REF:0000044
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0030449 regulation of complement activation
IEA
GO_REF:0000117
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:1903659 regulation of complement-dependent cytotoxicity
IEA
GO_REF:0000117
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0005515 protein binding
IPI
PMID:23460739
Deciphering complement receptor type 1 interactions with rec...
MODIFY
Summary: The MBL/ficolin/C1q interaction evidence supports complement-recognition ligand binding, but generic protein binding is too broad.
Reason: Replace generic protein binding with complement binding for CR1 interactions with recognition proteins of the lectin/classical complement pathways.
Proposed replacements: complement binding
GO:0005515 protein binding
IPI
PMID:29563915
C1q and Mannose-Binding Lectin Interact with CR1 in the Same...
MODIFY
Summary: The MBL/ficolin/C1q interaction evidence supports complement-recognition ligand binding, but generic protein binding is too broad.
Reason: Replace generic protein binding with complement binding for CR1 interactions with recognition proteins of the lectin/classical complement pathways.
Proposed replacements: complement binding
GO:0005886 plasma membrane
IDA
GO_REF:0000052
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0002251 organ or tissue specific immune response
IGI
PMID:20702729
Abnormal immune complex processing and spontaneous glomerulo...
KEEP AS NON CORE
Summary: This mouse disease/model annotation reflects complement-dependent immune-complex pathology and glomerular consequences rather than the primary molecular function of CR1.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0002434 immune complex clearance
IGI
PMID:20702729
Abnormal immune complex processing and spontaneous glomerulo...
ACCEPT
Summary: CR1 on erythrocytes and immune cells mediates immune adherence and clearance of complement-opsonized immune complexes/particles.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0006956 complement activation
IGI
PMID:20702729
Abnormal immune complex processing and spontaneous glomerulo...
KEEP AS NON CORE
Summary: This mouse disease/model annotation reflects complement-dependent immune-complex pathology and glomerular consequences rather than the primary molecular function of CR1.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0032835 glomerulus development
IGI
PMID:20702729
Abnormal immune complex processing and spontaneous glomerulo...
KEEP AS NON CORE
Summary: This mouse disease/model annotation reflects complement-dependent immune-complex pathology and glomerular consequences rather than the primary molecular function of CR1.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0016020 membrane
EXP
PMID:1385479
Expression and localization of proteins of the complement sy...
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:1903660 negative regulation of complement-dependent cytotoxicity
IDA
PMID:31862673
Complement Receptor 1 (CR1/CD35)-expressing retinal pigment ...
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0001970 positive regulation of activation of membrane attack complex
IMP
PMID:11981823
The role of complement receptors type 1 (CR1, CD35) and 2 (C...
ACCEPT
Summary: This complement activation/MAC annotation is supported in specific B-cell complement deposition contexts while remaining part of CR1 complement receptor biology.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0005515 protein binding
IPI
PMID:18684861
Ligation of erythrocyte CR1 induces its clustering in comple...
MARK AS OVER ANNOTATED
Summary: The FAP-1/scaffold interaction may be real in erythrocyte CR1 clustering, but generic protein binding does not define CR1 molecular function.
Reason: This annotation is less informative than specific CR1 complement-binding, complement receptor, immune-complex clearance, or complement-regulatory terms.
GO:0005856 cytoskeleton
IDA
PMID:18684861
Ligation of erythrocyte CR1 induces its clustering in comple...
KEEP AS NON CORE
Summary: This erythrocyte clustering, membrane organization, raft, ATP-release, or cytoskeletal annotation is relevant to immune-adherence transfer but is secondary to CR1 complement ligand binding.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0006957 complement activation, alternative pathway
IMP
PMID:11981823
The role of complement receptors type 1 (CR1, CD35) and 2 (C...
ACCEPT
Summary: This complement activation/MAC annotation is supported in specific B-cell complement deposition contexts while remaining part of CR1 complement receptor biology.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0008284 positive regulation of cell population proliferation
IDA
PMID:25742728
Complement receptor type 1 (CR1/CD35) expressed on activated...
KEEP AS NON CORE
Summary: This adaptive immune-cell annotation is supported or plausible but represents T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b receptor/cofactor role.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0009986 cell surface
IDA
PMID:25742728
Complement receptor type 1 (CR1/CD35) expressed on activated...
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0045591 positive regulation of regulatory T cell differentiation
IDA
PMID:25742728
Complement receptor type 1 (CR1/CD35) expressed on activated...
KEEP AS NON CORE
Summary: This adaptive immune-cell annotation is supported or plausible but represents T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b receptor/cofactor role.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0002435 immune complex clearance by erythrocytes
IDA
PMID:24022490
CR1-mediated ATP release by human red blood cells promotes C...
ACCEPT
Summary: CR1 on erythrocytes and immune cells mediates immune adherence and clearance of complement-opsonized immune complexes/particles.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0002430 complement receptor mediated signaling pathway
IDA
PMID:16360013
The complement receptor 1, CR1 (CD35), mediates inhibitory s...
ACCEPT
Summary: CR1 engagement can signal through complement receptor-mediated pathways in immune cells.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0002430 complement receptor mediated signaling pathway
IDA
PMID:22962438
Complement receptor type 1 (CR1, CD35) is a potent inhibitor...
ACCEPT
Summary: CR1 engagement can signal through complement receptor-mediated pathways in immune cells.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0002638 negative regulation of immunoglobulin production
IDA
PMID:22962438
Complement receptor type 1 (CR1, CD35) is a potent inhibitor...
KEEP AS NON CORE
Summary: This adaptive immune-cell annotation is supported or plausible but represents T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b receptor/cofactor role.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0007009 plasma membrane organization
IDA
PMID:24022490
CR1-mediated ATP release by human red blood cells promotes C...
KEEP AS NON CORE
Summary: This erythrocyte clustering, membrane organization, raft, ATP-release, or cytoskeletal annotation is relevant to immune-adherence transfer but is secondary to CR1 complement ligand binding.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0032689 negative regulation of type II interferon production
IDA
PMID:16360013
The complement receptor 1, CR1 (CD35), mediates inhibitory s...
KEEP AS NON CORE
Summary: This adaptive immune-cell annotation is supported or plausible but represents T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b receptor/cofactor role.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0032703 negative regulation of interleukin-2 production
IDA
PMID:16360013
The complement receptor 1, CR1 (CD35), mediates inhibitory s...
KEEP AS NON CORE
Summary: This adaptive immune-cell annotation is supported or plausible but represents T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b receptor/cofactor role.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0042130 negative regulation of T cell proliferation
IDA
PMID:16360013
The complement receptor 1, CR1 (CD35), mediates inhibitory s...
KEEP AS NON CORE
Summary: This adaptive immune-cell annotation is supported or plausible but represents T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b receptor/cofactor role.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0044853 plasma membrane raft
IDA
PMID:24022490
CR1-mediated ATP release by human red blood cells promotes C...
KEEP AS NON CORE
Summary: This erythrocyte clustering, membrane organization, raft, ATP-release, or cytoskeletal annotation is relevant to immune-adherence transfer but is secondary to CR1 complement ligand binding.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:1900099 negative regulation of plasma cell differentiation
IDA
PMID:22962438
Complement receptor type 1 (CR1, CD35) is a potent inhibitor...
KEEP AS NON CORE
Summary: This adaptive immune-cell annotation is supported or plausible but represents T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b receptor/cofactor role.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:1904669 ATP export
IDA
PMID:24022490
CR1-mediated ATP release by human red blood cells promotes C...
KEEP AS NON CORE
Summary: This erythrocyte clustering, membrane organization, raft, ATP-release, or cytoskeletal annotation is relevant to immune-adherence transfer but is secondary to CR1 complement ligand binding.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0001971 negative regulation of activation of membrane attack complex
IDA
PMID:31862673
Complement Receptor 1 (CR1/CD35)-expressing retinal pigment ...
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0045916 negative regulation of complement activation
IDA
PMID:31862673
Complement Receptor 1 (CR1/CD35)-expressing retinal pigment ...
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0030667 secretory granule membrane
TAS
Reactome:R-HSA-6798743
KEEP AS NON CORE
Summary: This intracellular/granule localization is compatible with immune-cell trafficking context but is not the main functional site of CR1 complement receptor activity.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0101003 ficolin-1-rich granule membrane
TAS
Reactome:R-HSA-6800426
KEEP AS NON CORE
Summary: This intracellular/granule localization is compatible with immune-cell trafficking context but is not the main functional site of CR1 complement receptor activity.
Reason: Retain as non-core because the annotation is supported or plausible but reflects cell-type-specific adaptive immune modulation, disease/model context, erythrocyte clustering mechanics, or intracellular/granule localization rather than the primary C3b/C4b receptor and complement-regulatory function.
GO:0070062 extracellular exosome
HDA
PMID:23533145
In-depth proteomic analyses of exosomes isolated from expres...
MARK AS OVER ANNOTATED
Summary: High-throughput extracellular exosome localization is not central to CR1 complement receptor biology and is less informative than membrane/cell-surface annotations.
Reason: This annotation is less informative than specific CR1 complement-binding, complement receptor, immune-complex clearance, or complement-regulatory terms.
GO:0070062 extracellular exosome
HDA
PMID:19056867
Large-scale proteomics and phosphoproteomics of urinary exos...
MARK AS OVER ANNOTATED
Summary: High-throughput extracellular exosome localization is not central to CR1 complement receptor biology and is less informative than membrane/cell-surface annotations.
Reason: This annotation is less informative than specific CR1 complement-binding, complement receptor, immune-complex clearance, or complement-regulatory terms.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-3266557
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-6798743
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-6800426
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-8939088
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-977375
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-977602
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-977615
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-977629
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0005886 plasma membrane
IDA
PMID:6910481
Complement receptor is an inhibitor of the complement cascad...
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0045957 negative regulation of complement activation, alternative pathway
IDA
PMID:10531307
Decay accelerating activity of complement receptor type 1 (C...
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0045959 negative regulation of complement activation, classical pathway
IDA
PMID:10531307
Decay accelerating activity of complement receptor type 1 (C...
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0045959 negative regulation of complement activation, classical pathway
IDA
PMID:6910481
Complement receptor is an inhibitor of the complement cascad...
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:1900004 negative regulation of serine-type endopeptidase activity
IDA
PMID:6910481
Complement receptor is an inhibitor of the complement cascad...
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0009986 cell surface
IDA
PMID:6978375
Complement receptor (CR1) deficiency in erythrocytes from pa...
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:0001851 complement component C3b binding
IDA
PMID:2972794
Identification of distinct C3b and C4b recognition sites in ...
ACCEPT
Summary: CR1 directly binds complement ligands, especially C3b and C4b, and functions as their cell-surface receptor.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0001855 complement component C4b binding
IDA
PMID:2972794
Identification of distinct C3b and C4b recognition sites in ...
ACCEPT
Summary: CR1 directly binds complement ligands, especially C3b and C4b, and functions as their cell-surface receptor.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0001861 complement component C4b receptor activity
IDA
PMID:2972794
Identification of distinct C3b and C4b recognition sites in ...
ACCEPT
Summary: CR1 directly binds complement ligands, especially C3b and C4b, and functions as their cell-surface receptor.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0004877 complement component C3b receptor activity
IDA
PMID:2972794
Identification of distinct C3b and C4b recognition sites in ...
ACCEPT
Summary: CR1 directly binds complement ligands, especially C3b and C4b, and functions as their cell-surface receptor.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.
GO:0009986 cell surface
IDA
PMID:2972794
Identification of distinct C3b and C4b recognition sites in ...
ACCEPT
Summary: This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing single-pass complement receptor.
Reason: This location is consistent with CR1 acting as a single-pass cell-surface/plasma-membrane complement receptor.
GO:1900005 positive regulation of serine-type endopeptidase activity
IDA
PMID:2972794
Identification of distinct C3b and C4b recognition sites in ...
ACCEPT
Summary: CR1 regulates complement by accelerating convertase decay and serving as a cofactor for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.
Reason: This term captures the core CR1/CD35 role as a cell-surface complement receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, and regulates complement convertase and factor I-dependent cleavage reactions.

Core Functions

CR1 acts as a cell-surface receptor for complement-opsonized ligands, binding C3b and C4b through extracellular CCP/Sushi domains to support immune adherence and clearance of immune complexes and complement-tagged particles, especially on human erythrocytes and myeloid/immune cells.

Supporting Evidence:
  • PMID:2972794
    mediated rosette formation with sheep erythrocytes bearing C4b and C3b
  • PMID:24022490
    This process is called immune-adherence clearance
  • PMID:18684861
    The primary identified function of complement receptor 1 (CR1/CD35) on primate erythrocytes is to bind complement-tagged inflammatory particles

CR1 binding to C3b/C4b also provides complement-regulatory activity: CR1 accelerates decay of classical and alternative pathway C3/C5 convertases and acts as a factor I cofactor for cleavage of C3b and C4b, limiting complement activation and complement-dependent cytotoxicity on cells bearing complement-tagged ligands.

Supporting Evidence:
  • PMID:6910481
    It promotes the dissociation of the alternative pathway C3 convertase C3b,Bb and the cleavage of C3b by C3b/C4b inactivator
  • PMID:6910481
    CR1 also inactivates the C3 and C5 convertases of the classical pathway
  • PMID:10531307
    Site 1 (CCPs 1-3) alone mediated the decay acceleration of the classical and alternative pathway C3 convertases

References

Annotation inferences using phylogenetic trees
Gene Ontology annotation based on UniProtKB/Swiss-Prot Subcellular Location vocabulary mapping, accompanied by conservative changes to GO terms applied by UniProt
Gene Ontology annotation based on curation of immunofluorescence data
Electronic Gene Ontology annotations created by ARBA machine learning models
Decay accelerating activity of complement receptor type 1 (CD35). Two active sites are required for dissociating C5 convertases.
The role of complement receptors type 1 (CR1, CD35) and 2 (CR2, CD21) in promoting C3 fragment deposition and membrane attack complex formation on normal peripheral human B cells.
Expression and localization of proteins of the complement system in human skin.
The complement receptor 1, CR1 (CD35), mediates inhibitory signals in human T-lymphocytes.
Ligation of erythrocyte CR1 induces its clustering in complex with scaffolding protein FAP-1.
Large-scale proteomics and phosphoproteomics of urinary exosomes.
Abnormal immune complex processing and spontaneous glomerulonephritis in complement factor H-deficient mice with human complement receptor 1 on erythrocytes.
Complement receptor type 1 (CR1, CD35) is a potent inhibitor of B-cell functions in rheumatoid arthritis patients.
Deciphering complement receptor type 1 interactions with recognition proteins of the lectin complement pathway.
In-depth proteomic analyses of exosomes isolated from expressed prostatic secretions in urine.
CR1-mediated ATP release by human red blood cells promotes CR1 clustering and modulates the immune transfer process.
Complement receptor type 1 (CR1/CD35) expressed on activated human CD4+ T cells contributes to generation of regulatory T cells.
C1q and Mannose-Binding Lectin Interact with CR1 in the Same Region on CCP24-25 Modules.
Identification of distinct C3b and C4b recognition sites in the human C3b/C4b receptor (CR1, CD35) by deletion mutagenesis.
Complement Receptor 1 (CR1/CD35)-expressing retinal pigment epithelial cells as a potential therapy for age-related macular degeneration.
Complement receptor is an inhibitor of the complement cascade.
Complement receptor (CR1) deficiency in erythrocytes from patients with systemic lupus erythematosus.
Reactome:R-HSA-3266557
Factor I cleaves iC3b
Reactome:R-HSA-6798743
Exocytosis of secretory granule membrane proteins
Reactome:R-HSA-6800426
Exocytosis of ficolin-rich granule membrane proteins
Reactome:R-HSA-8939088
CR1 gene expression is stimulated by RUNX1
Reactome:R-HSA-977375
CR1 binds C3bBb/C4bC2a
Reactome:R-HSA-977602
Complement factor I binds to MCP, CR1:C4b, C3b
Reactome:R-HSA-977615
Factor I inactivates MCP/CR1-bound C4b/C3b
Reactome:R-HSA-977629
Displacement of C2a/Bb by CR1

Suggested Questions for Experts

Q: Should CR1 interactions with C1q, MBL, and ficolins be curated with more specific complement-recognition binding terms rather than generic protein binding?

Suggested experts: Complement pathway experts, GO molecular-function curators

Q: Which CR1 immune-cell regulatory annotations should be considered core across species and cell types versus retained as human T-cell/B-cell context?

Suggested experts: CR1 immunology experts, GO immune process curators

Suggested Experiments

Experiment: Use CR1 variants disrupting C3b/C4b sites, CCP24-25 defense-collagen binding, and cytoplasmic FAP-1 interaction in erythrocytes or engineered immune cells, then measure immune-complex binding, transfer to macrophages, factor I cofactor activity, and convertase decay.

Hypothesis: CR1 immune-adherence clearance and complement-regulatory functions can be separated from defense-collagen binding and erythrocyte clustering modules.

Type: domain-resolved complement receptor function assay

Experiment: Compare CR1 engagement on primary human T-cell and B-cell subsets with C3b/C4b ligands versus antibody crosslinking, quantifying cytokines, proliferation, Treg differentiation, plasma-cell differentiation, and complement receptor signaling markers.

Hypothesis: Adaptive immune-cell effects of CR1 are cell-context-specific signaling outcomes of complement ligand engagement rather than the universal core function of CR1.

Type: primary immune-cell receptor signaling assay

📚 Additional Documentation

Notes

(CR1-notes.md)

CR1 curation notes

2026-06-19

  • Deep-research attempt with just deep-research-falcon human CR1 --fallback perplexity-lite timed out after 180 seconds with no generated research artifact, so this manual review uses cached UniProt, GOA, Reactome, PANTHER family, and publication evidence.
  • CR1/CD35 is a large cell-surface complement receptor whose core functions are C3b/C4b binding, immune-adherence clearance, and complement regulation. UniProt summarizes CR1 as a membrane immune adherence receptor for complement-opsonized pathogens/immune complexes and a complement inhibitor/cofactor.
  • C3b/C4b receptor activity is directly supported by deletion-mutagenesis work: recombinant CR1 mediated rosette formation with C3b/C4b-bearing erythrocytes and had factor I cofactor activity PMID:2972794.
  • CR1 complement-inhibitory activity is core. The classic receptor-inhibitor paper reports that CR1 promotes dissociation of the alternative pathway C3 convertase, supports C3b/C4b inactivator cleavage, and inactivates classical pathway C3/C5 convertases PMID:6910481.
  • C5 convertase decay-accelerating activity is supported by domain mapping; site 1 and a properly spaced site 2 are required for efficient activity PMID:10531307.
  • Erythrocyte immune-adherence clearance is a major in vivo role in primates. CR1 on red blood cells binds complement-tagged particles for transport to liver/spleen macrophages, and the paper explicitly names this process immune-adherence clearance PMID:24022490.
  • Erythrocyte CR1 cytoskeletal/FAP-1 association and ATP release are real but secondary mechanistic details for immune-adherence transfer, so I kept cytoskeleton, membrane organization, raft, and ATP export annotations non-core PMID:18684861.
  • T-cell and B-cell regulatory annotations are real immune-context biology but non-core relative to the general C3b/C4b receptor/cofactor function. T-cell CR1 triggering inhibits proliferation and cytokine protein synthesis PMID:16360013, and CD35/CR1 engagement can support Treg phenotypes PMID:25742728.
  • CR1 also acts as a B-cell inhibitory receptor in rheumatoid arthritis contexts, reducing BCR-induced proliferation, plasmablast differentiation, and immunoglobulin production PMID:22962438.
  • CR1 interactions with C1q, MBL, and ficolins are biologically meaningful complement-recognition interactions, but generic protein binding is too broad. I marked the corresponding generic annotations as MODIFY to complement binding PMID:23460739.
  • CR1 expression in engineered retinal pigment epithelial cells can inhibit C3b/C6 deposition and complement-mediated injury, supporting complement-inhibitory annotations while remaining a therapeutic/model context PMID:31862673.
  • High-throughput extracellular exosome annotations and generic FAP-1 protein binding were marked over-annotated because they do not define CR1 core molecular function compared with complement receptor, complement binding, and complement-regulatory terms.

2026-06-20 second-pass audit

The second-pass audit confirmed the existing CR1 review and manual reference metadata. No annotation action changes were needed: CR1 remains curated as a C3b/C4b complement receptor and immune-adherence/complement-regulatory cofactor, with T-cell, B-cell, retinal-therapeutic, cytoskeletal, and generic high-throughput interaction contexts retained as non-core, modified, or over-annotated as already reviewed.

📄 View Raw YAML

id: P17927
gene_symbol: CR1
product_type: PROTEIN
status: COMPLETE
taxon:
  id: NCBITaxon:9606
  label: Homo sapiens
description: 'CR1 encodes complement receptor type 1/CD35, a large single-pass membrane receptor
  composed of complement-control protein repeat domains and expressed on erythrocytes, monocytes/macrophages,
  granulocytes, B cells, subsets of T cells, follicular dendritic cells, and other immune-associated
  cells. CR1 binds complement-opsonized ligands, especially C3b and C4b, allowing immune adherence
  and clearance of immune complexes and particles from the circulation. CR1 also regulates
  complement by accelerating decay of classical and alternative pathway convertases and acting
  as a factor I cofactor for cleavage of C3b and C4b. In specific immune-cell contexts, CR1
  engagement can modulate T-cell, B-cell, and regulatory T-cell responses, while complement-pathway
  interactions with C1q, MBL, ficolins, and microbial ligands broaden its immune-recognition
  roles.'
existing_annotations:
- term:
    id: GO:0005576
    label: extracellular region
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: is_active_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: is_active_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0045959
    label: negative regulation of complement activation, classical pathway
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0002456
    label: T cell mediated immunity
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: involved_in
  review:
    summary: 'This adaptive immune-cell annotation is supported or plausible but represents
      T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b
      receptor/cofactor role.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0001848
    label: complement binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: enables
  review:
    summary: 'CR1 directly binds complement ligands, especially C3b and C4b, and functions
      as their cell-surface receptor.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0004875
    label: complement receptor activity
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: enables
  review:
    summary: 'CR1 directly binds complement ligands, especially C3b and C4b, and functions
      as their cell-surface receptor.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0006957
    label: complement activation, alternative pathway
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: involved_in
  review:
    summary: 'This complement activation/MAC annotation is supported in specific B-cell complement
      deposition contexts while remaining part of CR1 complement receptor biology.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0016020
    label: membrane
  evidence_type: IEA
  original_reference_id: GO_REF:0000044
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0030449
    label: regulation of complement activation
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:1903659
    label: regulation of complement-dependent cytotoxicity
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:23460739
  qualifier: enables
  review:
    summary: 'The MBL/ficolin/C1q interaction evidence supports complement-recognition ligand
      binding, but generic protein binding is too broad.'
    action: MODIFY
    reason: Replace generic protein binding with complement binding for CR1 interactions
      with recognition proteins of the lectin/classical complement pathways.
    proposed_replacement_terms:
    - id: GO:0001848
      label: complement binding
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:29563915
  qualifier: enables
  review:
    summary: 'The MBL/ficolin/C1q interaction evidence supports complement-recognition ligand
      binding, but generic protein binding is too broad.'
    action: MODIFY
    reason: Replace generic protein binding with complement binding for CR1 interactions
      with recognition proteins of the lectin/classical complement pathways.
    proposed_replacement_terms:
    - id: GO:0001848
      label: complement binding
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IDA
  original_reference_id: GO_REF:0000052
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0002251
    label: organ or tissue specific immune response
  evidence_type: IGI
  original_reference_id: PMID:20702729
  qualifier: acts_upstream_of_or_within
  review:
    summary: 'This mouse disease/model annotation reflects complement-dependent immune-complex
      pathology and glomerular consequences rather than the primary molecular function of
      CR1.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0002434
    label: immune complex clearance
  evidence_type: IGI
  original_reference_id: PMID:20702729
  qualifier: acts_upstream_of_or_within
  review:
    summary: 'CR1 on erythrocytes and immune cells mediates immune adherence and clearance
      of complement-opsonized immune complexes/particles.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0006956
    label: complement activation
  evidence_type: IGI
  original_reference_id: PMID:20702729
  qualifier: acts_upstream_of_or_within
  review:
    summary: 'This mouse disease/model annotation reflects complement-dependent immune-complex
      pathology and glomerular consequences rather than the primary molecular function of
      CR1.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0032835
    label: glomerulus development
  evidence_type: IGI
  original_reference_id: PMID:20702729
  qualifier: acts_upstream_of_or_within
  review:
    summary: 'This mouse disease/model annotation reflects complement-dependent immune-complex
      pathology and glomerular consequences rather than the primary molecular function of
      CR1.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0016020
    label: membrane
  evidence_type: EXP
  original_reference_id: PMID:1385479
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:1903660
    label: negative regulation of complement-dependent cytotoxicity
  evidence_type: IDA
  original_reference_id: PMID:31862673
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0001970
    label: positive regulation of activation of membrane attack complex
  evidence_type: IMP
  original_reference_id: PMID:11981823
  qualifier: involved_in
  review:
    summary: 'This complement activation/MAC annotation is supported in specific B-cell complement
      deposition contexts while remaining part of CR1 complement receptor biology.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:18684861
  qualifier: enables
  review:
    summary: 'The FAP-1/scaffold interaction may be real in erythrocyte CR1 clustering, but
      generic protein binding does not define CR1 molecular function.'
    action: MARK_AS_OVER_ANNOTATED
    reason: This annotation is less informative than specific CR1 complement-binding, 
      complement receptor, immune-complex clearance, or complement-regulatory terms.
- term:
    id: GO:0005856
    label: cytoskeleton
  evidence_type: IDA
  original_reference_id: PMID:18684861
  qualifier: located_in
  review:
    summary: 'This erythrocyte clustering, membrane organization, raft, ATP-release, or cytoskeletal
      annotation is relevant to immune-adherence transfer but is secondary to CR1 complement
      ligand binding.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0006957
    label: complement activation, alternative pathway
  evidence_type: IMP
  original_reference_id: PMID:11981823
  qualifier: involved_in
  review:
    summary: 'This complement activation/MAC annotation is supported in specific B-cell complement
      deposition contexts while remaining part of CR1 complement receptor biology.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0008284
    label: positive regulation of cell population proliferation
  evidence_type: IDA
  original_reference_id: PMID:25742728
  qualifier: involved_in
  review:
    summary: 'This adaptive immune-cell annotation is supported or plausible but represents
      T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b
      receptor/cofactor role.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0009986
    label: cell surface
  evidence_type: IDA
  original_reference_id: PMID:25742728
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0045591
    label: positive regulation of regulatory T cell differentiation
  evidence_type: IDA
  original_reference_id: PMID:25742728
  qualifier: involved_in
  review:
    summary: 'This adaptive immune-cell annotation is supported or plausible but represents
      T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b
      receptor/cofactor role.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0002435
    label: immune complex clearance by erythrocytes
  evidence_type: IDA
  original_reference_id: PMID:24022490
  qualifier: involved_in
  review:
    summary: 'CR1 on erythrocytes and immune cells mediates immune adherence and clearance
      of complement-opsonized immune complexes/particles.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0002430
    label: complement receptor mediated signaling pathway
  evidence_type: IDA
  original_reference_id: PMID:16360013
  qualifier: involved_in
  review:
    summary: 'CR1 engagement can signal through complement receptor-mediated pathways in immune
      cells.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0002430
    label: complement receptor mediated signaling pathway
  evidence_type: IDA
  original_reference_id: PMID:22962438
  qualifier: involved_in
  review:
    summary: 'CR1 engagement can signal through complement receptor-mediated pathways in immune
      cells.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0002638
    label: negative regulation of immunoglobulin production
  evidence_type: IDA
  original_reference_id: PMID:22962438
  qualifier: acts_upstream_of
  review:
    summary: 'This adaptive immune-cell annotation is supported or plausible but represents
      T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b
      receptor/cofactor role.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0007009
    label: plasma membrane organization
  evidence_type: IDA
  original_reference_id: PMID:24022490
  qualifier: involved_in
  review:
    summary: 'This erythrocyte clustering, membrane organization, raft, ATP-release, or cytoskeletal
      annotation is relevant to immune-adherence transfer but is secondary to CR1 complement
      ligand binding.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0032689
    label: negative regulation of type II interferon production
  evidence_type: IDA
  original_reference_id: PMID:16360013
  qualifier: acts_upstream_of_or_within
  review:
    summary: 'This adaptive immune-cell annotation is supported or plausible but represents
      T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b
      receptor/cofactor role.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0032703
    label: negative regulation of interleukin-2 production
  evidence_type: IDA
  original_reference_id: PMID:16360013
  qualifier: acts_upstream_of_or_within
  review:
    summary: 'This adaptive immune-cell annotation is supported or plausible but represents
      T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b
      receptor/cofactor role.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0042130
    label: negative regulation of T cell proliferation
  evidence_type: IDA
  original_reference_id: PMID:16360013
  qualifier: involved_in
  review:
    summary: 'This adaptive immune-cell annotation is supported or plausible but represents
      T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b
      receptor/cofactor role.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0044853
    label: plasma membrane raft
  evidence_type: IDA
  original_reference_id: PMID:24022490
  qualifier: located_in
  review:
    summary: 'This erythrocyte clustering, membrane organization, raft, ATP-release, or cytoskeletal
      annotation is relevant to immune-adherence transfer but is secondary to CR1 complement
      ligand binding.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:1900099
    label: negative regulation of plasma cell differentiation
  evidence_type: IDA
  original_reference_id: PMID:22962438
  qualifier: acts_upstream_of
  review:
    summary: 'This adaptive immune-cell annotation is supported or plausible but represents
      T-cell/B-cell regulatory consequences of CR1 engagement rather than the primary C3b/C4b
      receptor/cofactor role.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:1904669
    label: ATP export
  evidence_type: IDA
  original_reference_id: PMID:24022490
  qualifier: acts_upstream_of_or_within
  review:
    summary: 'This erythrocyte clustering, membrane organization, raft, ATP-release, or cytoskeletal
      annotation is relevant to immune-adherence transfer but is secondary to CR1 complement
      ligand binding.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0001971
    label: negative regulation of activation of membrane attack complex
  evidence_type: IDA
  original_reference_id: PMID:31862673
  qualifier: acts_upstream_of
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0045916
    label: negative regulation of complement activation
  evidence_type: IDA
  original_reference_id: PMID:31862673
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0030667
    label: secretory granule membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-6798743
  qualifier: located_in
  review:
    summary: 'This intracellular/granule localization is compatible with immune-cell trafficking
      context but is not the main functional site of CR1 complement receptor activity.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0101003
    label: ficolin-1-rich granule membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-6800426
  qualifier: located_in
  review:
    summary: 'This intracellular/granule localization is compatible with immune-cell trafficking
      context but is not the main functional site of CR1 complement receptor activity.'
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the annotation is supported or plausible but 
      reflects cell-type-specific adaptive immune modulation, disease/model context, 
      erythrocyte clustering mechanics, or intracellular/granule localization rather 
      than the primary C3b/C4b receptor and complement-regulatory function.
- term:
    id: GO:0070062
    label: extracellular exosome
  evidence_type: HDA
  original_reference_id: PMID:23533145
  qualifier: located_in
  review:
    summary: 'High-throughput extracellular exosome localization is not central to CR1 complement
      receptor biology and is less informative than membrane/cell-surface annotations.'
    action: MARK_AS_OVER_ANNOTATED
    reason: This annotation is less informative than specific CR1 complement-binding, 
      complement receptor, immune-complex clearance, or complement-regulatory terms.
- term:
    id: GO:0070062
    label: extracellular exosome
  evidence_type: HDA
  original_reference_id: PMID:19056867
  qualifier: located_in
  review:
    summary: 'High-throughput extracellular exosome localization is not central to CR1 complement
      receptor biology and is less informative than membrane/cell-surface annotations.'
    action: MARK_AS_OVER_ANNOTATED
    reason: This annotation is less informative than specific CR1 complement-binding, 
      complement receptor, immune-complex clearance, or complement-regulatory terms.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-3266557
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-6798743
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-6800426
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-8939088
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-977375
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-977602
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-977615
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-977629
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IDA
  original_reference_id: PMID:6910481
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0045957
    label: negative regulation of complement activation, alternative pathway
  evidence_type: IDA
  original_reference_id: PMID:10531307
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0045959
    label: negative regulation of complement activation, classical pathway
  evidence_type: IDA
  original_reference_id: PMID:10531307
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0045959
    label: negative regulation of complement activation, classical pathway
  evidence_type: IDA
  original_reference_id: PMID:6910481
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:1900004
    label: negative regulation of serine-type endopeptidase activity
  evidence_type: IDA
  original_reference_id: PMID:6910481
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0009986
    label: cell surface
  evidence_type: IDA
  original_reference_id: PMID:6978375
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:0001851
    label: complement component C3b binding
  evidence_type: IDA
  original_reference_id: PMID:2972794
  qualifier: enables
  review:
    summary: 'CR1 directly binds complement ligands, especially C3b and C4b, and functions
      as their cell-surface receptor.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0001855
    label: complement component C4b binding
  evidence_type: IDA
  original_reference_id: PMID:2972794
  qualifier: enables
  review:
    summary: 'CR1 directly binds complement ligands, especially C3b and C4b, and functions
      as their cell-surface receptor.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0001861
    label: complement component C4b receptor activity
  evidence_type: IDA
  original_reference_id: PMID:2972794
  qualifier: enables
  review:
    summary: 'CR1 directly binds complement ligands, especially C3b and C4b, and functions
      as their cell-surface receptor.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0004877
    label: complement component C3b receptor activity
  evidence_type: IDA
  original_reference_id: PMID:2972794
  qualifier: enables
  review:
    summary: 'CR1 directly binds complement ligands, especially C3b and C4b, and functions
      as their cell-surface receptor.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
- term:
    id: GO:0009986
    label: cell surface
  evidence_type: IDA
  original_reference_id: PMID:2972794
  qualifier: located_in
  review:
    summary: 'This membrane/extracellular location is consistent with CR1 as a large ectodomain-containing
      single-pass complement receptor.'
    action: ACCEPT
    reason: This location is consistent with CR1 acting as a single-pass 
      cell-surface/plasma-membrane complement receptor.
- term:
    id: GO:1900005
    label: positive regulation of serine-type endopeptidase activity
  evidence_type: IDA
  original_reference_id: PMID:2972794
  qualifier: involved_in
  review:
    summary: 'CR1 regulates complement by accelerating convertase decay and serving as a cofactor
      for factor I-mediated C3b/C4b cleavage, limiting complement activation and cytotoxicity.'
    action: ACCEPT
    reason: This term captures the core CR1/CD35 role as a cell-surface complement 
      receptor that binds C3b/C4b-bearing ligands, supports immune adherence/clearance, 
      and regulates complement convertase and factor I-dependent cleavage reactions.
references:
- id: GO_REF:0000033
  title: Annotation inferences using phylogenetic trees
  findings: []
- id: GO_REF:0000044
  title: Gene Ontology annotation based on UniProtKB/Swiss-Prot Subcellular Location 
    vocabulary mapping, accompanied by conservative changes to GO terms applied by 
    UniProt
  findings: []
- id: GO_REF:0000052
  title: Gene Ontology annotation based on curation of immunofluorescence data
  findings: []
- id: GO_REF:0000117
  title: Electronic Gene Ontology annotations created by ARBA machine learning models
  findings: []
- id: PMID:10531307
  title: Decay accelerating activity of complement receptor type 1 (CD35). Two active 
    sites are required for dissociating C5 convertases.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Abstract cached; supports decay-accelerating activity against C3/C5 
      convertases.
- id: PMID:11981823
  title: The role of complement receptors type 1 (CR1, CD35) and 2 (CR2, CD21) in 
    promoting C3 fragment deposition and membrane attack complex formation on normal 
    peripheral human B cells.
  findings: []
- id: PMID:1385479
  title: Expression and localization of proteins of the complement system in human skin.
  findings: []
- id: PMID:16360013
  title: The complement receptor 1, CR1 (CD35), mediates inhibitory signals in human 
    T-lymphocytes.
  findings: []
  reference_review:
    relevance: MEDIUM
    correctness: VERIFIED
    review_notes: Abstract cached; supports inhibitory CR1 signaling in human T 
      lymphocytes.
- id: PMID:18684861
  title: Ligation of erythrocyte CR1 induces its clustering in complex with scaffolding 
    protein FAP-1.
  findings: []
  reference_review:
    relevance: MEDIUM
    correctness: VERIFIED
    review_notes: Abstract cached; supports erythrocyte CR1 cytoskeletal association and
      FAP-1 scaffold interaction as non-core clustering context.
- id: PMID:19056867
  title: Large-scale proteomics and phosphoproteomics of urinary exosomes.
  findings: []
- id: PMID:20702729
  title: Abnormal immune complex processing and spontaneous glomerulonephritis in 
    complement factor H-deficient mice with human complement receptor 1 on erythrocytes.
  findings: []
  reference_review:
    relevance: MEDIUM
    correctness: VERIFIED
    review_notes: Abstract cached; supports human erythrocyte CR1 immune-complex 
      processing/glomerular disease model context.
- id: PMID:22962438
  title: Complement receptor type 1 (CR1, CD35) is a potent inhibitor of B-cell 
    functions in rheumatoid arthritis patients.
  findings: []
  reference_review:
    relevance: MEDIUM
    correctness: VERIFIED
    review_notes: Abstract cached; supports inhibitory B-cell effects of CR1 engagement.
- id: PMID:23460739
  title: Deciphering complement receptor type 1 interactions with recognition proteins 
    of the lectin complement pathway.
  findings: []
  reference_review:
    relevance: MEDIUM
    correctness: VERIFIED
    review_notes: Abstract cached; supports CR1 binding to MBL and ficolins through 
      CCP22-30/CCP24-25.
- id: PMID:23533145
  title: In-depth proteomic analyses of exosomes isolated from expressed prostatic 
    secretions in urine.
  findings: []
- id: PMID:24022490
  title: CR1-mediated ATP release by human red blood cells promotes CR1 clustering and 
    modulates the immune transfer process.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Abstract cached; supports erythrocyte immune-adherence clearance, ATP 
      release, membrane mobility, and CR1 clustering.
- id: PMID:25742728
  title: Complement receptor type 1 (CR1/CD35) expressed on activated human CD4+ T cells
    contributes to generation of regulatory T cells.
  findings: []
  reference_review:
    relevance: MEDIUM
    correctness: VERIFIED
    review_notes: Abstract cached; supports activated CD4 T-cell CR1 roles in regulatory
      T-cell generation.
- id: PMID:29563915
  title: C1q and Mannose-Binding Lectin Interact with CR1 in the Same Region on CCP24-25
    Modules.
  findings: []
  reference_review:
    relevance: MEDIUM
    correctness: VERIFIED
    review_notes: Full text cached; supports C1q and MBL interaction with CR1 CCP24-25 
      modules.
- id: PMID:2972794
  title: Identification of distinct C3b and C4b recognition sites in the human C3b/C4b 
    receptor (CR1, CD35) by deletion mutagenesis.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Abstract cached; supports C3b/C4b receptor activity, multivalent 
      binding sites, and factor I cofactor activity.
- id: PMID:31862673
  title: Complement Receptor 1 (CR1/CD35)-expressing retinal pigment epithelial cells as
    a potential therapy for age-related macular degeneration.
  findings: []
  reference_review:
    relevance: MEDIUM
    correctness: VERIFIED
    review_notes: Abstract cached; supports CR1 complement-inhibitory activity in 
      engineered RPE context.
- id: PMID:6910481
  title: Complement receptor is an inhibitor of the complement cascade.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Abstract/full abstract cached; supports CR1 inhibition of alternative 
      and classical convertases and factor I cofactor activity.
- id: PMID:6978375
  title: Complement receptor (CR1) deficiency in erythrocytes from patients with 
    systemic lupus erythematosus.
  findings: []
- id: Reactome:R-HSA-3266557
  title: Factor I cleaves iC3b
  findings: []
- id: Reactome:R-HSA-6798743
  title: Exocytosis of secretory granule membrane proteins
  findings: []
- id: Reactome:R-HSA-6800426
  title: Exocytosis of ficolin-rich granule membrane proteins
  findings: []
- id: Reactome:R-HSA-8939088
  title: CR1 gene expression is stimulated by RUNX1
  findings: []
- id: Reactome:R-HSA-977375
  title: CR1 binds C3bBb/C4bC2a
  findings: []
- id: Reactome:R-HSA-977602
  title: Complement factor I binds to MCP, CR1:C4b, C3b
  findings: []
- id: Reactome:R-HSA-977615
  title: Factor I inactivates MCP/CR1-bound C4b/C3b
  findings: []
- id: Reactome:R-HSA-977629
  title: Displacement of C2a/Bb by CR1
  findings: []
core_functions:
- molecular_function:
    id: GO:0004875
    label: complement receptor activity
  description: CR1 acts as a cell-surface receptor for complement-opsonized ligands, 
    binding C3b and C4b through extracellular CCP/Sushi domains to support immune 
    adherence and clearance of immune complexes and complement-tagged particles, 
    especially on human erythrocytes and myeloid/immune cells.
  directly_involved_in:
  - id: GO:0002434
    label: immune complex clearance
  - id: GO:0002435
    label: immune complex clearance by erythrocytes
  - id: GO:0002430
    label: complement receptor mediated signaling pathway
  locations:
  - id: GO:0005886
    label: plasma membrane
  - id: GO:0009986
    label: cell surface
  supported_by:
  - reference_id: PMID:2972794
    supporting_text: mediated rosette formation with sheep erythrocytes bearing C4b and 
      C3b
  - reference_id: PMID:24022490
    supporting_text: This process is called immune-adherence clearance
  - reference_id: PMID:18684861
    supporting_text: The primary identified function of complement receptor 1 (CR1/CD35)
      on primate erythrocytes is to bind complement-tagged inflammatory particles
- molecular_function:
    id: GO:0001851
    label: complement component C3b binding
  description: 'CR1 binding to C3b/C4b also provides complement-regulatory activity: CR1 accelerates
    decay of classical and alternative pathway C3/C5 convertases and acts as a factor I cofactor
    for cleavage of C3b and C4b, limiting complement activation and complement-dependent cytotoxicity
    on cells bearing complement-tagged ligands.'
  directly_involved_in:
  - id: GO:0045957
    label: negative regulation of complement activation, alternative pathway
  - id: GO:0045959
    label: negative regulation of complement activation, classical pathway
  - id: GO:0045916
    label: negative regulation of complement activation
  - id: GO:0001971
    label: negative regulation of activation of membrane attack complex
  - id: GO:1900005
    label: positive regulation of serine-type endopeptidase activity
  locations:
  - id: GO:0005886
    label: plasma membrane
  - id: GO:0009986
    label: cell surface
  supported_by:
  - reference_id: PMID:6910481
    supporting_text: It promotes the dissociation of the alternative pathway C3 
      convertase C3b,Bb and the cleavage of C3b by C3b/C4b inactivator
  - reference_id: PMID:6910481
    supporting_text: CR1 also inactivates the C3 and C5 convertases of the classical 
      pathway
  - reference_id: PMID:10531307
    supporting_text: Site 1 (CCPs 1-3) alone mediated the decay acceleration of the 
      classical and alternative pathway C3 convertases
proposed_new_terms: []
suggested_questions:
- question: Should CR1 interactions with C1q, MBL, and ficolins be curated with more 
    specific complement-recognition binding terms rather than generic protein binding?
  experts:
  - Complement pathway experts
  - GO molecular-function curators
- question: Which CR1 immune-cell regulatory annotations should be considered core 
    across species and cell types versus retained as human T-cell/B-cell context?
  experts:
  - CR1 immunology experts
  - GO immune process curators
suggested_experiments:
- description: Use CR1 variants disrupting C3b/C4b sites, CCP24-25 defense-collagen 
    binding, and cytoplasmic FAP-1 interaction in erythrocytes or engineered immune 
    cells, then measure immune-complex binding, transfer to macrophages, factor I 
    cofactor activity, and convertase decay.
  hypothesis: CR1 immune-adherence clearance and complement-regulatory functions can be 
    separated from defense-collagen binding and erythrocyte clustering modules.
  experiment_type: domain-resolved complement receptor function assay
- description: Compare CR1 engagement on primary human T-cell and B-cell subsets with 
    C3b/C4b ligands versus antibody crosslinking, quantifying cytokines, proliferation, 
    Treg differentiation, plasma-cell differentiation, and complement receptor signaling
    markers.
  hypothesis: Adaptive immune-cell effects of CR1 are cell-context-specific signaling 
    outcomes of complement ligand engagement rather than the universal core function of 
    CR1.
  experiment_type: primary immune-cell receptor signaling assay