DNMT3A

UniProt ID: Q9Y6K1
Organism: Homo sapiens
Review Status: IN PROGRESS
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Gene Description

DNMT3A is a de novo DNA cytosine-5 methyltransferase that establishes and remodels mammalian DNA methylation patterns. Its catalytic domain recognizes cytosines, preferentially in CpG contexts, while PWWP and ADD chromatin-reader domains and interacting proteins guide genomic targeting and activity. Chromatin methylation contributes to transcriptional regulation and cell identity. Alternative N-terminal isoforms differ in chromatin localization.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0000122 negative regulation of transcription by RNA polymerase II
IDA
PMID:23042785
Control of epigenetic states by WT1 via regulation of de nov...
ACCEPT
Summary: DNMT3A can repress transcription through targeted DNA methylation.
Reason: Recruitment by transcriptional regulators and promoter hypermethylation provide mechanistic support for negative transcriptional regulation. This does not imply that all DNMT3A-dependent methylation universally represses transcription.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:23042785
DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
GO:0000122 negative regulation of transcription by RNA polymerase II
IMP
PMID:19786833
Dnmt3/transcription factor interactions as crucial players i...
ACCEPT
Summary: DNMT3A can repress transcription through targeted DNA methylation.
Reason: Recruitment by transcriptional regulators and promoter hypermethylation provide mechanistic support for negative transcriptional regulation. This does not imply that all DNMT3A-dependent methylation universally represses transcription.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:23042785
DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
GO:0000775 chromosome, centromeric region
IEA
GO_REF:0000107
ACCEPT
Summary: DNMT3A associates with chromatin, including heterochromatic pools.
Reason: Isoform localization and chromatin-reader studies establish chromatin association. N-terminal isoforms differ in their distribution, so these gene-level location records should not be read as every isoform occupying identical chromatin.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
PMID:30478443
Substitutions in the PWWP domain abrogate binding to the histone modifications H3K36me2 and H3K36me3, and alter DNA methylation in patient cells.
GO:0000792 heterochromatin
IEA
GO_REF:0000107
ACCEPT
Summary: DNMT3A associates with chromatin, including heterochromatic pools.
Reason: Isoform localization and chromatin-reader studies establish chromatin association. N-terminal isoforms differ in their distribution, so these gene-level location records should not be read as every isoform occupying identical chromatin.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
PMID:30478443
Substitutions in the PWWP domain abrogate binding to the histone modifications H3K36me2 and H3K36me3, and alter DNA methylation in patient cells.
GO:0000978 RNA polymerase II cis-regulatory region sequence-specific DNA binding
IDA
PMID:19786833
Dnmt3/transcription factor interactions as crucial players i...
UNDECIDED
Summary: Promoter occupancy needs distinction from intrinsic transcription-factor-like sequence recognition.
Reason: The primary studies establish DNMT3A recruitment to regulatory DNA, including recruitment by MYC or changes downstream of WT1. The available evidence does not clearly resolve whether this specific cis-regulatory sequence-specific binding term describes intrinsic recognition rather than partner-mediated ChIP occupancy. Retain uncertainty rather than removing an experimental annotation from an incomplete assay assessment.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:29414941
Arg836 of the target recognition domain makes crucial contacts with CpG, ensuring DNMT3A enzymatic preference towards CpG sites in cells.
GO:0000978 RNA polymerase II cis-regulatory region sequence-specific DNA binding
IDA
PMID:23042785
Control of epigenetic states by WT1 via regulation of de nov...
UNDECIDED
Summary: Promoter occupancy needs distinction from intrinsic transcription-factor-like sequence recognition.
Reason: The primary studies establish DNMT3A recruitment to regulatory DNA, including recruitment by MYC or changes downstream of WT1. The available evidence does not clearly resolve whether this specific cis-regulatory sequence-specific binding term describes intrinsic recognition rather than partner-mediated ChIP occupancy. Retain uncertainty rather than removing an experimental annotation from an incomplete assay assessment.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:29414941
Arg836 of the target recognition domain makes crucial contacts with CpG, ensuring DNMT3A enzymatic preference towards CpG sites in cells.
GO:0001741 XY body
IEA
GO_REF:0000107
UNDECIDED
Summary: The XY-body localization transfer requires germ-cell and isoform-specific evidence.
Reason: Broad nuclear/chromatin localization does not establish the specialized meiotic sex-chromosome compartment. The source experiment underlying this ortholog transfer needs tracing.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0003677 DNA binding
EXP
PMID:29414941
Structural basis for DNMT3A-mediated de novo DNA methylation...
ACCEPT
Summary: DNMT3A binds DNA and catalyzes cytosine methylation.
Reason: DNA-bound structures and substrate assays establish catalytic recognition of unmethylated CpG DNA and the de novo methyltransferase reaction.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:29414941
Arg836 of the target recognition domain makes crucial contacts with CpG, ensuring DNMT3A enzymatic preference towards CpG sites in cells.
GO:0003677 DNA binding
IEA
GO_REF:0000107
ACCEPT
Summary: DNMT3A binds DNA and catalyzes cytosine methylation.
Reason: DNA-bound structures and substrate assays establish catalytic recognition of unmethylated CpG DNA and the de novo methyltransferase reaction.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:29414941
Arg836 of the target recognition domain makes crucial contacts with CpG, ensuring DNMT3A enzymatic preference towards CpG sites in cells.
GO:0003682 chromatin binding
IEA
GO_REF:0000107
ACCEPT
Summary: DNMT3A associates with chromatin, including heterochromatic pools.
Reason: Isoform localization and chromatin-reader studies establish chromatin association. N-terminal isoforms differ in their distribution, so these gene-level location records should not be read as every isoform occupying identical chromatin.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
PMID:30478443
Substitutions in the PWWP domain abrogate binding to the histone modifications H3K36me2 and H3K36me3, and alter DNA methylation in patient cells.
GO:0003714 transcription corepressor activity
IEA
GO_REF:0000117
ACCEPT
Summary: DNMT3A regulates gene expression as a recruited epigenetic corepressor.
Reason: Interaction with transcriptional regulators targets methylation and can repress promoter activity. Its enzymatic mechanism provides the specificity beyond a generic corepressor label.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:23042785
DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
GO:0003714 transcription corepressor activity
TAS
PMID:19786833
Dnmt3/transcription factor interactions as crucial players i...
ACCEPT
Summary: DNMT3A regulates gene expression as a recruited epigenetic corepressor.
Reason: Interaction with transcriptional regulators targets methylation and can repress promoter activity. Its enzymatic mechanism provides the specificity beyond a generic corepressor label.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:23042785
DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
GO:0003886 DNA (cytosine-5-)-methyltransferase activity
IBA
GO_REF:0000033
ACCEPT
Summary: DNMT3A binds DNA and catalyzes cytosine methylation.
Reason: DNA-bound structures and substrate assays establish catalytic recognition of unmethylated CpG DNA and the de novo methyltransferase reaction.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:29414941
Arg836 of the target recognition domain makes crucial contacts with CpG, ensuring DNMT3A enzymatic preference towards CpG sites in cells.
GO:0003886 DNA (cytosine-5-)-methyltransferase activity
IDA
PMID:12138111
A novel Dnmt3a isoform produced from an alternative promoter...
ACCEPT
Summary: DNMT3A binds DNA and catalyzes cytosine methylation.
Reason: DNA-bound structures and substrate assays establish catalytic recognition of unmethylated CpG DNA and the de novo methyltransferase reaction.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:29414941
Arg836 of the target recognition domain makes crucial contacts with CpG, ensuring DNMT3A enzymatic preference towards CpG sites in cells.
GO:0003886 DNA (cytosine-5-)-methyltransferase activity
IDA
PMID:19786833
Dnmt3/transcription factor interactions as crucial players i...
ACCEPT
Summary: DNMT3A binds DNA and catalyzes cytosine methylation.
Reason: DNA-bound structures and substrate assays establish catalytic recognition of unmethylated CpG DNA and the de novo methyltransferase reaction.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:29414941
Arg836 of the target recognition domain makes crucial contacts with CpG, ensuring DNMT3A enzymatic preference towards CpG sites in cells.
GO:0003886 DNA (cytosine-5-)-methyltransferase activity
IEA
GO_REF:0000120
ACCEPT
Summary: DNMT3A binds DNA and catalyzes cytosine methylation.
Reason: DNA-bound structures and substrate assays establish catalytic recognition of unmethylated CpG DNA and the de novo methyltransferase reaction.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:29414941
Arg836 of the target recognition domain makes crucial contacts with CpG, ensuring DNMT3A enzymatic preference towards CpG sites in cells.
GO:0005515 protein binding
IPI
PMID:16357870
The Polycomb group protein EZH2 directly controls DNA methyl...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:16357870
The Polycomb group protein EZH2 directly controls DNA methyl...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:16682412
The histone methyltransferase SETDB1 and the DNA methyltrans...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:16983096
Recruitment of the de novo DNA methyltransferase Dnmt3a by K...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:17713477
Structure of Dnmt3a bound to Dnmt3L suggests a model for de ...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:19234465
PRMT5-mediated methylation of histone H4R3 recruits DNMT3A, ...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:19798101
Np95 interacts with de novo DNA methyltransferases, Dnmt3a a...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:19834512
Structural basis for recognition of H3K4 methylation status ...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:21976679
cAMP-responsive element modulator (CREM)Ξ± protein signaling ...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:24074865
Dnmt3L antagonizes DNA methylation at bivalent promoters and...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:25383530
Structural insight into autoinhibition and histone H3-induce...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:25416956
A proteome-scale map of the human interactome network.
ACCEPT
Summary: DNMT3A participates in a regulated methyltransferase complex.
Reason: The DNMT3A-DNMT3L and histone-bound structures establish physical complex formation and catalytic regulation. Generic interaction annotations here are mechanistically grounded by the same structural study.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:25383530
Histone H3 (but not H3K4me3) disrupts ADD-CD interaction, induces a large movement of the ADD domain, and thus releases the autoinhibition of DNMT3A.
GO:0005515 protein binding
IPI
PMID:25416956
A proteome-scale map of the human interactome network.
ACCEPT
Summary: DNMT3A participates in a regulated methyltransferase complex.
Reason: The DNMT3A-DNMT3L and histone-bound structures establish physical complex formation and catalytic regulation. Generic interaction annotations here are mechanistically grounded by the same structural study.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:25383530
Histone H3 (but not H3K4me3) disrupts ADD-CD interaction, induces a large movement of the ADD domain, and thus releases the autoinhibition of DNMT3A.
GO:0005515 protein binding
IPI
PMID:31515488
Extensive disruption of protein interactions by genetic vari...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:32051553
The EGFR-ZNF263 signaling axis silences SIX3 in glioblastoma...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:32814053
Interactome Mapping Provides a Network of Neurodegenerative ...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:32968275
Structure of nucleosome-boundΒ DNA methyltransferases DNMT3A ...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:33961781
Dual proteome-scale networks reveal cell-specific remodeling...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005515 protein binding
IPI
PMID:33961781
Dual proteome-scale networks reveal cell-specific remodeling...
UNDECIDED
Summary: The particular interaction needs partner-level source verification.
Reason: DNMT3A has well-characterized regulatory partners, but that does not validate every seeded interaction. The exact gene/partner assay for this row has not been verified in the retrieved publication or supplementary dataset; retain the curated interaction pending that evidence check.
GO:0005634 nucleus
IBA
GO_REF:0000033
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005634 nucleus
IDA
PMID:12138111
A novel Dnmt3a isoform produced from an alternative promoter...
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005634 nucleus
IDA
PMID:25743254
Proteomic analyses reveal that loss of TDP-43 affects RNA pr...
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005634 nucleus
IEA
GO_REF:0000120
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005634 nucleus
NAS
PMID:25383530
Structural insight into autoinhibition and histone H3-induce...
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005654 nucleoplasm
IDA
GO_REF:0000052
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005654 nucleoplasm
TAS
Reactome:R-HSA-212222
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005654 nucleoplasm
TAS
Reactome:R-HSA-212269
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005654 nucleoplasm
TAS
Reactome:R-HSA-5212679
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005654 nucleoplasm
TAS
Reactome:R-HSA-9710490
ACCEPT
Summary: DNMT3A acts in the nucleus and nucleoplasm.
Reason: Direct human tissue imaging and chromatin-localization studies support the nuclear methylation machinery, independently of individual high-throughput location rows.
Supporting Evidence:
PMID:23042785
co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005694 chromosome
EXP
PMID:12138111
A novel Dnmt3a isoform produced from an alternative promoter...
ACCEPT
Summary: DNMT3A associates with chromatin, including heterochromatic pools.
Reason: Isoform localization and chromatin-reader studies establish chromatin association. N-terminal isoforms differ in their distribution, so these gene-level location records should not be read as every isoform occupying identical chromatin.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
PMID:30478443
Substitutions in the PWWP domain abrogate binding to the histone modifications H3K36me2 and H3K36me3, and alter DNA methylation in patient cells.
GO:0005694 chromosome
IEA
GO_REF:0000044
ACCEPT
Summary: DNMT3A associates with chromatin, including heterochromatic pools.
Reason: Isoform localization and chromatin-reader studies establish chromatin association. N-terminal isoforms differ in their distribution, so these gene-level location records should not be read as every isoform occupying identical chromatin.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
PMID:30478443
Substitutions in the PWWP domain abrogate binding to the histone modifications H3K36me2 and H3K36me3, and alter DNA methylation in patient cells.
GO:0005737 cytoplasm
IBA
GO_REF:0000033
KEEP AS NON CORE
Summary: DNMT3A can occupy a cytoplasmic pool.
Reason: Retain the experimentally curated additional localization; its principal catalytic and chromatin-targeting role is nuclear. The source study also examines distinct isoforms, so the annotation does not mean all DNMT3A isoforms act primarily in the cytoplasm.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005737 cytoplasm
IDA
PMID:12138111
A novel Dnmt3a isoform produced from an alternative promoter...
KEEP AS NON CORE
Summary: DNMT3A can occupy a cytoplasmic pool.
Reason: Retain the experimentally curated additional localization; its principal catalytic and chromatin-targeting role is nuclear. The source study also examines distinct isoforms, so the annotation does not mean all DNMT3A isoforms act primarily in the cytoplasm.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0005737 cytoplasm
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: DNMT3A can occupy a cytoplasmic pool.
Reason: Retain the experimentally curated additional localization; its principal catalytic and chromatin-targeting role is nuclear. The source study also examines distinct isoforms, so the annotation does not mean all DNMT3A isoforms act primarily in the cytoplasm.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0006346 DNA methylation-dependent constitutive heterochromatin formation
IMP
PMID:30478443
Gain-of-function DNMT3A mutations cause microcephalic dwarfi...
ACCEPT
Summary: DNMT3A-dependent methylation changes chromatin silencing states.
Reason: The PWWP mutant study directly measures aberrant methylation of normally protected Polycomb-associated domains during differentiation, establishing a role in methylation-dependent chromatin organization.
Supporting Evidence:
PMID:30478443
Substitutions in the PWWP domain abrogate binding to the histone modifications H3K36me2 and H3K36me3, and alter DNA methylation in patient cells.
GO:0008168 methyltransferase activity
IEA
GO_REF:0000002
MODIFY
Summary: DNA cytosine-5 methyltransferase activity is the informative catalytic term.
Reason: Structures and assays resolve the methyl acceptor as DNA cytosine; replace the generic methyltransferase ancestor.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
GO:0009410 response to xenobiotic stimulus
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0009636 response to toxic substance
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0010212 response to ionizing radiation
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0010288 response to lead ion
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0010468 regulation of gene expression
IEA
GO_REF:0000002
ACCEPT
Summary: DNMT3A regulates gene expression as a recruited epigenetic corepressor.
Reason: Interaction with transcriptional regulators targets methylation and can repress promoter activity. Its enzymatic mechanism provides the specificity beyond a generic corepressor label.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:23042785
DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
GO:0016363 nuclear matrix
IDA
PMID:12138111
A novel Dnmt3a isoform produced from an alternative promoter...
UNDECIDED
Summary: Nuclear-matrix association needs the original fractionation/localization details.
Reason: The source study establishes chromatin and isoform-dependent localization; the accessible abstract does not resolve the separate nuclear-matrix fraction. Nuclear residence alone is insufficient.
Supporting Evidence:
PMID:12138111
Unlike Dnmt3a, which was concentrated on heterochromatin, Dnmt3a2 displayed a localization pattern suggestive of euchromatin association.
GO:0030182 neuron differentiation
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0031048 regulatory ncRNA-mediated heterochromatin formation
IGI
PMID:23435381
A pseudogene long-noncoding-RNA network regulates PTEN trans...
ACCEPT
Summary: DNMT3A associates with an antisense lncRNA in a repressive chromatin pathway.
Reason: The primary human study uses RNA immunoprecipitation and perturbation to connect DNMT3A with PTENpg1 antisense RNA and PTEN repression. Co-immunoprecipitation establishes complex association rather than purified direct RNA affinity.
Supporting Evidence:
PMID:23435381
RNA immunoprecipitation (IP) of DNMT3a showed DNMT3a and PTENpg1 asRNA Ξ± to co-IP and thus interact within the same protein:RNA complex
GO:0031667 response to nutrient levels
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0032355 response to estradiol
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0033189 response to vitamin A
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0042220 response to cocaine
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0042802 identical protein binding
IPI
PMID:24656771
The R882H DNMT3A mutation associated with AML dominantly inh...
ACCEPT
Summary: DNMT3A self-association is integral to catalysis.
Reason: Mutant/WT coexpression perturbs homotetramerization and methylation, while DNA-bound structures show the homodimeric DNA-recognition interface.
Supporting Evidence:
PMID:24656771
coexpression of the two proteins in cells profoundly inhibits the WT enzyme by disrupting its ability to homotetramerize.
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
GO:0044027 negative regulation of gene expression via chromosomal CpG island methylation
IDA
PMID:12138111
A novel Dnmt3a isoform produced from an alternative promoter...
ACCEPT
Summary: DNMT3A can repress transcription through targeted DNA methylation.
Reason: Recruitment by transcriptional regulators and promoter hypermethylation provide mechanistic support for negative transcriptional regulation. This does not imply that all DNMT3A-dependent methylation universally represses transcription.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:23042785
DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
GO:0044027 negative regulation of gene expression via chromosomal CpG island methylation
IDA
PMID:19786833
Dnmt3/transcription factor interactions as crucial players i...
ACCEPT
Summary: DNMT3A can repress transcription through targeted DNA methylation.
Reason: Recruitment by transcriptional regulators and promoter hypermethylation provide mechanistic support for negative transcriptional regulation. This does not imply that all DNMT3A-dependent methylation universally represses transcription.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:23042785
DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
GO:0044027 negative regulation of gene expression via chromosomal CpG island methylation
IDA
PMID:23042785
Control of epigenetic states by WT1 via regulation of de nov...
ACCEPT
Summary: DNMT3A can repress transcription through targeted DNA methylation.
Reason: Recruitment by transcriptional regulators and promoter hypermethylation provide mechanistic support for negative transcriptional regulation. This does not imply that all DNMT3A-dependent methylation universally represses transcription.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:23042785
DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
GO:0045322 unmethylated CpG binding
EXP
PMID:29414941
Structural basis for DNMT3A-mediated de novo DNA methylation...
ACCEPT
Summary: DNMT3A binds DNA and catalyzes cytosine methylation.
Reason: DNA-bound structures and substrate assays establish catalytic recognition of unmethylated CpG DNA and the de novo methyltransferase reaction.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:29414941
Arg836 of the target recognition domain makes crucial contacts with CpG, ensuring DNMT3A enzymatic preference towards CpG sites in cells.
GO:0045471 response to ethanol
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0045892 negative regulation of DNA-templated transcription
IBA
GO_REF:0000033
ACCEPT
Summary: DNMT3A can repress transcription through targeted DNA methylation.
Reason: Recruitment by transcriptional regulators and promoter hypermethylation provide mechanistic support for negative transcriptional regulation. This does not imply that all DNMT3A-dependent methylation universally represses transcription.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
PMID:23042785
DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
GO:0061629 RNA polymerase II-specific DNA-binding transcription factor binding
IPI
PMID:19786833
Dnmt3/transcription factor interactions as crucial players i...
ACCEPT
Summary: Transcription-factor binding helps target DNMT3A.
Reason: The MYC recruitment and interaction study directly connects transcription-factor contacts to site-selective promoter methylation.
Supporting Evidence:
PMID:19786833
the invalidation of c-myc reveals that c-myc allows the Dnmt3a recruitment on c-myc box of c-myc-regulated genes.
GO:0071361 cellular response to ethanol
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0071456 cellular response to hypoxia
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0097284 hepatocyte apoptotic process
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:0106222 lncRNA binding
IDA
PMID:23435381
A pseudogene long-noncoding-RNA network regulates PTEN trans...
ACCEPT
Summary: DNMT3A associates with an antisense lncRNA in a repressive chromatin pathway.
Reason: The primary human study uses RNA immunoprecipitation and perturbation to connect DNMT3A with PTENpg1 antisense RNA and PTEN repression. Co-immunoprecipitation establishes complex association rather than purified direct RNA affinity.
Supporting Evidence:
PMID:23435381
RNA immunoprecipitation (IP) of DNMT3a showed DNMT3a and PTENpg1 asRNA Ξ± to co-IP and thus interact within the same protein:RNA complex
GO:0106222 lncRNA binding
IEA
GO_REF:0000107
ACCEPT
Summary: DNMT3A associates with an antisense lncRNA in a repressive chromatin pathway.
Reason: The primary human study uses RNA immunoprecipitation and perturbation to connect DNMT3A with PTENpg1 antisense RNA and PTEN repression. Co-immunoprecipitation establishes complex association rather than purified direct RNA affinity.
Supporting Evidence:
PMID:23435381
RNA immunoprecipitation (IP) of DNMT3a showed DNMT3a and PTENpg1 asRNA Ξ± to co-IP and thus interact within the same protein:RNA complex
GO:0106363 protein-cysteine methyltransferase activity
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: Cysteine automethylation is an observed biochemical side reaction.
Reason: The mouse catalytic-domain study demonstrates slow automethylation in the absence of DNA; natural CpG DNA suppresses this reaction. It does not establish an evolved protein-methylation function in vivo. Retain as non-core biochemical capacity, not a second central catalytic role.
Supporting Evidence:
PMID:21481189
In the presence of CpG containing dsDNA, which is the natural substrate for Dnmt3a, the transfer of the methyl group from AdoMet to the flipped target base was preferred and auto-methylation was not detected.
GO:0106363 protein-cysteine methyltransferase activity
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: Cysteine automethylation is an observed biochemical side reaction.
Reason: The mouse catalytic-domain study demonstrates slow automethylation in the absence of DNA; natural CpG DNA suppresses this reaction. It does not establish an evolved protein-methylation function in vivo. Retain as non-core biochemical capacity, not a second central catalytic role.
Supporting Evidence:
PMID:21481189
In the presence of CpG containing dsDNA, which is the natural substrate for Dnmt3a, the transfer of the methyl group from AdoMet to the flipped target base was preferred and auto-methylation was not detected.
GO:0141196 transposable element silencing by piRNA-mediated DNA methylation
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:1900039 positive regulation of cellular response to hypoxia
IEA
GO_REF:0000107
UNDECIDED
Summary: This context-specific transfer needs its underlying experimental evidence.
Reason: The general role in epigenetic gene regulation does not independently establish this particular exposure response, developmental context or germline pathway. Trace the source ortholog experiment and distinguish causal participation from altered DNMT3A expression or methylation as a readout.
GO:1902494 catalytic complex
IPI
PMID:25383530
Structural insight into autoinhibition and histone H3-induce...
ACCEPT
Summary: DNMT3A participates in a regulated methyltransferase complex.
Reason: The DNMT3A-DNMT3L and histone-bound structures establish physical complex formation and catalytic regulation. Generic interaction annotations here are mechanistically grounded by the same structural study.
Supporting Evidence:
PMID:29414941
the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
PMID:25383530
Histone H3 (but not H3K4me3) disrupts ADD-CD interaction, induces a large movement of the ADD domain, and thus releases the autoinhibition of DNMT3A.

Core Functions

Methylates DNA cytosines to regulate chromatin and gene expression.

Supporting Evidence:
  • PMID:29414941
    the DNMT3A-DNMT3L-DNA complex in which two DNMT3A monomers simultaneously attack two cytosine-phosphate-guanine (CpG) dinucleotides
  • PMID:23042785
    DNMT3A recruitment is increased, with hypermethylation being associated with silencing of gene expression.
  • PMID:23042785
    co-expression of WT1 and DNMT3A proteins in nuclei of blastemal cells in human fetal kidney and Wilms' tumours.

References

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Deep Research

Falcon

(DNMT3A-deep-research-falcon.md)

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πŸ“š Additional Documentation

Notes

(DNMT3A-notes.md)

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