SORL1

UniProt ID: Q92673
Organism: Homo sapiens
Review Status: COMPLETE
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Gene Description

SORL1 encodes sortilin-related receptor, a large type I membrane receptor of the VPS10P/LDLR family that cycles through the trans-Golgi network, endosomes, recycling endosomes, and the plasma membrane. It binds and sorts cargo proteins and peptides, including amyloid precursor protein, amyloid-beta peptides, lipoprotein-related ligands, neurotrophic factors and receptors, insulin receptor, and cytokine-receptor complexes. In neurons, SORL1 retains and retrieves APP-containing complexes through Golgi/endosomal routes and can direct amyloid-beta toward lysosomal degradation, thereby reducing amyloidogenic APP processing. Other cargo-specific roles in lipid metabolism, trophic signaling, cytokine signaling, vascular smooth muscle behavior, and receptor recycling reflect the same receptor-mediated sorting biology in different cell contexts.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0005794 Golgi apparatus
IBA
GO_REF:0000033
ACCEPT
Summary: SorLA resides in the Golgi where, with AP-1, it executes TGN-endosome sorting; a major steady-state pool of the receptor is Golgi-associated.
Reason: Golgi is a core compartment for SorLA's cargo-sorting cycle (UniProt subcellular location).
GO:0006892 post-Golgi vesicle-mediated transport
IBA
GO_REF:0000033
ACCEPT
Summary: SorLA participates in vesicular transport from the TGN onward, dispatching itself and cargo to endosomes and the cell surface.
Reason: Core trafficking step in SorLA's sorting itinerary (IBA/IDA).
GO:0016020 membrane
IBA
GO_REF:0000033
ACCEPT
Summary: Generic membrane association reflects SorLA's nature as an integral single-pass type I membrane protein distributed across the secretory/endosomal system.
Reason: Correct but generic; consistent with the transmembrane receptor.
GO:0006622 protein targeting to lysosome
IBA
GO_REF:0000033
ACCEPT
Summary: SorLA routes select cargo (LPL, GDNF, IL6, APOA5, amyloid-beta) to lysosomes for degradation, a defined branch of its sorting output.
Reason: Core degradative sorting branch of the receptor (IBA/IDA).
GO:0045053 protein retention in Golgi apparatus
IBA
GO_REF:0000033
ACCEPT
Summary: SorLA retains APP (and other cargo) in the TGN, the key event that limits APP's transit to amyloidogenic late-endosomal processing.
Reason: Defining sorting outcome that links SorLA to reduced amyloidogenesis (IBA/IDA).
GO:0000139 Golgi membrane
IEA
GO_REF:0000044
ACCEPT
Summary: As a single-pass type I membrane protein, SorLA sits in the Golgi/TGN membrane where it engages adaptors (GGA, AP-1) and retains cargo such as APP.
Reason: Correct membrane sub-localization for a transmembrane Golgi sorting receptor.
GO:0005041 low-density lipoprotein particle receptor activity
IEA
GO_REF:0000117
ACCEPT
Summary: SorLA functions as an LDL-receptor-family member with complement-type ligand-binding repeats that bind lipoprotein and LRPAP1/RAP ligands.
Reason: Core receptor activity defining SorLA's LDLR-family identity.
GO:0005576 extracellular region
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: ADAM17-mediated shedding releases the soluble SorLA ectodomain (sLR11) into the extracellular space, where it acts as a secreted ligand-binding fragment.
Reason: Reflects shed soluble ectodomain, a secondary product of the membrane receptor.
GO:0005764 lysosome
IEA
GO_REF:0000108
KEEP AS NON CORE
Summary: SorLA routes select cargo (LPL, GDNF, APOA5, amyloid-beta) toward lysosomes for catabolism; the lysosome is a destination of its degradative sorting branch.
Reason: Terminal destination compartment downstream of SorLA's sorting decisions.
GO:0005789 endoplasmic reticulum membrane
IEA
GO_REF:0000044
KEEP AS NON CORE
Summary: ER-membrane residence reflects the newly synthesized receptor before propeptide cleavage and export; not where SorLA performs cargo sorting.
Reason: Biosynthetic localization, peripheral to the endosome/TGN sorting role.
GO:0005794 Golgi apparatus
IEA
GO_REF:0000044
ACCEPT
Summary: SorLA resides in the Golgi where, with AP-1, it executes TGN-endosome sorting; a major steady-state pool of the receptor is Golgi-associated.
Reason: Golgi is a core compartment for SorLA's cargo-sorting cycle (UniProt subcellular location).
GO:0005829 cytosol
IEA
GO_REF:0000108
MARK AS OVER ANNOTATED
Summary: SorLA is a single-pass type I membrane protein; only a small RIP-released C-terminal fragment is cytosolic, so a bulk cytosol annotation overstates its distribution.
Reason: IEA cytosol overstates localization of an integral membrane protein.
GO:0005886 plasma membrane
IEA
GO_REF:0000044
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:0010008 endosome membrane
IEA
GO_REF:0000044
ACCEPT
Summary: The transmembrane receptor is embedded in the endosomal limiting membrane, where its cytosolic tail binds retromer/GGA adaptors for sorting.
Reason: Membrane localization underpins SorLA's endosomal sorting function.
GO:0016020 membrane
IEA
GO_REF:0000120
ACCEPT
Summary: Generic membrane association reflects SorLA's nature as an integral single-pass type I membrane protein distributed across the secretory/endosomal system.
Reason: Correct but generic; consistent with the transmembrane receptor.
GO:0030658 transport vesicle membrane
IEA
GO_REF:0000044
ACCEPT
Summary: SorLA is carried in transport/secretory and endosome-to-TGN retrograde vesicles that shuttle it and its cargo between compartments.
Reason: Vesicle-membrane localization matches its trafficking itinerary (UniProt: retrograde vesicles).
GO:0031901 early endosome membrane
IEA
GO_REF:0000044
ACCEPT
Summary: SorLA occupies the early-endosome membrane where its cytoplasmic domain recruits sorting adaptors for cargo retrieval.
Reason: Membrane sub-localization supporting endosomal cargo sorting.
GO:0032585 multivesicular body membrane
IEA
GO_REF:0000044
KEEP AS NON CORE
Summary: MVB-membrane localization is consistent with SorLA delivering cargo to the lysosomal degradation pathway.
Reason: Late-endosomal membrane, secondary to the core TGN/endosome retrieval role.
GO:0055038 recycling endosome membrane
IEA
GO_REF:0000044
ACCEPT
Summary: Membrane localization to recycling endosomes is consistent with SorLA promoting endocytic recycling of internalized receptors.
Reason: Membrane sub-localization for SorLA's recycling function.
GO:0005515 protein binding
IPI
PMID:11821067
The sorLA cytoplasmic domain interacts with GGA1 and -2 and ...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:16174740
Neuronal sorting protein-related receptor sorLA/LR11 regulat...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:16407538
Interaction of the cytosolic domains of sorLA/LR11 with the ...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:17326667
Apolipoprotein A-V interaction with members of the low densi...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:23333276
SorLA controls neurotrophic activity by sorting of GDNF and ...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:25643321
Structural basis for amyloidogenic peptide recognition by so...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:26053850
The Golgi-Localized Ξ³-Ear-Containing ARF-Binding (GGA) Prote...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:26858303
Cytokine-Like Factor 1, an Essential Facilitator of Cardiotr...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:28265003
SorLA in Interleukin-6 Signaling and Turnover.
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:28514442
Architecture of the human interactome defines protein commun...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:30679749
HSPA12A targets the cytoplasmic domain and affects the traff...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:32814053
Interactome Mapping Provides a Network of Neurodegenerative ...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005515 protein binding
IPI
PMID:33961781
Dual proteome-scale networks reveal cell-specific remodeling...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0002024 diet induced thermogenesis
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: SorLA's metabolic regulation of adipose energy expenditure extends to diet-induced thermogenesis, a systemic consequence of its ectodomain activity.
Reason: Pleiotropic energy-balance phenotype, downstream of metabolic signaling roles.
GO:0005641 nuclear envelope lumen
IEA
GO_REF:0000107
UNDECIDED
Summary: RIP releases a SorLA C-terminal fragment that localizes to the nucleus, but a specific nuclear-envelope-lumen assignment is not clearly supported by the evidence reviewed.
Reason: Cannot verify the nuclear-envelope-lumen call; nuclear targeting is only of a cleaved fragment.
GO:0005771 multivesicular body
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: SorLA is seen in MVBs/late endosomes as part of cargo routing toward lysosomal degradation (e.g. LPL, GDNF).
Reason: Downstream late-endosomal station of the degradative arm of SorLA sorting.
GO:0006605 protein targeting
IEA
GO_REF:0000107
ACCEPT
Summary: SorLA directs bound client proteins to specific destinations (TGN retention, lysosomal degradation or surface recycling), the essence of its sorting role.
Reason: Core process: directing proteins to their correct intracellular location.
GO:0010897 negative regulation of triglyceride catabolic process
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: By trafficking LPL to lysosomes and degrading it, SorLA reduces lipase activity and triglyceride catabolism; a metabolic consequence of degradative sorting.
Reason: Lipid-metabolism effect downstream of SorLA-mediated LPL turnover.
GO:0016477 cell migration
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: SorLA modulates migration of smooth-muscle and myeloid cells through uPAR/PLAUR surface regulation, a pleiotropic consequence of its receptor trafficking.
Reason: Generic pleiotropic migration role secondary to ligand trafficking.
GO:0019828 aspartic-type endopeptidase inhibitor activity
IEA
GO_REF:0000107
MODIFY
Summary: SorLA reduces BACE1-dependent APP cleavage by sequestering APP and interfering with BACE1-APP encounter; this is competitive cargo sorting, not direct aspartic-protease inhibition.
Reason: Effect on BACE1 is indirect via trafficking; better captured as negative regulation of amyloid-beta formation.
GO:0030514 negative regulation of BMP signaling pathway
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: The SorLA N-terminal ectodomain inhibits BMP/Smad signaling in adipose tissue to restrain energy expenditure; a metabolic, ectodomain-mediated role.
Reason: Pleiotropic metabolic signaling effect of the soluble ectodomain (IEA/ISS).
GO:0038020 insulin receptor recycling
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: SorLA recycles internalized INSR via the Golgi to the surface, raising INSR surface levels in adipocytes; a tissue-specific instance of its recycling-sorting role.
Reason: Cargo-specific recycling event downstream of SorLA's general recycling activity.
GO:0043025 neuronal cell body
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: SorLA is enriched in neuronal soma, consistent with its high brain expression and role in neuronal cargo (APP, TrkB, GDNF) trafficking.
Reason: Cell-type-specific localization reflecting neuronal expression, not a distinct core compartment.
GO:0045053 protein retention in Golgi apparatus
IEA
GO_REF:0000107
ACCEPT
Summary: SorLA retains APP (and other cargo) in the TGN, the key event that limits APP's transit to amyloidogenic late-endosomal processing.
Reason: Defining sorting outcome that links SorLA to reduced amyloidogenesis (IBA/IDA).
GO:0046628 positive regulation of insulin receptor signaling pathway
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: By increasing INSR surface expression through recycling, SorLA strengthens insulin signaling in adipose tissue, a pleiotropic downstream consequence.
Reason: Signaling effect secondary to SorLA-mediated INSR recycling.
GO:0050768 negative regulation of neurogenesis
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: SorLA influences neuronal development consistent with its high brain expression and trafficking of neurotrophic cargo (BDNF/TrkB, GDNF).
Reason: Developmental role downstream of neurotrophic-receptor sorting (IEA/ISS).
GO:0097356 perinucleolar compartment
IEA
GO_REF:0000107
MARK AS OVER ANNOTATED
Summary: No evidence places this transmembrane sorting receptor in the perinucleolar compartment; the IEA call is implausible for SorLA.
Reason: Implausible nuclear sub-compartment for a membrane sorting receptor; electronic over-annotation.
GO:0099638 endosome to plasma membrane protein transport
IEA
GO_REF:0000107
ACCEPT
Summary: SorLA promotes return of internalized receptors (INSR, HER2) from endosomes to the plasma membrane, a recycling arm of its sorting activity.
Reason: Core recycling route mediated by SorLA (UniProt: HER2/INSR recycling).
GO:1900168 positive regulation of glial cell-derived neurotrophic factor production
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: SorLA controls regulated (vs constitutive) GDNF secretion by sorting GDNF and its receptors, influencing GDNF availability; a neurotrophic-specific role.
Reason: Cargo-specific neurotrophic effect downstream of GDNF sorting.
GO:1902430 negative regulation of amyloid-beta formation
IEA
GO_REF:0000107
ACCEPT
Summary: SorLA lowers Abeta40/Abeta42 generation by sequestering APP from amyloidogenic processing and routing Abeta to lysosomes, the basis of its AD-protective role.
Reason: Central disease-relevant output directly downstream of SorLA APP sorting (IDA/IMP).
GO:1902992 negative regulation of amyloid precursor protein catabolic process
IEA
GO_REF:0000107
ACCEPT
Summary: By retaining APP in the TGN and limiting its access to secretases, SorLA reduces amyloidogenic APP catabolism, the best-characterized physiological output of its sorting.
Reason: Hallmark functional consequence of SorLA's APP retention (IDA/IMP).
GO:1902997 negative regulation of neurofibrillary tangle assembly
IEA
GO_REF:0000107
MARK AS OVER ANNOTATED
Summary: SorLA's primary action is on APP/Abeta trafficking; an effect on tau neurofibrillary-tangle assembly is at best indirect and rests only on electronic/ISS inference.
Reason: Tau-tangle effect is speculative and indirect; electronic over-annotation.
GO:1904179 positive regulation of adipose tissue development
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Through its metabolic actions (INSR recycling, BMP inhibition, LPL turnover) SorLA influences adipose tissue development; a pleiotropic systemic role.
Reason: Systemic metabolic/developmental role secondary to its trafficking functions.
GO:1990845 adaptive thermogenesis
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Soluble SorLA represses adipose thermogenesis via BMP/Smad inhibition, shifting energy balance toward storage; a metabolic role of the ectodomain.
Reason: Pleiotropic metabolic phenotype of the shed ectodomain (IEA/ISS).
GO:0038024 cargo receptor activity
IDA
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: This is SorLA's defining activity: a VPS10P-domain cargo receptor that binds and sorts client proteins (APP, LPL, INSR, HER2, GDNF) among the TGN, endosomes and surface.
Reason: Core defining molecular function of SorLA as an intracellular sorting receptor.
GO:0038024 cargo receptor activity
IDA
PMID:11294867
Activation and functional characterization of the mosaic rec...
ACCEPT
Summary: This is SorLA's defining activity: a VPS10P-domain cargo receptor that binds and sorts client proteins (APP, LPL, INSR, HER2, GDNF) among the TGN, endosomes and surface.
Reason: Core defining molecular function of SorLA as an intracellular sorting receptor.
GO:0038024 cargo receptor activity
TAS
PMID:9157966
A novel mosaic protein containing LDL receptor elements is h...
ACCEPT
Summary: This is SorLA's defining activity: a VPS10P-domain cargo receptor that binds and sorts client proteins (APP, LPL, INSR, HER2, GDNF) among the TGN, endosomes and surface.
Reason: Core defining molecular function of SorLA as an intracellular sorting receptor.
GO:0000139 Golgi membrane
EXP
PMID:11294867
Activation and functional characterization of the mosaic rec...
ACCEPT
Summary: As a single-pass type I membrane protein, SorLA sits in the Golgi/TGN membrane where it engages adaptors (GGA, AP-1) and retains cargo such as APP.
Reason: Correct membrane sub-localization for a transmembrane Golgi sorting receptor.
GO:0000139 Golgi membrane
EXP
PMID:16174740
Neuronal sorting protein-related receptor sorLA/LR11 regulat...
ACCEPT
Summary: As a single-pass type I membrane protein, SorLA sits in the Golgi/TGN membrane where it engages adaptors (GGA, AP-1) and retains cargo such as APP.
Reason: Correct membrane sub-localization for a transmembrane Golgi sorting receptor.
GO:0000139 Golgi membrane
EXP
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: As a single-pass type I membrane protein, SorLA sits in the Golgi/TGN membrane where it engages adaptors (GGA, AP-1) and retains cargo such as APP.
Reason: Correct membrane sub-localization for a transmembrane Golgi sorting receptor.
GO:0000139 Golgi membrane
EXP
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
ACCEPT
Summary: As a single-pass type I membrane protein, SorLA sits in the Golgi/TGN membrane where it engages adaptors (GGA, AP-1) and retains cargo such as APP.
Reason: Correct membrane sub-localization for a transmembrane Golgi sorting receptor.
GO:0000139 Golgi membrane
EXP
PMID:21994944
Sorting protein-related receptor SorLA controls regulated se...
ACCEPT
Summary: As a single-pass type I membrane protein, SorLA sits in the Golgi/TGN membrane where it engages adaptors (GGA, AP-1) and retains cargo such as APP.
Reason: Correct membrane sub-localization for a transmembrane Golgi sorting receptor.
GO:0005576 extracellular region
EXP
PMID:11082041
Ectodomain shedding, translocation and synthesis of SorLA ar...
KEEP AS NON CORE
Summary: ADAM17-mediated shedding releases the soluble SorLA ectodomain (sLR11) into the extracellular space, where it acts as a secreted ligand-binding fragment.
Reason: Reflects shed soluble ectodomain, a secondary product of the membrane receptor.
GO:0005576 extracellular region
EXP
PMID:15053742
The mosaic receptor sorLA/LR11 binds components of the plasm...
KEEP AS NON CORE
Summary: ADAM17-mediated shedding releases the soluble SorLA ectodomain (sLR11) into the extracellular space, where it acts as a secreted ligand-binding fragment.
Reason: Reflects shed soluble ectodomain, a secondary product of the membrane receptor.
GO:0005576 extracellular region
EXP
PMID:16393139
Tumour necrosis factor alpha-converting enzyme mediates ecto...
KEEP AS NON CORE
Summary: ADAM17-mediated shedding releases the soluble SorLA ectodomain (sLR11) into the extracellular space, where it acts as a secreted ligand-binding fragment.
Reason: Reflects shed soluble ectodomain, a secondary product of the membrane receptor.
GO:0005576 extracellular region
EXP
PMID:16531402
SorLA signaling by regulated intramembrane proteolysis.
KEEP AS NON CORE
Summary: ADAM17-mediated shedding releases the soluble SorLA ectodomain (sLR11) into the extracellular space, where it acts as a secreted ligand-binding fragment.
Reason: Reflects shed soluble ectodomain, a secondary product of the membrane receptor.
GO:0005789 endoplasmic reticulum membrane
EXP
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
KEEP AS NON CORE
Summary: ER-membrane residence reflects the newly synthesized receptor before propeptide cleavage and export; not where SorLA performs cargo sorting.
Reason: Biosynthetic localization, peripheral to the endosome/TGN sorting role.
GO:0005789 endoplasmic reticulum membrane
EXP
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
KEEP AS NON CORE
Summary: ER-membrane residence reflects the newly synthesized receptor before propeptide cleavage and export; not where SorLA performs cargo sorting.
Reason: Biosynthetic localization, peripheral to the endosome/TGN sorting role.
GO:0005886 plasma membrane
EXP
PMID:11294867
Activation and functional characterization of the mosaic rec...
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:0005886 plasma membrane
EXP
PMID:14764453
LR11, an LDL receptor gene family member, is a novel regulat...
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:0005886 plasma membrane
EXP
PMID:15053742
The mosaic receptor sorLA/LR11 binds components of the plasm...
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:0005886 plasma membrane
EXP
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:0005886 plasma membrane
EXP
PMID:31138794
SORLA regulates endosomal trafficking and oncogenic fitness ...
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:0010008 endosome membrane
EXP
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
ACCEPT
Summary: The transmembrane receptor is embedded in the endosomal limiting membrane, where its cytosolic tail binds retromer/GGA adaptors for sorting.
Reason: Membrane localization underpins SorLA's endosomal sorting function.
GO:0010008 endosome membrane
EXP
PMID:23333276
SorLA controls neurotrophic activity by sorting of GDNF and ...
ACCEPT
Summary: The transmembrane receptor is embedded in the endosomal limiting membrane, where its cytosolic tail binds retromer/GGA adaptors for sorting.
Reason: Membrane localization underpins SorLA's endosomal sorting function.
GO:0031901 early endosome membrane
EXP
PMID:17646382
Sorting by the cytoplasmic domain of the amyloid precursor p...
ACCEPT
Summary: SorLA occupies the early-endosome membrane where its cytoplasmic domain recruits sorting adaptors for cargo retrieval.
Reason: Membrane sub-localization supporting endosomal cargo sorting.
GO:0031901 early endosome membrane
EXP
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
ACCEPT
Summary: SorLA occupies the early-endosome membrane where its cytoplasmic domain recruits sorting adaptors for cargo retrieval.
Reason: Membrane sub-localization supporting endosomal cargo sorting.
GO:0031901 early endosome membrane
EXP
PMID:31138794
SORLA regulates endosomal trafficking and oncogenic fitness ...
ACCEPT
Summary: SorLA occupies the early-endosome membrane where its cytoplasmic domain recruits sorting adaptors for cargo retrieval.
Reason: Membrane sub-localization supporting endosomal cargo sorting.
GO:0055038 recycling endosome membrane
EXP
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: Membrane localization to recycling endosomes is consistent with SorLA promoting endocytic recycling of internalized receptors.
Reason: Membrane sub-localization for SorLA's recycling function.
GO:0055038 recycling endosome membrane
EXP
PMID:31138794
SORLA regulates endosomal trafficking and oncogenic fitness ...
ACCEPT
Summary: Membrane localization to recycling endosomes is consistent with SorLA promoting endocytic recycling of internalized receptors.
Reason: Membrane sub-localization for SorLA's recycling function.
GO:0001960 negative regulation of cytokine-mediated signaling pathway
IDA
PMID:26858303
Cytokine-Like Factor 1, an Essential Facilitator of Cardiotr...
KEEP AS NON CORE
Summary: SorLA dampens cytokine signaling by internalizing and degrading IL6/CLCF1-CRLF1-CNTFR complexes, a downstream effect of its endocytic-degradative sorting.
Reason: Signaling modulation secondary to SorLA cargo internalization and turnover.
GO:0005886 plasma membrane
IDA
PMID:26858303
Cytokine-Like Factor 1, an Essential Facilitator of Cardiotr...
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:0038024 cargo receptor activity
IDA
PMID:26858303
Cytokine-Like Factor 1, an Essential Facilitator of Cardiotr...
ACCEPT
Summary: This is SorLA's defining activity: a VPS10P-domain cargo receptor that binds and sorts client proteins (APP, LPL, INSR, HER2, GDNF) among the TGN, endosomes and surface.
Reason: Core defining molecular function of SorLA as an intracellular sorting receptor.
GO:1905146 lysosomal protein catabolic process
IDA
PMID:26858303
Cytokine-Like Factor 1, an Essential Facilitator of Cardiotr...
KEEP AS NON CORE
Summary: Lysosomal degradation of SorLA-delivered cargo is a downstream consequence of its degradative sorting branch rather than the proximal sorting activity itself.
Reason: Downstream catabolic outcome of SorLA's lysosomal targeting.
GO:0031901 early endosome membrane
IDA
PMID:16174740
Neuronal sorting protein-related receptor sorLA/LR11 regulat...
ACCEPT
Summary: SorLA occupies the early-endosome membrane where its cytoplasmic domain recruits sorting adaptors for cargo retrieval.
Reason: Membrane sub-localization supporting endosomal cargo sorting.
GO:0042147 retrograde transport, endosome to Golgi
IDA
PMID:16174740
Neuronal sorting protein-related receptor sorLA/LR11 regulat...
ACCEPT
Summary: SorLA's cytosolic tail engages retromer (VPS29) and GGA adaptors to retrieve itself and cargo from endosomes back to the TGN.
Reason: Central retrieval route defining SorLA's sorting behaviour (IDA).
GO:0140318 protein transporter activity
IDA
PMID:16174740
Neuronal sorting protein-related receptor sorLA/LR11 regulat...
ACCEPT
Summary: SorLA physically carries bound cargo proteins between compartments, consistent with its cargo-receptor role in directing protein localization.
Reason: Captures the cargo-carrying activity central to SorLA's sorting function (IDA).
GO:1902430 negative regulation of amyloid-beta formation
IDA
PMID:16174740
Neuronal sorting protein-related receptor sorLA/LR11 regulat...
ACCEPT
Summary: SorLA lowers Abeta40/Abeta42 generation by sequestering APP from amyloidogenic processing and routing Abeta to lysosomes, the basis of its AD-protective role.
Reason: Central disease-relevant output directly downstream of SorLA APP sorting (IDA/IMP).
GO:0019828 aspartic-type endopeptidase inhibitor activity
IDA
PMID:16407538
Interaction of the cytosolic domains of sorLA/LR11 with the ...
MODIFY
Summary: SorLA reduces BACE1-dependent APP cleavage by sequestering APP and interfering with BACE1-APP encounter; this is competitive cargo sorting, not direct aspartic-protease inhibition.
Reason: Effect on BACE1 is indirect via trafficking; better captured as negative regulation of amyloid-beta formation.
GO:0019828 aspartic-type endopeptidase inhibitor activity
IMP
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
MODIFY
Summary: SorLA reduces BACE1-dependent APP cleavage by sequestering APP and interfering with BACE1-APP encounter; this is competitive cargo sorting, not direct aspartic-protease inhibition.
Reason: Effect on BACE1 is indirect via trafficking; better captured as negative regulation of amyloid-beta formation.
GO:0019828 aspartic-type endopeptidase inhibitor activity
IMP
PMID:21989385
Quantitative modelling of amyloidogenic processing and its i...
MODIFY
Summary: SorLA reduces BACE1-dependent APP cleavage by sequestering APP and interfering with BACE1-APP encounter; this is competitive cargo sorting, not direct aspartic-protease inhibition.
Reason: Effect on BACE1 is indirect via trafficking; better captured as negative regulation of amyloid-beta formation.
GO:1902992 negative regulation of amyloid precursor protein catabolic process
IDA
PMID:16407538
Interaction of the cytosolic domains of sorLA/LR11 with the ...
ACCEPT
Summary: By retaining APP in the TGN and limiting its access to secretases, SorLA reduces amyloidogenic APP catabolism, the best-characterized physiological output of its sorting.
Reason: Hallmark functional consequence of SorLA's APP retention (IDA/IMP).
GO:1902992 negative regulation of amyloid precursor protein catabolic process
IMP
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: By retaining APP in the TGN and limiting its access to secretases, SorLA reduces amyloidogenic APP catabolism, the best-characterized physiological output of its sorting.
Reason: Hallmark functional consequence of SorLA's APP retention (IDA/IMP).
GO:1902992 negative regulation of amyloid precursor protein catabolic process
IMP
PMID:21989385
Quantitative modelling of amyloidogenic processing and its i...
ACCEPT
Summary: By retaining APP in the TGN and limiting its access to secretases, SorLA reduces amyloidogenic APP catabolism, the best-characterized physiological output of its sorting.
Reason: Hallmark functional consequence of SorLA's APP retention (IDA/IMP).
GO:0005515 protein binding
IPI
PMID:11294867
Activation and functional characterization of the mosaic rec...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005794 Golgi apparatus
IDA
PMID:11294867
Activation and functional characterization of the mosaic rec...
ACCEPT
Summary: SorLA resides in the Golgi where, with AP-1, it executes TGN-endosome sorting; a major steady-state pool of the receptor is Golgi-associated.
Reason: Golgi is a core compartment for SorLA's cargo-sorting cycle (UniProt subcellular location).
GO:0006898 receptor-mediated endocytosis
IDA
PMID:11294867
Activation and functional characterization of the mosaic rec...
ACCEPT
Summary: SorLA mediates AP-2/clathrin-dependent internalization of surface ligands (uPA-PAI1, APOA5, IL6, HER2), feeding them into its endosomal sorting itinerary.
Reason: Core uptake step of SorLA's cargo-sorting cycle (IDA/TAS).
GO:0007218 neuropeptide signaling pathway
IDA
PMID:11294867
Activation and functional characterization of the mosaic rec...
KEEP AS NON CORE
Summary: SorLA's interaction with neuropeptides (HA) links it to neuropeptide signaling, a downstream context of its ligand-binding capacity.
Reason: Signaling context downstream of SorLA neuropeptide binding.
GO:0009986 cell surface
IDA
PMID:11294867
Activation and functional characterization of the mosaic rec...
ACCEPT
Summary: Surface-exposed SorLA binds ligands (e.g. APP, IL6, uPAR) and mediates their internalization; the surface pool also undergoes ectodomain shedding.
Reason: Cell-surface presence is required for SorLA receptor-mediated endocytosis (IDA).
GO:0042923 neuropeptide binding
IPI
PMID:11082041
Ectodomain shedding, translocation and synthesis of SorLA ar...
KEEP AS NON CORE
Summary: The VPS10P domain binds neuropeptides such as head activator (HA) and neurotensin; a specific partner-binding activity peripheral to the core sorting role.
Reason: Partner-specific ligand binding; ancillary to the defining cargo-receptor activity.
GO:0042923 neuropeptide binding
IPI
PMID:11294867
Activation and functional characterization of the mosaic rec...
KEEP AS NON CORE
Summary: The VPS10P domain binds neuropeptides such as head activator (HA) and neurotensin; a specific partner-binding activity peripheral to the core sorting role.
Reason: Partner-specific ligand binding; ancillary to the defining cargo-receptor activity.
GO:0031333 negative regulation of protein-containing complex assembly
IMP
PMID:21989385
Quantitative modelling of amyloidogenic processing and its i...
KEEP AS NON CORE
Summary: By sequestering APP and modulating BACE1 access, SorLA can limit assembly of processing complexes; an indirect effect of its cargo-binding behaviour.
Reason: Indirect complex-assembly effect downstream of SorLA ligand sequestration (IMP).
GO:0005515 protein binding
IPI
PMID:21994944
Sorting protein-related receptor SorLA controls regulated se...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005794 Golgi apparatus
IDA
PMID:21994944
Sorting protein-related receptor SorLA controls regulated se...
ACCEPT
Summary: SorLA resides in the Golgi where, with AP-1, it executes TGN-endosome sorting; a major steady-state pool of the receptor is Golgi-associated.
Reason: Golgi is a core compartment for SorLA's cargo-sorting cycle (UniProt subcellular location).
GO:0005886 plasma membrane
IDA
PMID:21994944
Sorting protein-related receptor SorLA controls regulated se...
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:1900168 positive regulation of glial cell-derived neurotrophic factor production
IDA
PMID:21994944
Sorting protein-related receptor SorLA controls regulated se...
KEEP AS NON CORE
Summary: SorLA controls regulated (vs constitutive) GDNF secretion by sorting GDNF and its receptors, influencing GDNF availability; a neurotrophic-specific role.
Reason: Cargo-specific neurotrophic effect downstream of GDNF sorting.
GO:0005515 protein binding
IPI
PMID:30448281
Soluble LR11 competes with amyloid Ξ² in binding to cerebrosp...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0006605 protein targeting
IDA
PMID:27322061
SORLA facilitates insulin receptor signaling in adipocytes a...
ACCEPT
Summary: SorLA directs bound client proteins to specific destinations (TGN retention, lysosomal degradation or surface recycling), the essence of its sorting role.
Reason: Core process: directing proteins to their correct intracellular location.
GO:0010897 negative regulation of triglyceride catabolic process
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: By trafficking LPL to lysosomes and degrading it, SorLA reduces lipase activity and triglyceride catabolism; a metabolic consequence of degradative sorting.
Reason: Lipid-metabolism effect downstream of SorLA-mediated LPL turnover.
GO:0038020 insulin receptor recycling
IDA
PMID:27322061
SORLA facilitates insulin receptor signaling in adipocytes a...
KEEP AS NON CORE
Summary: SorLA recycles internalized INSR via the Golgi to the surface, raising INSR surface levels in adipocytes; a tissue-specific instance of its recycling-sorting role.
Reason: Cargo-specific recycling event downstream of SorLA's general recycling activity.
GO:0046628 positive regulation of insulin receptor signaling pathway
IDA
PMID:27322061
SORLA facilitates insulin receptor signaling in adipocytes a...
KEEP AS NON CORE
Summary: By increasing INSR surface expression through recycling, SorLA strengthens insulin signaling in adipose tissue, a pleiotropic downstream consequence.
Reason: Signaling effect secondary to SorLA-mediated INSR recycling.
GO:1904179 positive regulation of adipose tissue development
IDA
PMID:27322061
SORLA facilitates insulin receptor signaling in adipocytes a...
KEEP AS NON CORE
Summary: Through its metabolic actions (INSR recycling, BMP inhibition, LPL turnover) SorLA influences adipose tissue development; a pleiotropic systemic role.
Reason: Systemic metabolic/developmental role secondary to its trafficking functions.
GO:0002024 diet induced thermogenesis
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: SorLA's metabolic regulation of adipose energy expenditure extends to diet-induced thermogenesis, a systemic consequence of its ectodomain activity.
Reason: Pleiotropic energy-balance phenotype, downstream of metabolic signaling roles.
GO:0005515 protein binding
IPI
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005768 endosome
IDA
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
ACCEPT
Summary: SorLA cycles through endosomes, capturing internalized cargo (APP, LPL, INSR, HER2) for retrieval to the TGN or routing to lysosomes.
Reason: Endosome is a defining compartment of SorLA's retrieval/sorting itinerary.
GO:0005769 early endosome
IDA
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
ACCEPT
Summary: SorLA reaches early endosomes after internalization and directs cargo such as APOA5 and APP toward retrieval or onward lysosomal routing.
Reason: Early endosome is an early station of SorLA's sorting itinerary (IDA/IMP).
GO:0005771 multivesicular body
IDA
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
KEEP AS NON CORE
Summary: SorLA is seen in MVBs/late endosomes as part of cargo routing toward lysosomal degradation (e.g. LPL, GDNF).
Reason: Downstream late-endosomal station of the degradative arm of SorLA sorting.
GO:0005783 endoplasmic reticulum
IDA
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
KEEP AS NON CORE
Summary: SorLA transits the ER during biosynthesis and maturation, but its sorting activity acts downstream in the TGN/endosomal system rather than in the ER.
Reason: Biosynthetic transit compartment, upstream of and peripheral to the core sorting cycle.
GO:0005794 Golgi apparatus
IDA
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
ACCEPT
Summary: SorLA resides in the Golgi where, with AP-1, it executes TGN-endosome sorting; a major steady-state pool of the receptor is Golgi-associated.
Reason: Golgi is a core compartment for SorLA's cargo-sorting cycle (UniProt subcellular location).
GO:0005886 plasma membrane
IDA
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:0006605 protein targeting
IDA
PMID:21385844
SorLA regulates the activity of lipoprotein lipase by intrac...
ACCEPT
Summary: SorLA directs bound client proteins to specific destinations (TGN retention, lysosomal degradation or surface recycling), the essence of its sorting role.
Reason: Core process: directing proteins to their correct intracellular location.
GO:0030514 negative regulation of BMP signaling pathway
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: The SorLA N-terminal ectodomain inhibits BMP/Smad signaling in adipose tissue to restrain energy expenditure; a metabolic, ectodomain-mediated role.
Reason: Pleiotropic metabolic signaling effect of the soluble ectodomain (IEA/ISS).
GO:1990845 adaptive thermogenesis
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: Soluble SorLA represses adipose thermogenesis via BMP/Smad inhibition, shifting energy balance toward storage; a metabolic role of the ectodomain.
Reason: Pleiotropic metabolic phenotype of the shed ectodomain (IEA/ISS).
GO:0005041 low-density lipoprotein particle receptor activity
TAS
PMID:20005821
Overexpression of low-density lipoprotein receptor in the br...
ACCEPT
Summary: SorLA functions as an LDL-receptor-family member with complement-type ligand-binding repeats that bind lipoprotein and LRPAP1/RAP ligands.
Reason: Core receptor activity defining SorLA's LDLR-family identity.
GO:0000139 Golgi membrane
TAS
Reactome:R-HSA-8871506
ACCEPT
Summary: As a single-pass type I membrane protein, SorLA sits in the Golgi/TGN membrane where it engages adaptors (GGA, AP-1) and retains cargo such as APP.
Reason: Correct membrane sub-localization for a transmembrane Golgi sorting receptor.
GO:0010008 endosome membrane
TAS
Reactome:R-HSA-8871494
ACCEPT
Summary: The transmembrane receptor is embedded in the endosomal limiting membrane, where its cytosolic tail binds retromer/GGA adaptors for sorting.
Reason: Membrane localization underpins SorLA's endosomal sorting function.
GO:0010008 endosome membrane
TAS
Reactome:R-HSA-8871506
ACCEPT
Summary: The transmembrane receptor is embedded in the endosomal limiting membrane, where its cytosolic tail binds retromer/GGA adaptors for sorting.
Reason: Membrane localization underpins SorLA's endosomal sorting function.
GO:0050768 negative regulation of neurogenesis
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: SorLA influences neuronal development consistent with its high brain expression and trafficking of neurotrophic cargo (BDNF/TrkB, GDNF).
Reason: Developmental role downstream of neurotrophic-receptor sorting (IEA/ISS).
GO:1902997 negative regulation of neurofibrillary tangle assembly
ISS
GO_REF:0000024
MARK AS OVER ANNOTATED
Summary: SorLA's primary action is on APP/Abeta trafficking; an effect on tau neurofibrillary-tangle assembly is at best indirect and rests only on electronic/ISS inference.
Reason: Tau-tangle effect is speculative and indirect; electronic over-annotation.
GO:1902953 positive regulation of ER to Golgi vesicle-mediated transport
IMP
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: SorLA promotes anterograde ER-to-Golgi transport of cargo, complementing its retrieval activity in establishing correct cargo localization.
Reason: Trafficking-regulatory role consistent with its sorting function (IMP).
GO:1902966 positive regulation of protein localization to early endosome
IMP
PMID:22621900
GGA1-mediated endocytic traffic of LR11/SorLA alters APP int...
ACCEPT
Summary: SorLA drives delivery of cargo into early endosomes en route to retrieval or degradation, an integral step of its sorting cycle.
Reason: Direct manifestation of SorLA endosomal cargo sorting (IMP).
GO:0001540 amyloid-beta binding
IDA
PMID:24523320
Lysosomal sorting of amyloid-Ξ² by the SORLA receptor is impa...
ACCEPT
Summary: SorLA directly binds amyloid-beta (and APP C99/Abeta40) via its ectodomain, enabling capture of newly produced Abeta for lysosomal clearance.
Reason: Defining ligand interaction underlying SorLA's anti-amyloidogenic sorting (IDA/IPI).
GO:0005515 protein binding
IPI
PMID:24001769
SORLA-dependent and -independent functions for PACS1 in cont...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005769 early endosome
IMP
PMID:22621900
GGA1-mediated endocytic traffic of LR11/SorLA alters APP int...
ACCEPT
Summary: SorLA reaches early endosomes after internalization and directs cargo such as APOA5 and APP toward retrieval or onward lysosomal routing.
Reason: Early endosome is an early station of SorLA's sorting itinerary (IDA/IMP).
GO:0005769 early endosome
IDA
PMID:24001769
SORLA-dependent and -independent functions for PACS1 in cont...
ACCEPT
Summary: SorLA reaches early endosomes after internalization and directs cargo such as APOA5 and APP toward retrieval or onward lysosomal routing.
Reason: Early endosome is an early station of SorLA's sorting itinerary (IDA/IMP).
GO:0005802 trans-Golgi network
IDA
PMID:24001769
SORLA-dependent and -independent functions for PACS1 in cont...
ACCEPT
Summary: SorLA is predominantly localized to the TGN, the compartment where it retains APP and from which it dispatches cargo to endosomes.
Reason: TGN is the principal steady-state site of this sorting receptor (UniProt: 'predominantly in the TGN').
GO:0006622 protein targeting to lysosome
IDA
PMID:24523320
Lysosomal sorting of amyloid-Ξ² by the SORLA receptor is impa...
ACCEPT
Summary: SorLA routes select cargo (LPL, GDNF, IL6, APOA5, amyloid-beta) to lysosomes for degradation, a defined branch of its sorting output.
Reason: Core degradative sorting branch of the receptor (IBA/IDA).
GO:0045732 positive regulation of protein catabolic process
IDA
PMID:24523320
Lysosomal sorting of amyloid-Ξ² by the SORLA receptor is impa...
ACCEPT
Summary: By routing clients (LPL, GDNF, IL6, amyloid-beta) to lysosomes, SorLA promotes their catabolism as a downstream output of degradative sorting.
Reason: Direct outcome of SorLA's lysosomal targeting branch (IDA).
GO:0055037 recycling endosome
IMP
PMID:22621900
GGA1-mediated endocytic traffic of LR11/SorLA alters APP int...
ACCEPT
Summary: SorLA traffics through recycling endosomes, enabling cargo such as INSR and HER2 to be returned to the cell surface.
Reason: Recycling-endosome residence supports SorLA-mediated receptor recycling (IMP).
GO:1902430 negative regulation of amyloid-beta formation
IDA
PMID:22621900
GGA1-mediated endocytic traffic of LR11/SorLA alters APP int...
ACCEPT
Summary: SorLA lowers Abeta40/Abeta42 generation by sequestering APP from amyloidogenic processing and routing Abeta to lysosomes, the basis of its AD-protective role.
Reason: Central disease-relevant output directly downstream of SorLA APP sorting (IDA/IMP).
GO:1902430 negative regulation of amyloid-beta formation
IMP
PMID:24001769
SORLA-dependent and -independent functions for PACS1 in cont...
ACCEPT
Summary: SorLA lowers Abeta40/Abeta42 generation by sequestering APP from amyloidogenic processing and routing Abeta to lysosomes, the basis of its AD-protective role.
Reason: Central disease-relevant output directly downstream of SorLA APP sorting (IDA/IMP).
GO:1902430 negative regulation of amyloid-beta formation
IMP
PMID:24523320
Lysosomal sorting of amyloid-Ξ² by the SORLA receptor is impa...
ACCEPT
Summary: SorLA lowers Abeta40/Abeta42 generation by sequestering APP from amyloidogenic processing and routing Abeta to lysosomes, the basis of its AD-protective role.
Reason: Central disease-relevant output directly downstream of SorLA APP sorting (IDA/IMP).
GO:2001137 positive regulation of endocytic recycling
IMP
PMID:22621900
GGA1-mediated endocytic traffic of LR11/SorLA alters APP int...
ACCEPT
Summary: SorLA enhances recycling of internalized cargo back to the surface, exemplified by HER2 recycling that sustains PI3K-dependent signaling.
Reason: Core recycling-promoting activity of the sorting receptor (IMP).
GO:0001540 amyloid-beta binding
IPI
PMID:16407538
Interaction of the cytosolic domains of sorLA/LR11 with the ...
ACCEPT
Summary: SorLA directly binds amyloid-beta (and APP C99/Abeta40) via its ectodomain, enabling capture of newly produced Abeta for lysosomal clearance.
Reason: Defining ligand interaction underlying SorLA's anti-amyloidogenic sorting (IDA/IPI).
GO:0005515 protein binding
IPI
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005641 nuclear envelope lumen
IDA
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
UNDECIDED
Summary: RIP releases a SorLA C-terminal fragment that localizes to the nucleus, but a specific nuclear-envelope-lumen assignment is not clearly supported by the evidence reviewed.
Reason: Cannot verify the nuclear-envelope-lumen call; nuclear targeting is only of a cleaved fragment.
GO:0005641 nuclear envelope lumen
IDA
PMID:21989385
Quantitative modelling of amyloidogenic processing and its i...
UNDECIDED
Summary: RIP releases a SorLA C-terminal fragment that localizes to the nucleus, but a specific nuclear-envelope-lumen assignment is not clearly supported by the evidence reviewed.
Reason: Cannot verify the nuclear-envelope-lumen call; nuclear targeting is only of a cleaved fragment.
GO:0005783 endoplasmic reticulum
IDA
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
KEEP AS NON CORE
Summary: SorLA transits the ER during biosynthesis and maturation, but its sorting activity acts downstream in the TGN/endosomal system rather than in the ER.
Reason: Biosynthetic transit compartment, upstream of and peripheral to the core sorting cycle.
GO:0006605 protein targeting
IDA
PMID:16407538
Interaction of the cytosolic domains of sorLA/LR11 with the ...
ACCEPT
Summary: SorLA directs bound client proteins to specific destinations (TGN retention, lysosomal degradation or surface recycling), the essence of its sorting role.
Reason: Core process: directing proteins to their correct intracellular location.
GO:0006892 post-Golgi vesicle-mediated transport
IDA
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: SorLA participates in vesicular transport from the TGN onward, dispatching itself and cargo to endosomes and the cell surface.
Reason: Core trafficking step in SorLA's sorting itinerary (IBA/IDA).
GO:0031267 small GTPase binding
IPI
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
KEEP AS NON CORE
Summary: Reported binding to a small GTPase (e.g. RHOH) is a specific partner interaction not central to SorLA's cargo-sorting mechanism.
Reason: Specific partner binding, peripheral to the core sorting function (IPI).
GO:0031985 Golgi cisterna
IDA
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: IDA localization to Golgi cisternae is consistent with SorLA's residence in the Golgi stack during cargo sorting and retrieval.
Reason: Cisternal localization refines the core Golgi pool of this sorting receptor.
GO:0034067 protein localization to Golgi apparatus
IDA
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: By capturing and retrieving cargo to the Golgi/TGN, SorLA establishes the Golgi localization of clients such as APP.
Reason: Direct consequence of SorLA's retrieval/retention activity (IDA).
GO:0045053 protein retention in Golgi apparatus
IDA
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: SorLA retains APP (and other cargo) in the TGN, the key event that limits APP's transit to amyloidogenic late-endosomal processing.
Reason: Defining sorting outcome that links SorLA to reduced amyloidogenesis (IBA/IDA).
GO:1902430 negative regulation of amyloid-beta formation
IDA
PMID:16407538
Interaction of the cytosolic domains of sorLA/LR11 with the ...
ACCEPT
Summary: SorLA lowers Abeta40/Abeta42 generation by sequestering APP from amyloidogenic processing and routing Abeta to lysosomes, the basis of its AD-protective role.
Reason: Central disease-relevant output directly downstream of SorLA APP sorting (IDA/IMP).
GO:1902430 negative regulation of amyloid-beta formation
IMP
PMID:17855360
SorLA/LR11 regulates processing of amyloid precursor protein...
ACCEPT
Summary: SorLA lowers Abeta40/Abeta42 generation by sequestering APP from amyloidogenic processing and routing Abeta to lysosomes, the basis of its AD-protective role.
Reason: Central disease-relevant output directly downstream of SorLA APP sorting (IDA/IMP).
GO:1902430 negative regulation of amyloid-beta formation
IMP
PMID:21989385
Quantitative modelling of amyloidogenic processing and its i...
ACCEPT
Summary: SorLA lowers Abeta40/Abeta42 generation by sequestering APP from amyloidogenic processing and routing Abeta to lysosomes, the basis of its AD-protective role.
Reason: Central disease-relevant output directly downstream of SorLA APP sorting (IDA/IMP).
GO:0070062 extracellular exosome
HDA
PMID:19056867
Large-scale proteomics and phosphoproteomics of urinary exos...
KEEP AS NON CORE
Summary: HDA detection of SorLA in exosomes is consistent with proteomic surveys; it is incidental to the receptor's intracellular sorting function.
Reason: High-throughput proteomic localization; not a core functional site.
GO:0005515 protein binding
IPI
PMID:14764453
LR11, an LDL receptor gene family member, is a novel regulat...
MARK AS OVER ANNOTATED
Summary: Bare 'protein binding' (n=21 IPI) records many partner interactions but conveys no specific molecular function for this cargo receptor.
Reason: Uninformative generic MF; specific ligand-binding terms already capture SorLA's activity.
GO:0005576 extracellular region
IDA
PMID:14764453
LR11, an LDL receptor gene family member, is a novel regulat...
KEEP AS NON CORE
Summary: ADAM17-mediated shedding releases the soluble SorLA ectodomain (sLR11) into the extracellular space, where it acts as a secreted ligand-binding fragment.
Reason: Reflects shed soluble ectodomain, a secondary product of the membrane receptor.
GO:0014910 regulation of smooth muscle cell migration
IDA
PMID:14764453
LR11, an LDL receptor gene family member, is a novel regulat...
KEEP AS NON CORE
Summary: The shed SorLA ectodomain promotes smooth-muscle migration via increased uPAR surface display, a pleiotropic vascular role of the soluble fragment.
Reason: Vascular phenotype downstream of ectodomain/uPAR handling, not core sorting.
GO:0016020 membrane
IDA
PMID:14764453
LR11, an LDL receptor gene family member, is a novel regulat...
ACCEPT
Summary: Generic membrane association reflects SorLA's nature as an integral single-pass type I membrane protein distributed across the secretory/endosomal system.
Reason: Correct but generic; consistent with the transmembrane receptor.
GO:0005768 endosome
IDA
PMID:16174740
Neuronal sorting protein-related receptor sorLA/LR11 regulat...
ACCEPT
Summary: SorLA cycles through endosomes, capturing internalized cargo (APP, LPL, INSR, HER2) for retrieval to the TGN or routing to lysosomes.
Reason: Endosome is a defining compartment of SorLA's retrieval/sorting itinerary.
GO:0005794 Golgi apparatus
IDA
PMID:16174740
Neuronal sorting protein-related receptor sorLA/LR11 regulat...
ACCEPT
Summary: SorLA resides in the Golgi where, with AP-1, it executes TGN-endosome sorting; a major steady-state pool of the receptor is Golgi-associated.
Reason: Golgi is a core compartment for SorLA's cargo-sorting cycle (UniProt subcellular location).
GO:0006605 protein targeting
IMP
PMID:16174740
Neuronal sorting protein-related receptor sorLA/LR11 regulat...
ACCEPT
Summary: SorLA directs bound client proteins to specific destinations (TGN retention, lysosomal degradation or surface recycling), the essence of its sorting role.
Reason: Core process: directing proteins to their correct intracellular location.
GO:0030169 low-density lipoprotein particle binding
IPI
PMID:17326667
Apolipoprotein A-V interaction with members of the low densi...
ACCEPT
Summary: Through its LDLR-class complement repeats SorLA binds LDL particles, reflecting its membership in the LDL-receptor gene family.
Reason: Direct ligand-binding activity characteristic of this LDLR-family receptor (IPI).
GO:0005886 plasma membrane
TAS
PMID:9157966
A novel mosaic protein containing LDL receptor elements is h...
ACCEPT
Summary: A minor surface pool of SorLA captures extracellular ligands and undergoes AP-2-dependent endocytosis; surface levels are low but functionally important.
Reason: Cell-surface localization is the entry point for SorLA's endocytic sorting cycle.
GO:0006898 receptor-mediated endocytosis
TAS
PMID:9157966
A novel mosaic protein containing LDL receptor elements is h...
ACCEPT
Summary: SorLA mediates AP-2/clathrin-dependent internalization of surface ligands (uPA-PAI1, APOA5, IL6, HER2), feeding them into its endosomal sorting itinerary.
Reason: Core uptake step of SorLA's cargo-sorting cycle (IDA/TAS).

Core Functions

SORL1/SORLA is a type I membrane cargo receptor that binds APP and other ligands and routes them through trans-Golgi, endosomal, recycling, lysosomal, and plasma-membrane pathways via adaptor-dependent sorting.

Supporting Evidence:
  • PMID:16174740
    sorLA acts as a sorting receptor that protects APP from processing into Abeta and thereby reduces the burden of amyloidogenic peptide formation
  • PMID:17855360
    sorLA acts as a retention factor for APP in trans-Golgi compartments/trans-Golgi network, preventing release of the precursor into regular processing pathways
  • PMID:16407538
    sorLA acts as a trafficking receptor that prevents BACE-APP interactions and hence BACE cleavage of APP

SORL1 binds amyloid-beta peptides through its VPS10P domain and can route nascent amyloid-beta toward lysosomal catabolism, complementing its APP-retention role in lowering amyloidogenic peptide production.

Supporting Evidence:
  • PMID:24523320
    this receptor to direct lysosomal targeting of nascent AΞ² peptides
  • PMID:24523320
    AΞ² binds to the amino-terminal VPS10P domain of SORLA
  • PMID:22621900
    LR11 modulates AΞ² levels by promoting APP traffic to the endocytic recycling compartment

References

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Suggested Questions for Experts

Q: Should GO add a more specific APP/amyloid-beta cargo sorting receptor activity term for SORL1-like receptor functions, or is cargo receptor activity plus cargo-specific binding/process annotations sufficient?

Suggested experts: GO receptor activity curators, Alzheimer trafficking experts

Q: Which SORL1 cargo-specific phenotypes, such as insulin receptor recycling, GDNF/GFRA1 turnover, IL6-family cytokine regulation, or HER2 recycling, should be retained as non-core versus represented only through generic cargo sorting?

Suggested experts: VPS10P-domain receptor experts, GO annotation curators

Q: Are nuclear-envelope or perinucleolar localization annotations for SORL1 supported by direct full-text evidence, or are they artifacts of perinuclear Golgi/endosomal localization descriptions?

Suggested experts: cellular trafficking curators, SORL1 cell biology experts

Suggested Experiments

Experiment: Use endogenous SORL1 knock-in/knockout neurons to compare APP, BACE1, and nascent amyloid-beta routing across TGN, early endosome, recycling endosome, and lysosomal compartments.

Hypothesis: SORL1 lowers amyloidogenic processing primarily by retaining or retrieving APP/BACE1-containing cargo away from compartments that favor BACE cleavage and by routing nascent amyloid-beta to lysosomes.

Type: live-cell cargo trafficking and compartment-resolved APP processing assay

Experiment: Introduce Alzheimer-associated SORL1 variants into the endogenous locus and assay cargo binding, adaptor recruitment, endosome-to-Golgi retrieval, and A-beta lysosomal targeting.

Hypothesis: Pathogenic SORL1 variants impair specific cargo-binding or adaptor-dependent trafficking steps without eliminating the general sorting-receptor scaffold.

Type: endogenous variant knock-in functional rescue assay

Experiment: Compare APP/A-beta, insulin receptor, GDNF/GFRA1, IL6-family cytokine complexes, and HER2 as parallel cargoes in matched cellular systems.

Hypothesis: SORL1 uses a shared trafficking architecture for many cargoes, but Alzheimer-relevant APP/A-beta handling depends on cargo-specific binding and adaptor requirements.

Type: multi-cargo receptor sorting assay

πŸ“š Additional Documentation

Notes

(SORL1-notes.md)

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