SPI1

UniProt ID: P17947
Organism: Homo sapiens
Review Status: COMPLETE
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Gene Description

SPI1 encodes PU.1, an ETS-family pioneer transcription factor expressed mainly in hematopoietic and immune lineages. PU.1 localizes to nucleus, nucleoplasm, chromatin, and transcription-regulator complexes, where its ETS domain binds purine-rich PU-box/cis-regulatory DNA motifs and works with other transcription factors and chromatin cofactors to activate or repress RNA polymerase II transcription. By opening or selecting regulatory chromatin and coordinating lineage-specific transcriptional programs, PU.1 controls myeloid, macrophage/microglial, dendritic-cell, B-cell, and broader hematopoietic differentiation and immune-cell functional states.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0005634 nucleus
IBA
GO_REF:0000033
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0030154 cell differentiation
IBA
GO_REF:0000033
ACCEPT
Summary: PU.1 regulates hematopoietic lineage fate, especially myeloid/macrophage, dendritic-cell, and B-cell development.
Reason: Retain as core or near-core developmental biology because PU.1 deficiency or mutation disrupts lymphoid and myeloid development, while iPSC macrophage differentiation is SPI1-dependent (PMID:28111278, PMID:33951726).
GO:0000981 DNA-binding transcription factor activity, RNA polymerase II-specific
IBA
GO_REF:0000033
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0006357 regulation of transcription by RNA polymerase II
IBA
GO_REF:0000033
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0003700 DNA-binding transcription factor activity
IEA
GO_REF:0000120
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0005634 nucleus
IEA
GO_REF:0000120
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0006355 regulation of DNA-templated transcription
IEA
GO_REF:0000002
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0006357 regulation of transcription by RNA polymerase II
IEA
GO_REF:0000002
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0043565 sequence-specific DNA binding
IEA
GO_REF:0000120
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0005515 protein binding
IPI
PMID:18250304
Gain-of-function mutation of GATA-2 in acute myeloid transfo...
MODIFY
Summary: The cited interaction reflects PU.1 cooperation with transcription factors or transcriptional cofactors rather than generic protein binding.
Reason: Modify generic protein binding to transcription-factor/cofactor binding where the evidence supports PU.1 physical or functional interaction with another transcriptional regulator.
GO:0005515 protein binding
IPI
PMID:18692240
Physical and functional interactions between hematopoietic c...
MODIFY
Summary: The cited interaction reflects PU.1 cooperation with transcription factors or transcriptional cofactors rather than generic protein binding.
Reason: Modify generic protein binding to transcription-factor/cofactor binding where the evidence supports PU.1 physical or functional interaction with another transcriptional regulator.
GO:0005515 protein binding
IPI
PMID:20159610
PML/RARalpha targets promoter regions containing PU.1 consen...
MODIFY
Summary: The cited interaction reflects PU.1 cooperation with transcription factors or transcriptional cofactors rather than generic protein binding.
Reason: Modify generic protein binding to transcription-factor/cofactor binding where the evidence supports PU.1 physical or functional interaction with another transcriptional regulator.
GO:0005515 protein binding
IPI
PMID:21575865
Mechanistic rationale for inhibition of poly(ADP-ribose) pol...
UNDECIDED
Summary: The cached abstract is abstract-only or focused on another ETS/fusion context and does not expose enough evidence to verify the SPI1-specific interaction.
Reason: Use UNDECIDED rather than removing the experimental annotation because the curator may have used full-text evidence not present in the cache; the abstract alone does not establish this SPI1-specific generic protein-binding assertion.
GO:0000122 negative regulation of transcription by RNA polymerase II
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000785 chromatin
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000978 RNA polymerase II cis-regulatory region sequence-specific DNA binding
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000981 DNA-binding transcription factor activity, RNA polymerase II-specific
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000987 cis-regulatory region sequence-specific DNA binding
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0001227 DNA-binding transcription repressor activity, RNA polymerase II-specific
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0001228 DNA-binding transcription activator activity, RNA polymerase II-specific
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0002314 germinal center B cell differentiation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0002316 follicular B cell differentiation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0002327 immature B cell differentiation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0002572 pro-T cell differentiation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0003677 DNA binding
IEA
GO_REF:0000107
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0003682 chromatin binding
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1 acts in nucleus/chromatin as a pioneer/master transcription factor that selects accessible and inaccessible binding sites and shapes hematopoietic chromatin accessibility.
Reason: Retain as core because PU.1 occupies active chromatin domains, can bind high-affinity sites in DNase-inaccessible regions, and controls hematopoietic euchromatin accessibility and transcription-factor access (PMID:23658224, PMID:33951726).
GO:0007179 transforming growth factor beta receptor signaling pathway
IEA
GO_REF:0000107
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:0010628 positive regulation of gene expression
IEA
GO_REF:0000107
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0010629 negative regulation of gene expression
IEA
GO_REF:0000107
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0030316 osteoclast differentiation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0031663 lipopolysaccharide-mediated signaling pathway
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0042826 histone deacetylase binding
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0043314 negative regulation of neutrophil degranulation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0045347 negative regulation of MHC class II biosynthetic process
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0045471 response to ethanol
IEA
GO_REF:0000107
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:0045815 transcription initiation-coupled chromatin remodeling
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1 acts in nucleus/chromatin as a pioneer/master transcription factor that selects accessible and inaccessible binding sites and shapes hematopoietic chromatin accessibility.
Reason: Retain as core because PU.1 occupies active chromatin domains, can bind high-affinity sites in DNase-inaccessible regions, and controls hematopoietic euchromatin accessibility and transcription-factor access (PMID:23658224, PMID:33951726).
GO:0045892 negative regulation of DNA-templated transcription
IEA
GO_REF:0000120
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045893 positive regulation of DNA-templated transcription
IEA
GO_REF:0000107
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0045944 positive regulation of transcription by RNA polymerase II
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0051525 NFAT protein binding
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0061629 RNA polymerase II-specific DNA-binding transcription factor binding
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0071361 cellular response to ethanol
IEA
GO_REF:0000107
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:0090402 oncogene-induced cell senescence
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0098508 endothelial to hematopoietic transition
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0120186 negative regulation of protein localization to chromatin
IEA
GO_REF:0000107
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:0140297 DNA-binding transcription factor binding
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0140311 protein sequestering activity
IEA
GO_REF:0000107
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:1900745 positive regulation of p38MAPK cascade
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:1902895 positive regulation of miRNA transcription
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:1904151 positive regulation of microglial cell mediated cytotoxicity
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:1904178 negative regulation of adipose tissue development
IEA
GO_REF:0000107
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:1904238 pericyte cell differentiation
IEA
GO_REF:0000107
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:1905036 positive regulation of antifungal innate immune response
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:1905453 regulation of myeloid progenitor cell differentiation
IEA
GO_REF:0000107
ACCEPT
Summary: PU.1 regulates hematopoietic lineage fate, especially myeloid/macrophage, dendritic-cell, and B-cell development.
Reason: Retain as core or near-core developmental biology because PU.1 deficiency or mutation disrupts lymphoid and myeloid development, while iPSC macrophage differentiation is SPI1-dependent (PMID:28111278, PMID:33951726).
GO:2000529 positive regulation of myeloid dendritic cell chemotaxis
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0005654 nucleoplasm
IDA
GO_REF:0000052
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0043124 negative regulation of canonical NF-kappaB signal transduction
IDA
PMID:28362429
PU.1 supports TRAIL-induced cell death by inhibiting NF-ΞΊB-m...
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0045815 transcription initiation-coupled chromatin remodeling
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1 acts in nucleus/chromatin as a pioneer/master transcription factor that selects accessible and inaccessible binding sites and shapes hematopoietic chromatin accessibility.
Reason: Retain as core because PU.1 occupies active chromatin domains, can bind high-affinity sites in DNase-inaccessible regions, and controls hematopoietic euchromatin accessibility and transcription-factor access (PMID:23658224, PMID:33951726).
GO:0005515 protein binding
IPI
PMID:33951726
Constrained chromatin accessibility in PU.1-mutated agammagl...
MODIFY
Summary: The cited interaction reflects PU.1 cooperation with transcription factors or transcriptional cofactors rather than generic protein binding.
Reason: Modify generic protein binding to transcription-factor/cofactor binding where the evidence supports PU.1 physical or functional interaction with another transcriptional regulator.
GO:0005634 nucleus
IDA
PMID:33951726
Constrained chromatin accessibility in PU.1-mutated agammagl...
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0043565 sequence-specific DNA binding
IDA
PMID:33951726
Constrained chromatin accessibility in PU.1-mutated agammagl...
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0045579 positive regulation of B cell differentiation
IMP
PMID:33951726
Constrained chromatin accessibility in PU.1-mutated agammagl...
ACCEPT
Summary: PU.1 regulates hematopoietic lineage fate, especially myeloid/macrophage, dendritic-cell, and B-cell development.
Reason: Retain as core or near-core developmental biology because PU.1 deficiency or mutation disrupts lymphoid and myeloid development, while iPSC macrophage differentiation is SPI1-dependent (PMID:28111278, PMID:33951726).
GO:0045815 transcription initiation-coupled chromatin remodeling
IMP
PMID:33951726
Constrained chromatin accessibility in PU.1-mutated agammagl...
ACCEPT
Summary: PU.1 acts in nucleus/chromatin as a pioneer/master transcription factor that selects accessible and inaccessible binding sites and shapes hematopoietic chromatin accessibility.
Reason: Retain as core because PU.1 occupies active chromatin domains, can bind high-affinity sites in DNase-inaccessible regions, and controls hematopoietic euchromatin accessibility and transcription-factor access (PMID:23658224, PMID:33951726).
GO:0002357 defense response to tumor cell
IMP
PMID:28362429
PU.1 supports TRAIL-induced cell death by inhibiting NF-ΞΊB-m...
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0036462 TRAIL-activated apoptotic signaling pathway
IMP
PMID:28362429
PU.1 supports TRAIL-induced cell death by inhibiting NF-ΞΊB-m...
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0061629 RNA polymerase II-specific DNA-binding transcription factor binding
IPI
PMID:24429361
Hepatitis C virus-induced changes in microRNA 107 (miRNA-107...
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0097677 STAT family protein binding
IPI
PMID:24429361
Hepatitis C virus-induced changes in microRNA 107 (miRNA-107...
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0000987 cis-regulatory region sequence-specific DNA binding
IDA
PMID:28481873
Restoring PU.1 induces apoptosis and modulates viral transac...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0002357 defense response to tumor cell
IMP
PMID:28481873
Restoring PU.1 induces apoptosis and modulates viral transac...
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0045944 positive regulation of transcription by RNA polymerase II
IDA
PMID:28481873
Restoring PU.1 induces apoptosis and modulates viral transac...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000122 negative regulation of transcription by RNA polymerase II
IMP
PMID:28362429
PU.1 supports TRAIL-induced cell death by inhibiting NF-ΞΊB-m...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
IDA
PMID:12833137
CDK6 blocks differentiation: coupling cell proliferation to ...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
IDA
PMID:20139074
Metallothionein-1 isoforms and vimentin are direct PU.1 down...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
IDA
PMID:27506447
Transcriptional regulation of the proto-oncogene Zfp521 by S...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
IGI
PMID:27506447
Transcriptional regulation of the proto-oncogene Zfp521 by S...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
IMP
PMID:28362429
PU.1 supports TRAIL-induced cell death by inhibiting NF-ΞΊB-m...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
IMP
PMID:30718772
A transcription factor PU.1 is critical for Ccl22 gene expre...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
IMP
PMID:31146003
Androgen upregulates NR4A1 via the TFAP2A and ETS signaling ...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
IMP
PMID:31314592
PU.1 plays a pivotal role in dendritic cell migration from t...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:1902895 positive regulation of miRNA transcription
IDA
PMID:20972335
Hypoxia-induced microRNA-424 expression in human endothelial...
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0001216 DNA-binding transcription activator activity
IDA
PMID:20139074
Metallothionein-1 isoforms and vimentin are direct PU.1 down...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0001217 DNA-binding transcription repressor activity
IDA
PMID:20139074
Metallothionein-1 isoforms and vimentin are direct PU.1 down...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0098508 endothelial to hematopoietic transition
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:1905453 regulation of myeloid progenitor cell differentiation
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1 regulates hematopoietic lineage fate, especially myeloid/macrophage, dendritic-cell, and B-cell development.
Reason: Retain as core or near-core developmental biology because PU.1 deficiency or mutation disrupts lymphoid and myeloid development, while iPSC macrophage differentiation is SPI1-dependent (PMID:28111278, PMID:33951726).
GO:0061629 RNA polymerase II-specific DNA-binding transcription factor binding
IPI
PMID:28362429
PU.1 supports TRAIL-induced cell death by inhibiting NF-ΞΊB-m...
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:1901223 negative regulation of non-canonical NF-kappaB signal transduction
IMP
PMID:28362429
PU.1 supports TRAIL-induced cell death by inhibiting NF-ΞΊB-m...
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:1904178 negative regulation of adipose tissue development
ISS
GO_REF:0000024
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:0000987 cis-regulatory region sequence-specific DNA binding
IDA
PMID:30718772
A transcription factor PU.1 is critical for Ccl22 gene expre...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000987 cis-regulatory region sequence-specific DNA binding
IDA
PMID:31314592
PU.1 plays a pivotal role in dendritic cell migration from t...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:1904238 pericyte cell differentiation
ISS
GO_REF:0000024
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:0000976 transcription cis-regulatory region binding
IDA
PMID:31146003
Androgen upregulates NR4A1 via the TFAP2A and ETS signaling ...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000976 transcription cis-regulatory region binding
IMP
PMID:27506447
Transcriptional regulation of the proto-oncogene Zfp521 by S...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0003700 DNA-binding transcription factor activity
IDA
PMID:27506447
Transcriptional regulation of the proto-oncogene Zfp521 by S...
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0140297 DNA-binding transcription factor binding
IPI
PMID:27506447
Transcriptional regulation of the proto-oncogene Zfp521 by S...
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0002357 defense response to tumor cell
IMP
PMID:30429596
In vitro conversion of adult murine endothelial cells to hem...
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0005634 nucleus
IDA
PMID:24504023
Dual regulation of SPI1/PU.1 transcription factor by heat sh...
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000987 cis-regulatory region sequence-specific DNA binding
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0002572 pro-T cell differentiation
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0003682 chromatin binding
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1 acts in nucleus/chromatin as a pioneer/master transcription factor that selects accessible and inaccessible binding sites and shapes hematopoietic chromatin accessibility.
Reason: Retain as core because PU.1 occupies active chromatin domains, can bind high-affinity sites in DNase-inaccessible regions, and controls hematopoietic euchromatin accessibility and transcription-factor access (PMID:23658224, PMID:33951726).
GO:0003700 DNA-binding transcription factor activity
ISS
GO_REF:0000024
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0002573 myeloid leukocyte differentiation
IMP
PMID:28111278
Human Induced Pluripotent Stem Cell-Derived Macrophages Shar...
ACCEPT
Summary: PU.1 regulates hematopoietic lineage fate, especially myeloid/macrophage, dendritic-cell, and B-cell development.
Reason: Retain as core or near-core developmental biology because PU.1 deficiency or mutation disrupts lymphoid and myeloid development, while iPSC macrophage differentiation is SPI1-dependent (PMID:28111278, PMID:33951726).
GO:1904151 positive regulation of microglial cell mediated cytotoxicity
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0000987 cis-regulatory region sequence-specific DNA binding
IDA
PMID:31018969
A Recurrent Activating Missense Mutation in WaldenstrΓΆm Macr...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0003700 DNA-binding transcription factor activity
IDA
PMID:31018969
A Recurrent Activating Missense Mutation in WaldenstrΓΆm Macr...
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0006355 regulation of DNA-templated transcription
IDA
PMID:31018969
A Recurrent Activating Missense Mutation in WaldenstrΓΆm Macr...
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0045892 negative regulation of DNA-templated transcription
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0140297 DNA-binding transcription factor binding
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0140311 protein sequestering activity
ISS
GO_REF:0000024
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:0000122 negative regulation of transcription by RNA polymerase II
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0042826 histone deacetylase binding
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0043314 negative regulation of neutrophil degranulation
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0120186 negative regulation of protein localization to chromatin
ISS
GO_REF:0000024
MARK AS OVER ANNOTATED
Summary: This annotation is a narrow disease, cell-line, high-throughput, or non-core downstream context and does not define SPI1/PU.1 core function.
Reason: Mark as over-annotated because the reviewed evidence supports PU.1 as a hematopoietic ETS transcription factor, while this term is peripheral, context-specific, or too downstream for core function curation.
GO:1905036 positive regulation of antifungal innate immune response
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0010629 negative regulation of gene expression
IMP
PMID:29290617
METTL14 Inhibits Hematopoietic Stem/Progenitor Differentiati...
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0090402 oncogene-induced cell senescence
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:1900745 positive regulation of p38MAPK cascade
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0000785 chromatin
ISA
GO_REF:0000113
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000981 DNA-binding transcription factor activity, RNA polymerase II-specific
ISA
GO_REF:0000113
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0005667 transcription regulator complex
IDA
PMID:24429361
Hepatitis C virus-induced changes in microRNA 107 (miRNA-107...
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045944 positive regulation of transcription by RNA polymerase II
IGI
PMID:24429361
Hepatitis C virus-induced changes in microRNA 107 (miRNA-107...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0070102 interleukin-6-mediated signaling pathway
IDA
PMID:24429361
Hepatitis C virus-induced changes in microRNA 107 (miRNA-107...
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0000978 RNA polymerase II cis-regulatory region sequence-specific DNA binding
IDA
PMID:20139074
Metallothionein-1 isoforms and vimentin are direct PU.1 down...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0005634 nucleus
IDA
PMID:20972335
Hypoxia-induced microRNA-424 expression in human endothelial...
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0005515 protein binding
IPI
PMID:26019275
A Novel In-Frame Deletion in the Leucine Zipper Domain of C/...
MODIFY
Summary: The cited interaction reflects PU.1 cooperation with transcription factors or transcriptional cofactors rather than generic protein binding.
Reason: Modify generic protein binding to transcription-factor/cofactor binding where the evidence supports PU.1 physical or functional interaction with another transcriptional regulator.
GO:0005654 nucleoplasm
TAS
Reactome:R-HSA-9617064
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0005654 nucleoplasm
TAS
Reactome:R-HSA-9617207
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:1902895 positive regulation of miRNA transcription
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0000978 RNA polymerase II cis-regulatory region sequence-specific DNA binding
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0000981 DNA-binding transcription factor activity, RNA polymerase II-specific
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0051525 NFAT protein binding
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0061629 RNA polymerase II-specific DNA-binding transcription factor binding
ISS
GO_REF:0000024
ACCEPT
Summary: PU.1 physically or functionally cooperates with transcription factors and chromatin/transcriptional cofactors.
Reason: Retain because PU.1 function includes cooperative binding with other transcriptional regulators and cofactors at hematopoietic regulatory elements (PMID:23658224, PMID:33951726).
GO:0000785 chromatin
IDA
PMID:15304486
Essential role of p38 mitogen-activated protein kinase in ca...
ACCEPT
Summary: PU.1 is a nuclear/chromatin transcription factor.
Reason: Retain because PU.1 functions in the nucleus, nucleoplasm, chromatin, and transcription-regulator complexes to bind regulatory DNA and control gene expression (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0045892 negative regulation of DNA-templated transcription
IDA
PMID:20139074
Metallothionein-1 isoforms and vimentin are direct PU.1 down...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0003700 DNA-binding transcription factor activity
IDA
PMID:12833137
CDK6 blocks differentiation: coupling cell proliferation to ...
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0045646 regulation of erythrocyte differentiation
IMP
PMID:12833137
CDK6 blocks differentiation: coupling cell proliferation to ...
KEEP AS NON CORE
Summary: This lineage or immune-cell phenotype is compatible with PU.1 biology but is downstream or cell-type-specific.
Reason: Keep as non-core because it reflects a particular hematopoietic, immune, inflammatory, or disease-context outcome of PU.1 transcriptional control rather than the defining DNA-binding/chromatin-regulatory molecular function.
GO:0005515 protein binding
IPI
PMID:10207087
Functional and physical interactions between AML1 proteins a...
UNDECIDED
Summary: The cached abstract is abstract-only or focused on another ETS/fusion context and does not expose enough evidence to verify the SPI1-specific interaction.
Reason: Use UNDECIDED rather than removing the experimental annotation because the curator may have used full-text evidence not present in the cache; the abstract alone does not establish this SPI1-specific generic protein-binding assertion.
GO:0000122 negative regulation of transcription by RNA polymerase II
TAS
PMID:10867017
The hematopoietic transcription factor PU.1 represses gelati...
ACCEPT
Summary: PU.1/SPI1 is an ETS-domain transcription factor that binds PU-box/cis-regulatory DNA motifs and activates or represses RNA polymerase II transcription.
Reason: Retain as core because PU.1 recognizes the 5'-GAGGAA-3' PU-box/ETS motif, binds cis-regulatory regions in vivo, and controls hematopoietic gene expression as a DNA-binding transcription factor (PMID:2180582, PMID:23658224, PMID:33951726).
GO:0003700 DNA-binding transcription factor activity
TAS
PMID:10867017
The hematopoietic transcription factor PU.1 represses gelati...
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.
GO:0003700 DNA-binding transcription factor activity
TAS
PMID:2180582
The macrophage and B cell-specific transcription factor PU.1...
MODIFY
Summary: The annotation is directionally correct but too broad for PU.1; the specific biology is RNA polymerase II cis-regulatory DNA binding and transcription factor activity.
Reason: Modify to a more informative PU.1 transcription-factor term because generic DNA binding or DNA-templated transcription regulation obscures the ETS-domain cis-regulatory/RNA polymerase II mechanism.

Core Functions

ETS-domain cis-regulatory DNA binding and RNA polymerase II transcription-factor activity at PU-box/ETS motifs in hematopoietic regulatory elements.

Supporting Evidence:
  • PMID:2180582
    the PU.1 protein recognized a purine-rich sequence, 5'-GAGGAA-3' (PU box)
  • PMID:2180582
    The PU.1 protein was shown to be a transcriptional activator that is expressed in macrophages and B cells
  • PMID:23658224
    PU.1 selects its binding sites primarily based on sequence affinity

Pioneer/chromatin-regulatory activity that opens or selects hematopoietic regulatory chromatin and permits cooperative transcription-factor access.

Supporting Evidence:
  • PMID:23658224
    Occupied sites were predominantly detected in active chromatin domains
  • PMID:33951726
    decompacting stem cell heterochromatin and allowing nonpioneer TFs to enter
  • PMID:33951726
    destabilized PU.1 proteins unable to nuclear localize or bind target DNA

Hematopoietic lineage specification and differentiation, especially B-cell, myeloid/macrophage, dendritic-cell, and microglia-related macrophage programs, through PU.1-dependent transcriptional control.

Supporting Evidence:
  • PMID:33951726
    Affected patients lacked circulating B cells and possessed few conventional dendritic cells
  • PMID:33951726
    mutations into human hematopoietic stem and progenitor cells impaired early in vitro B cell and myeloid cell differentiation
  • PMID:28111278
    human iPSC-derived monocytes/macrophages develop in an MYB-independent, RUNX1-, and SPI1 (PU.1)-dependent fashion

References

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Suggested Questions for Experts

Q: Which SPI1-dependent microglial terms should be curated as direct lineage-maintenance functions rather than downstream immune-cell outcomes?

Q: Can full-text evidence for the abstract-only ETS-family interaction annotations be reviewed to decide whether SPI1-specific protein-binding assertions should be accepted or removed?

Suggested Experiments

Experiment: Perform matched PU.1 ChIP-seq/CUT&RUN and ATAC-seq in primary or iPSC-derived human microglia carrying relevant SPI1 regulatory haplotypes.

Hypothesis: Alzheimer-associated SPI1 regulatory variation alters PU.1 occupancy and chromatin accessibility in microglia.

Type: microglial chromatin profiling

Experiment: Compare SPI1 titration or degron perturbation across human B-cell precursors and microglia-like cells with RNA-seq and chromatin accessibility readouts.

Hypothesis: PU.1 dosage controls distinct B-cell and microglial target-gene programs through shared ETS motif occupancy but different cooperative factors.

Type: dose-response perturbation genomics

πŸ“š Additional Documentation

Notes

(SPI1-notes.md)

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πŸ“„ View Raw YAML

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