TREM2

UniProt ID: Q9NZC2
Organism: Homo sapiens
Review Status: COMPLETE
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Gene Description

TREM2 is a type I plasma-membrane immunoglobulin-superfamily receptor expressed mainly by microglia and other myeloid-lineage cells. Together with the TYROBP/DAP12 adaptor, TREM2 converts ligand binding at the cell surface into intracellular signaling through kinases and PLCG2-linked pathways that support cell survival, calcium/ERK signaling, phagocytosis, chemotaxis, microglial activation, and lipid handling. TREM2 recognizes lipid-rich and damage-associated ligands, including aminophospholipids exposed on apoptotic cells, lipoprotein particles and apolipoproteins, and amyloid-beta-lipoprotein complexes. Disease-model annotations involving amyloid plaques, synapse pruning, neuroinflammation, osteoclast biology, and dendritic cell maturation are important contexts but are secondary to the core receptor-ligand signaling and phagocytic myeloid-cell functions.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0005886 plasma membrane
IBA
GO_REF:0000033
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0004888 transmembrane signaling receptor activity
IBA
GO_REF:0000033
ACCEPT
Summary: TREM2 transmembrane signaling receptor activity is a core plasma-membrane receptor function coupled to TYROBP/DAP12 signaling.
Reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives intracellular signaling through DAP12/TYROBP-linked kinase pathways.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0007165 signal transduction
IBA
GO_REF:0000033
ACCEPT
Summary: signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0045088 regulation of innate immune response
IBA
GO_REF:0000033
ACCEPT
Summary: regulation of innate immune response is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0050850 positive regulation of calcium-mediated signaling
IBA
GO_REF:0000033
ACCEPT
Summary: positive regulation of calcium-mediated signaling is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0060100 positive regulation of phagocytosis, engulfment
IBA
GO_REF:0000033
ACCEPT
Summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0070374 positive regulation of ERK1 and ERK2 cascade
IBA
GO_REF:0000033
ACCEPT
Summary: positive regulation of ERK1 and ERK2 cascade is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0120035 regulation of plasma membrane bounded cell projection organization
IBA
GO_REF:0000033
KEEP AS NON CORE
Summary: regulation of plasma membrane bounded cell projection organization is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1903980 positive regulation of microglial cell activation
IBA
GO_REF:0000033
ACCEPT
Summary: positive regulation of microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0004888 transmembrane signaling receptor activity
IEA
GO_REF:0000117
ACCEPT
Summary: TREM2 transmembrane signaling receptor activity is a core plasma-membrane receptor function coupled to TYROBP/DAP12 signaling.
Reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives intracellular signaling through DAP12/TYROBP-linked kinase pathways.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0005576 extracellular region
IEA
GO_REF:0000044
KEEP AS NON CORE
Summary: The extracellular-region annotation likely reflects secreted or shed TREM2 isoforms/fragments rather than the core membrane receptor.
Reason: Retain as non-core because soluble TREM2 biology is relevant, but the primary TREM2 function is cell-surface receptor signaling.
GO:0005886 plasma membrane
IEA
GO_REF:0000120
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0006909 phagocytosis
IEA
GO_REF:0000117
ACCEPT
Summary: phagocytosis is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0008289 lipid binding
IEA
GO_REF:0000117
ACCEPT
Summary: TREM2 lipid binding is part of its ligand-sensing receptor biology.
Reason: TREM2 ligand recognition includes lipid-rich particles and supports microglial lipid handling.
Supporting Evidence:
PMID:27477018
we identified a set of lipoprotein particles (including LDL) and apolipoproteins (including CLU/APOJ and APOE) as ligands
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
GO:0035176 social behavior
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: social behavior is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0048468 cell development
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: cell development is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0050850 positive regulation of calcium-mediated signaling
IEA
GO_REF:0000117
ACCEPT
Summary: positive regulation of calcium-mediated signaling is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0071396 cellular response to lipid
IEA
GO_REF:0000117
ACCEPT
Summary: cellular response to lipid is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0097367 carbohydrate derivative binding
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: carbohydrate derivative binding is retained as non-core TREM2-associated biology pending deeper reference-specific adjudication.
Reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic microglial function set, but the local evidence was not sufficient to remove it.
GO:1900226 negative regulation of NLRP3 inflammasome complex assembly
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: negative regulation of NLRP3 inflammasome complex assembly is retained as non-core TREM2-associated biology pending deeper reference-specific adjudication.
Reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic microglial function set, but the local evidence was not sufficient to remove it.
GO:0005515 protein binding
IPI
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
MARK AS OVER ANNOTATED
Summary: Generic protein binding annotation for TREM2.
Reason: The interaction may be real, but this term is too generic. Specific ligand-binding, receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
GO:0005515 protein binding
IPI
PMID:28490631
A split-luciferase complementation, real-time reporting assa...
MARK AS OVER ANNOTATED
Summary: Generic protein binding annotation for TREM2.
Reason: The interaction may be real, but this term is too generic. Specific ligand-binding, receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
GO:0005515 protein binding
IPI
PMID:29611543
Intracellular trafficking of TREM2 is regulated by presenili...
MARK AS OVER ANNOTATED
Summary: Generic protein binding annotation for TREM2.
Reason: The interaction may be real, but this term is too generic. Specific ligand-binding, receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
GO:0005515 protein binding
IPI
PMID:30341064
High-affinity interactions and signal transduction between A...
MARK AS OVER ANNOTATED
Summary: Generic protein binding annotation for TREM2.
Reason: The interaction may be real, but this term is too generic. Specific ligand-binding, receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
GO:0001530 lipopolysaccharide binding
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: lipopolysaccharide binding is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0001774 microglial cell activation
IEA
GO_REF:0000107
ACCEPT
Summary: microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0002282 microglial cell activation involved in immune response
IEA
GO_REF:0000107
ACCEPT
Summary: microglial cell activation involved in immune response is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0002291 T cell activation via T cell receptor contact with antigen bound to MHC molecule on antigen presenting cell
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: T cell activation via T cell receptor contact with antigen bound to MHC molecule on antigen presenting cell is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0002862 negative regulation of inflammatory response to antigenic stimulus
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of inflammatory response to antigenic stimulus is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0002931 response to ischemia
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: response to ischemia is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0010507 negative regulation of autophagy
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of autophagy is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0010628 positive regulation of gene expression
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: positive regulation of gene expression is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0010875 positive regulation of cholesterol efflux
IEA
GO_REF:0000107
ACCEPT
Summary: positive regulation of cholesterol efflux is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0010887 negative regulation of cholesterol storage
IEA
GO_REF:0000107
ACCEPT
Summary: negative regulation of cholesterol storage is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0010891 negative regulation of triglyceride storage
IEA
GO_REF:0000107
ACCEPT
Summary: negative regulation of triglyceride storage is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0010983 positive regulation of high-density lipoprotein particle clearance
IEA
GO_REF:0000107
ACCEPT
Summary: positive regulation of high-density lipoprotein particle clearance is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0017154 semaphorin receptor activity
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: semaphorin receptor activity is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0030215 semaphorin receptor binding
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: semaphorin receptor binding is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0032006 regulation of TOR signaling
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: regulation of TOR signaling is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0032008 positive regulation of TOR signaling
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of TOR signaling is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0032497 detection of lipopolysaccharide
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: detection of lipopolysaccharide is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0032499 detection of peptidoglycan
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: detection of peptidoglycan is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0032675 regulation of interleukin-6 production
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: regulation of interleukin-6 production is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0032691 negative regulation of interleukin-1 beta production
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of interleukin-1 beta production is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0032720 negative regulation of tumor necrosis factor production
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of tumor necrosis factor production is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0032733 positive regulation of interleukin-10 production
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of interleukin-10 production is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0034136 negative regulation of toll-like receptor 2 signaling pathway
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of toll-like receptor 2 signaling pathway is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0034144 negative regulation of toll-like receptor 4 signaling pathway
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of toll-like receptor 4 signaling pathway is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0034151 regulation of toll-like receptor 6 signaling pathway
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: regulation of toll-like receptor 6 signaling pathway is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0034241 positive regulation of macrophage fusion
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of macrophage fusion is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0034351 negative regulation of glial cell apoptotic process
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of glial cell apoptotic process is retained as non-core TREM2-associated biology pending deeper reference-specific adjudication.
Reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic microglial function set, but the local evidence was not sufficient to remove it.
GO:0038023 signaling receptor activity
IEA
GO_REF:0000107
ACCEPT
Summary: TREM2 signaling receptor activity is a core plasma-membrane receptor function coupled to TYROBP/DAP12 signaling.
Reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives intracellular signaling through DAP12/TYROBP-linked kinase pathways.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0042742 defense response to bacterium
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: defense response to bacterium is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0042834 peptidoglycan binding
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: peptidoglycan binding is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0043066 negative regulation of apoptotic process
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of apoptotic process is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0043124 negative regulation of canonical NF-kappaB signal transduction
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of canonical NF-kappaB signal transduction is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0043268 positive regulation of potassium ion transport
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of potassium ion transport is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0043277 apoptotic cell clearance
IEA
GO_REF:0000107
ACCEPT
Summary: apoptotic cell clearance is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0045088 regulation of innate immune response
IEA
GO_REF:0000107
ACCEPT
Summary: regulation of innate immune response is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0045672 positive regulation of osteoclast differentiation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of osteoclast differentiation is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
Supporting Evidence:
PMID:12925681
loss of function mutations in DAP12 and TREM2 result in an inefficient and delayed differentiation of osteoclasts
GO:0045728 respiratory burst after phagocytosis
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: respiratory burst after phagocytosis is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0045960 positive regulation of complement activation, classical pathway
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of complement activation, classical pathway is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0048143 astrocyte activation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: astrocyte activation is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0048678 response to axon injury
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: response to axon injury is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0050766 positive regulation of phagocytosis
IEA
GO_REF:0000107
ACCEPT
Summary: positive regulation of phagocytosis is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0050866 negative regulation of cell activation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of cell activation is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0050921 positive regulation of chemotaxis
IEA
GO_REF:0000107
ACCEPT
Summary: positive regulation of chemotaxis is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0051897 positive regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
IEA
GO_REF:0000107
ACCEPT
Summary: positive regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0051898 negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
IEA
GO_REF:0000120
ACCEPT
Summary: negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0055088 lipid homeostasis
IEA
GO_REF:0000107
ACCEPT
Summary: lipid homeostasis is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0060075 regulation of resting membrane potential
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: regulation of resting membrane potential is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0060100 positive regulation of phagocytosis, engulfment
IEA
GO_REF:0000120
ACCEPT
Summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0061518 microglial cell proliferation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: microglial cell proliferation is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0061889 negative regulation of astrocyte activation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of astrocyte activation is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0070345 negative regulation of fat cell proliferation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of fat cell proliferation is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0070374 positive regulation of ERK1 and ERK2 cascade
IEA
GO_REF:0000120
ACCEPT
Summary: positive regulation of ERK1 and ERK2 cascade is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0070392 detection of lipoteichoic acid
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: detection of lipoteichoic acid is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0070891 lipoteichoic acid binding
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: lipoteichoic acid binding is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0071223 cellular response to lipoteichoic acid
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: cellular response to lipoteichoic acid is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0071224 cellular response to peptidoglycan
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: cellular response to peptidoglycan is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0071333 cellular response to glucose stimulus
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: cellular response to glucose stimulus is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0071456 cellular response to hypoxia
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: cellular response to hypoxia is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0071526 semaphorin-plexin signaling pathway
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: semaphorin-plexin signaling pathway is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0071640 regulation of macrophage inflammatory protein 1 alpha production
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: regulation of macrophage inflammatory protein 1 alpha production is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0097062 dendritic spine maintenance
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: dendritic spine maintenance is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0097242 amyloid-beta clearance
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0098657 import into cell
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: import into cell is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0110089 regulation of hippocampal neuron apoptotic process
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: regulation of hippocampal neuron apoptotic process is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0120035 regulation of plasma membrane bounded cell projection organization
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: regulation of plasma membrane bounded cell projection organization is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0150062 complement-mediated synapse pruning
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: complement-mediated synapse pruning is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0150076 neuroinflammatory response
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: neuroinflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0150078 positive regulation of neuroinflammatory response
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of neuroinflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0150079 negative regulation of neuroinflammatory response
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: negative regulation of neuroinflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0150094 amyloid-beta clearance by cellular catabolic process
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: amyloid-beta clearance by cellular catabolic process is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1900015 regulation of cytokine production involved in inflammatory response
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: regulation of cytokine production involved in inflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1900016 negative regulation of cytokine production involved in inflammatory response
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of cytokine production involved in inflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1900223 positive regulation of amyloid-beta clearance
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: positive regulation of amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1901224 positive regulation of non-canonical NF-kappaB signal transduction
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of non-canonical NF-kappaB signal transduction is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1901800 positive regulation of proteasomal protein catabolic process
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of proteasomal protein catabolic process is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1902533 positive regulation of intracellular signal transduction
IEA
GO_REF:0000107
ACCEPT
Summary: positive regulation of intracellular signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:1903376 regulation of oxidative stress-induced neuron intrinsic apoptotic signaling pathway
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: regulation of oxidative stress-induced neuron intrinsic apoptotic signaling pathway is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1903753 negative regulation of p38MAPK cascade
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of p38MAPK cascade is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1903980 positive regulation of microglial cell activation
IEA
GO_REF:0000120
ACCEPT
Summary: positive regulation of microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1904093 negative regulation of autophagic cell death
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of autophagic cell death is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1904141 positive regulation of microglial cell migration
IEA
GO_REF:0000120
ACCEPT
Summary: positive regulation of microglial cell migration is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1904646 cellular response to amyloid-beta
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: cellular response to amyloid-beta is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1904951 positive regulation of establishment of protein localization
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: positive regulation of establishment of protein localization is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1905291 positive regulation of CAMKK-AMPK signaling cascade
IEA
GO_REF:0000107
ACCEPT
Summary: positive regulation of CAMKK-AMPK signaling cascade is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:1905581 positive regulation of low-density lipoprotein particle clearance
IEA
GO_REF:0000107
ACCEPT
Summary: positive regulation of low-density lipoprotein particle clearance is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:1905805 excitatory synapse pruning
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: excitatory synapse pruning is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1905907 negative regulation of amyloid fibril formation
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: negative regulation of amyloid fibril formation is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1990782 protein tyrosine kinase binding
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: protein tyrosine kinase binding is a specific interaction context for TREM2 but not the main molecular-function term.
Reason: Retain as non-core because the interaction may be real, while more informative core terms describe receptor activity and lipid/lipoprotein/aminophospholipid ligand binding.
GO:2001171 positive regulation of ATP biosynthetic process
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: positive regulation of ATP biosynthetic process is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1902533 positive regulation of intracellular signal transduction
IDA
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
ACCEPT
Summary: positive regulation of intracellular signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0005886 plasma membrane
EXP
PMID:25615530
Disease-Associated Mutations of TREM2 Alter the Processing o...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
EXP
PMID:28768830
Neurodegeneration-associated mutant TREM2 proteins abortivel...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
EXP
PMID:28923481
ADAM17 is the main sheddase for the generation of human trig...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0010468 regulation of gene expression
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
KEEP AS NON CORE
Summary: regulation of gene expression is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0032675 regulation of interleukin-6 production
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
KEEP AS NON CORE
Summary: regulation of interleukin-6 production is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0043268 positive regulation of potassium ion transport
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
KEEP AS NON CORE
Summary: positive regulation of potassium ion transport is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0071640 regulation of macrophage inflammatory protein 1 alpha production
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
KEEP AS NON CORE
Summary: regulation of macrophage inflammatory protein 1 alpha production is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1902533 positive regulation of intracellular signal transduction
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
ACCEPT
Summary: positive regulation of intracellular signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:1902533 positive regulation of intracellular signal transduction
IDA
PMID:31902528
TREM2 Regulates Microglial Cholesterol Metabolism upon Chron...
ACCEPT
Summary: positive regulation of intracellular signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0002116 semaphorin receptor complex
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: semaphorin receptor complex is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0002291 T cell activation via T cell receptor contact with antigen bound to MHC molecule on antigen presenting cell
IDA
PMID:16715077
Plexin-A1 and its interaction with DAP12 in immune responses...
KEEP AS NON CORE
Summary: T cell activation via T cell receptor contact with antigen bound to MHC molecule on antigen presenting cell is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0005886 plasma membrane
IDA
PMID:16715077
Plexin-A1 and its interaction with DAP12 in immune responses...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0017154 semaphorin receptor activity
IDA
PMID:16715077
Plexin-A1 and its interaction with DAP12 in immune responses...
KEEP AS NON CORE
Summary: semaphorin receptor activity is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0071526 semaphorin-plexin signaling pathway
IDA
PMID:16715077
Plexin-A1 and its interaction with DAP12 in immune responses...
KEEP AS NON CORE
Summary: semaphorin-plexin signaling pathway is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0004888 transmembrane signaling receptor activity
IDA
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
ACCEPT
Summary: TREM2 transmembrane signaling receptor activity is a core plasma-membrane receptor function coupled to TYROBP/DAP12 signaling.
Reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives intracellular signaling through DAP12/TYROBP-linked kinase pathways.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0004888 transmembrane signaling receptor activity
IDA
PMID:30333625
LILRB4 signalling in leukaemia cells mediates T cell suppres...
ACCEPT
Summary: TREM2 transmembrane signaling receptor activity is a core plasma-membrane receptor function coupled to TYROBP/DAP12 signaling.
Reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives intracellular signaling through DAP12/TYROBP-linked kinase pathways.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0005886 plasma membrane
IDA
PMID:24078628
Sequential proteolytic processing of the triggering receptor...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
IDA
PMID:27477018
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ,...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0071402 cellular response to lipoprotein particle stimulus
IDA
PMID:27477018
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ,...
ACCEPT
Summary: cellular response to lipoprotein particle stimulus is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0097242 amyloid-beta clearance
IDA
PMID:27477018
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ,...
KEEP AS NON CORE
Summary: amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0008013 beta-catenin binding
IPI
PMID:30683932
TREM2 acts as a tumor suppressor in hepatocellular carcinoma...
KEEP AS NON CORE
Summary: beta-catenin binding is a specific interaction context for TREM2 but not the main molecular-function term.
Reason: Retain as non-core because the interaction may be real, while more informative core terms describe receptor activity and lipid/lipoprotein/aminophospholipid ligand binding.
GO:0051898 negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
IMP
PMID:30683932
TREM2 acts as a tumor suppressor in hepatocellular carcinoma...
ACCEPT
Summary: negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0050921 positive regulation of chemotaxis
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of chemotaxis is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0120035 regulation of plasma membrane bounded cell projection organization
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: regulation of plasma membrane bounded cell projection organization is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1904141 positive regulation of microglial cell migration
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of microglial cell migration is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0010507 negative regulation of autophagy
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of autophagy is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0032006 regulation of TOR signaling
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: regulation of TOR signaling is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0032008 positive regulation of TOR signaling
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: positive regulation of TOR signaling is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1904093 negative regulation of autophagic cell death
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of autophagic cell death is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1905291 positive regulation of CAMKK-AMPK signaling cascade
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of CAMKK-AMPK signaling cascade is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:2001171 positive regulation of ATP biosynthetic process
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: positive regulation of ATP biosynthetic process is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1900223 positive regulation of amyloid-beta clearance
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: positive regulation of amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0006910 phagocytosis, recognition
IMP
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
ACCEPT
Summary: phagocytosis, recognition is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0010628 positive regulation of gene expression
IMP
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
KEEP AS NON CORE
Summary: positive regulation of gene expression is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0019216 regulation of lipid metabolic process
IMP
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
ACCEPT
Summary: regulation of lipid metabolic process is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0038023 signaling receptor activity
IMP
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
ACCEPT
Summary: TREM2 signaling receptor activity is a core plasma-membrane receptor function coupled to TYROBP/DAP12 signaling.
Reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives intracellular signaling through DAP12/TYROBP-linked kinase pathways.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0048678 response to axon injury
IMP
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
KEEP AS NON CORE
Summary: response to axon injury is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0060100 positive regulation of phagocytosis, engulfment
IMP
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
ACCEPT
Summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0071396 cellular response to lipid
IMP
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
ACCEPT
Summary: cellular response to lipid is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0150079 negative regulation of neuroinflammatory response
IMP
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
KEEP AS NON CORE
Summary: negative regulation of neuroinflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1900226 negative regulation of NLRP3 inflammasome complex assembly
IMP
PMID:32514138
Alzheimer's-associated PLCγ2 is a signaling node required fo...
KEEP AS NON CORE
Summary: negative regulation of NLRP3 inflammasome complex assembly is retained as non-core TREM2-associated biology pending deeper reference-specific adjudication.
Reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic microglial function set, but the local evidence was not sufficient to remove it.
GO:0038160 CXCL12-activated CXCR4 signaling pathway
IMP
PMID:33097708
Gene expression and functional deficits underlie TREM2-knock...
KEEP AS NON CORE
Summary: CXCL12-activated CXCR4 signaling pathway is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0050829 defense response to Gram-negative bacterium
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: defense response to Gram-negative bacterium is electronically inferred pathogen-associated ligand or defense biology.
Reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and microglial phagocytic ligands.
GO:0070269 pyroptotic inflammatory response
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: pyroptotic inflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1900016 negative regulation of cytokine production involved in inflammatory response
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of cytokine production involved in inflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1900226 negative regulation of NLRP3 inflammasome complex assembly
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of NLRP3 inflammasome complex assembly is retained as non-core TREM2-associated biology pending deeper reference-specific adjudication.
Reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic microglial function set, but the local evidence was not sufficient to remove it.
GO:0150079 negative regulation of neuroinflammatory response
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of neuroinflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1903753 negative regulation of p38MAPK cascade
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of p38MAPK cascade is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0034144 negative regulation of toll-like receptor 4 signaling pathway
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of toll-like receptor 4 signaling pathway is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0050866 negative regulation of cell activation
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of cell activation is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0051897 positive regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0061889 negative regulation of astrocyte activation
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of astrocyte activation is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0110089 regulation of hippocampal neuron apoptotic process
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: regulation of hippocampal neuron apoptotic process is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1903376 regulation of oxidative stress-induced neuron intrinsic apoptotic signaling pathway
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: regulation of oxidative stress-induced neuron intrinsic apoptotic signaling pathway is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0001786 phosphatidylserine binding
IDA
PMID:31101881
Aminophospholipids are signal-transducing TREM2 ligands on a...
ACCEPT
Summary: TREM2 phosphatidylserine binding is part of its ligand-sensing receptor biology.
Reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function for apoptotic-cell and debris responses.
Supporting Evidence:
PMID:31101881
TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
GO:0002282 microglial cell activation involved in immune response
ISS
GO_REF:0000024
ACCEPT
Summary: microglial cell activation involved in immune response is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
IDA
PMID:31462511
Increased soluble TREM2 in cerebrospinal fluid is associated...
KEEP AS NON CORE
Summary: memory is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0008429 phosphatidylethanolamine binding
IDA
PMID:31101881
Aminophospholipids are signal-transducing TREM2 ligands on a...
ACCEPT
Summary: TREM2 phosphatidylethanolamine binding is part of its ligand-sensing receptor biology.
Reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function for apoptotic-cell and debris responses.
Supporting Evidence:
PMID:31101881
TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
GO:0034136 negative regulation of toll-like receptor 2 signaling pathway
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of toll-like receptor 2 signaling pathway is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0034151 regulation of toll-like receptor 6 signaling pathway
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: regulation of toll-like receptor 6 signaling pathway is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0097062 dendritic spine maintenance
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: dendritic spine maintenance is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0150094 amyloid-beta clearance by cellular catabolic process
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: amyloid-beta clearance by cellular catabolic process is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0150094 amyloid-beta clearance by cellular catabolic process
IMP
PMID:33097708
Gene expression and functional deficits underlie TREM2-knock...
KEEP AS NON CORE
Summary: amyloid-beta clearance by cellular catabolic process is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0001774 microglial cell activation
ISS
GO_REF:0000024
ACCEPT
Summary: microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0002862 negative regulation of inflammatory response to antigenic stimulus
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of inflammatory response to antigenic stimulus is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
IMP
PMID:29518357
Elevated TREM2 Gene Dosage Reprograms Microglia Responsivity...
KEEP AS NON CORE
Summary: memory is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0034351 negative regulation of glial cell apoptotic process
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of glial cell apoptotic process is retained as non-core TREM2-associated biology pending deeper reference-specific adjudication.
Reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic microglial function set, but the local evidence was not sufficient to remove it.
GO:0043124 negative regulation of canonical NF-kappaB signal transduction
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of canonical NF-kappaB signal transduction is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0050766 positive regulation of phagocytosis
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of phagocytosis is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0050766 positive regulation of phagocytosis
IMP
PMID:29518357
Elevated TREM2 Gene Dosage Reprograms Microglia Responsivity...
ACCEPT
Summary: positive regulation of phagocytosis is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0051898 negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
ISS
GO_REF:0000024
ACCEPT
Summary: negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:1900015 regulation of cytokine production involved in inflammatory response
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: regulation of cytokine production involved in inflammatory response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0060100 positive regulation of phagocytosis, engulfment
IMP
PMID:33097708
Gene expression and functional deficits underlie TREM2-knock...
ACCEPT
Summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1902227 negative regulation of macrophage colony-stimulating factor signaling pathway
IMP
PMID:33097708
Gene expression and functional deficits underlie TREM2-knock...
KEEP AS NON CORE
Summary: negative regulation of macrophage colony-stimulating factor signaling pathway is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:1904141 positive regulation of microglial cell migration
IMP
PMID:33097708
Gene expression and functional deficits underlie TREM2-knock...
ACCEPT
Summary: positive regulation of microglial cell migration is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1905808 positive regulation of synapse pruning
IMP
PMID:33097708
Gene expression and functional deficits underlie TREM2-knock...
KEEP AS NON CORE
Summary: positive regulation of synapse pruning is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0035176 social behavior
IMP
PMID:29752066
The Microglial Innate Immune Receptor TREM2 Is Required for ...
KEEP AS NON CORE
Summary: social behavior is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0045088 regulation of innate immune response
ISS
GO_REF:0000024
ACCEPT
Summary: regulation of innate immune response is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0045728 respiratory burst after phagocytosis
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: respiratory burst after phagocytosis is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0045960 positive regulation of complement activation, classical pathway
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: positive regulation of complement activation, classical pathway is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0061518 microglial cell proliferation
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: microglial cell proliferation is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0150062 complement-mediated synapse pruning
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: complement-mediated synapse pruning is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0060100 positive regulation of phagocytosis, engulfment
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1905805 excitatory synapse pruning
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: excitatory synapse pruning is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0048678 response to axon injury
IMP
PMID:31235932
TREM2 function impedes tau seeding in neuritic plaques.
KEEP AS NON CORE
Summary: response to axon injury is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1900223 positive regulation of amyloid-beta clearance
IMP
PMID:31235932
TREM2 function impedes tau seeding in neuritic plaques.
KEEP AS NON CORE
Summary: positive regulation of amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0001786 phosphatidylserine binding
IDA
PMID:31902528
TREM2 Regulates Microglial Cholesterol Metabolism upon Chron...
ACCEPT
Summary: TREM2 phosphatidylserine binding is part of its ligand-sensing receptor biology.
Reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function for apoptotic-cell and debris responses.
Supporting Evidence:
PMID:31101881
TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
GO:0010875 positive regulation of cholesterol efflux
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of cholesterol efflux is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0010887 negative regulation of cholesterol storage
ISS
GO_REF:0000024
ACCEPT
Summary: negative regulation of cholesterol storage is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0010891 negative regulation of triglyceride storage
ISS
GO_REF:0000024
ACCEPT
Summary: negative regulation of triglyceride storage is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0048678 response to axon injury
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: response to axon injury is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0120146 sulfatide binding
IDA
PMID:31902528
TREM2 Regulates Microglial Cholesterol Metabolism upon Chron...
ACCEPT
Summary: TREM2 sulfatide binding is part of its ligand-sensing receptor biology.
Reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function for apoptotic-cell and debris responses.
Supporting Evidence:
PMID:31101881
TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
GO:0140052 cellular response to oxidised low-density lipoprotein particle stimulus
IDA
PMID:31902528
TREM2 Regulates Microglial Cholesterol Metabolism upon Chron...
ACCEPT
Summary: cellular response to oxidised low-density lipoprotein particle stimulus is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:1903980 positive regulation of microglial cell activation
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1904951 positive regulation of establishment of protein localization
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: positive regulation of establishment of protein localization is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1905907 negative regulation of amyloid fibril formation
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of amyloid fibril formation is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0019209 kinase activator activity
IMP
PMID:31902528
TREM2 Regulates Microglial Cholesterol Metabolism upon Chron...
ACCEPT
Summary: kinase activator activity is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0010983 positive regulation of high-density lipoprotein particle clearance
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of high-density lipoprotein particle clearance is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0055088 lipid homeostasis
ISS
GO_REF:0000024
ACCEPT
Summary: lipid homeostasis is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0070345 negative regulation of fat cell proliferation
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: negative regulation of fat cell proliferation is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1905581 positive regulation of low-density lipoprotein particle clearance
ISS
GO_REF:0000024
ACCEPT
Summary: positive regulation of low-density lipoprotein particle clearance is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol handling.
Supporting Evidence:
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
PMID:31902528
Our studies identify TREM2 as a key transcriptional regulator of cholesterol transport and metabolism under conditions of chronic myelin phagocytic activity
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0044853 plasma membrane raft
IDA
PMID:31413141
The MS4A gene cluster is a key modulator of soluble TREM2 an...
ACCEPT
Summary: TREM2 localization to plasma membrane raft is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0001774 microglial cell activation
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
ACCEPT
Summary: microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0048143 astrocyte activation
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: astrocyte activation is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0005886 plasma membrane
IMP
PMID:24990881
TREM2 mutations implicated in neurodegeneration impair cell ...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
IMP
PMID:27589997
Rare TREM2 variants associated with Alzheimer's disease disp...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
IMP
PMID:28855300
An Alzheimer-associated TREM2 variant occurs at the ADAM cle...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0060100 positive regulation of phagocytosis, engulfment
IMP
PMID:28855300
An Alzheimer-associated TREM2 variant occurs at the ADAM cle...
ACCEPT
Summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1901076 positive regulation of engulfment of apoptotic cell
IMP
PMID:24990881
TREM2 mutations implicated in neurodegeneration impair cell ...
ACCEPT
Summary: positive regulation of engulfment of apoptotic cell is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1903980 positive regulation of microglial cell activation
IMP
PMID:24990881
TREM2 mutations implicated in neurodegeneration impair cell ...
ACCEPT
Summary: positive regulation of microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0030316 osteoclast differentiation
IMP
PMID:12925681
DAP12/TREM2 deficiency results in impaired osteoclast differ...
KEEP AS NON CORE
Summary: osteoclast differentiation is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
Supporting Evidence:
PMID:12925681
loss of function mutations in DAP12 and TREM2 result in an inefficient and delayed differentiation of osteoclasts
GO:0005886 plasma membrane
IMP
PMID:28855301
TREM2 shedding by cleavage at the H157-S158 bond is accelera...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0050714 positive regulation of protein secretion
IMP
PMID:27044754
FRMD4A-cytohesin signaling modulates the cellular release of...
KEEP AS NON CORE
Summary: positive regulation of protein secretion is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0043277 apoptotic cell clearance
ISS
GO_REF:0000024
ACCEPT
Summary: apoptotic cell clearance is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:0008289 lipid binding
TAS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
ACCEPT
Summary: TREM2 lipid binding is part of its ligand-sensing receptor biology.
Reason: TREM2 ligand recognition includes lipid-rich particles and supports microglial lipid handling.
Supporting Evidence:
PMID:27477018
we identified a set of lipoprotein particles (including LDL) and apolipoproteins (including CLU/APOJ and APOE) as ligands
PMID:31902528
TREM2 senses lipids and mediates myelin phagocytosis
GO:0038023 signaling receptor activity
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
ACCEPT
Summary: TREM2 signaling receptor activity is a core plasma-membrane receptor function coupled to TYROBP/DAP12 signaling.
Reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives intracellular signaling through DAP12/TYROBP-linked kinase pathways.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0060075 regulation of resting membrane potential
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
KEEP AS NON CORE
Summary: regulation of resting membrane potential is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0120035 regulation of plasma membrane bounded cell projection organization
IGI
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
KEEP AS NON CORE
Summary: regulation of plasma membrane bounded cell projection organization is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1901800 positive regulation of proteasomal protein catabolic process
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
KEEP AS NON CORE
Summary: positive regulation of proteasomal protein catabolic process is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1904141 positive regulation of microglial cell migration
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
ACCEPT
Summary: positive regulation of microglial cell migration is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
Reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, activation, and migration responses.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1904646 cellular response to amyloid-beta
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
KEEP AS NON CORE
Summary: cellular response to amyloid-beta is Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than the broad core receptor function.
Reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial function; do not treat them as disease-progression hypotheses in the core function.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
PMID:33097708
TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
GO:1990782 protein tyrosine kinase binding
ISS
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
KEEP AS NON CORE
Summary: protein tyrosine kinase binding is a specific interaction context for TREM2 but not the main molecular-function term.
Reason: Retain as non-core because the interaction may be real, while more informative core terms describe receptor activity and lipid/lipoprotein/aminophospholipid ligand binding.
GO:0001540 amyloid-beta binding
IPI
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
ACCEPT
Summary: TREM2 amyloid-beta binding is part of its ligand-sensing receptor biology.
Reason: Amyloid-beta binding is disease-relevant but mechanistically tied to TREM2 ligand uptake through lipoprotein-associated complexes; retain as part of ligand recognition.
Supporting Evidence:
PMID:27477018
β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up by microglia in a TREM2-dependent fashion.
GO:0044877 protein-containing complex binding
IPI
PMID:29518356
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial F...
MARK AS OVER ANNOTATED
Summary: Generic protein-containing complex binding annotation for TREM2.
Reason: The interaction may be real, but this term is too generic. Specific ligand-binding, receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
GO:0010628 positive regulation of gene expression
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: positive regulation of gene expression is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0098657 import into cell
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: import into cell is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0008035 high-density lipoprotein particle binding
IDA
PMID:27477018
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ,...
ACCEPT
Summary: TREM2 high-density lipoprotein particle binding is part of its ligand-sensing receptor biology.
Reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by microglia or receptor-expressing cells.
Supporting Evidence:
PMID:27477018
we identified a set of lipoprotein particles (including LDL) and apolipoproteins (including CLU/APOJ and APOE) as ligands
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0030169 low-density lipoprotein particle binding
IDA
PMID:27477018
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ,...
ACCEPT
Summary: TREM2 low-density lipoprotein particle binding is part of its ligand-sensing receptor biology.
Reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by microglia or receptor-expressing cells.
Supporting Evidence:
PMID:27477018
we identified a set of lipoprotein particles (including LDL) and apolipoproteins (including CLU/APOJ and APOE) as ligands
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0034185 apolipoprotein binding
IPI
PMID:27477018
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ,...
ACCEPT
Summary: TREM2 apolipoprotein binding is part of its ligand-sensing receptor biology.
Reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by microglia or receptor-expressing cells.
Supporting Evidence:
PMID:27477018
we identified a set of lipoprotein particles (including LDL) and apolipoproteins (including CLU/APOJ and APOE) as ligands
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0034186 apolipoprotein A-I binding
IPI
PMID:27477018
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ,...
ACCEPT
Summary: TREM2 apolipoprotein A-I binding is part of its ligand-sensing receptor biology.
Reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by microglia or receptor-expressing cells.
Supporting Evidence:
PMID:27477018
we identified a set of lipoprotein particles (including LDL) and apolipoproteins (including CLU/APOJ and APOE) as ligands
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0034189 very-low-density lipoprotein particle binding
IDA
PMID:27477018
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ,...
ACCEPT
Summary: TREM2 very-low-density lipoprotein particle binding is part of its ligand-sensing receptor biology.
Reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by microglia or receptor-expressing cells.
Supporting Evidence:
PMID:27477018
we identified a set of lipoprotein particles (including LDL) and apolipoproteins (including CLU/APOJ and APOE) as ligands
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0071813 lipoprotein particle binding
IDA
PMID:27477018
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ,...
ACCEPT
Summary: TREM2 lipoprotein particle binding is part of its ligand-sensing receptor biology.
Reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by microglia or receptor-expressing cells.
Supporting Evidence:
PMID:27477018
we identified a set of lipoprotein particles (including LDL) and apolipoproteins (including CLU/APOJ and APOE) as ligands
PMID:27477018
Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, CLU, and APOE in heterologous cells
GO:0034241 positive regulation of macrophage fusion
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: positive regulation of macrophage fusion is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0030316 osteoclast differentiation
IMP
PMID:21841309
OSCAR is a collagen receptor that costimulates osteoclastoge...
KEEP AS NON CORE
Summary: osteoclast differentiation is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
Supporting Evidence:
PMID:12925681
loss of function mutations in DAP12 and TREM2 result in an inefficient and delayed differentiation of osteoclasts
GO:0005543 phospholipid binding
IDA
PMID:27995897
Neurodegenerative disease mutations in TREM2 reveal a functi...
ACCEPT
Summary: TREM2 phospholipid binding is part of its ligand-sensing receptor biology.
Reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function for apoptotic-cell and debris responses.
Supporting Evidence:
PMID:31101881
TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
GO:0005886 plasma membrane
IDA
PMID:27995897
Neurodegenerative disease mutations in TREM2 reveal a functi...
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0070374 positive regulation of ERK1 and ERK2 cascade
IMP
PMID:27995897
Neurodegenerative disease mutations in TREM2 reveal a functi...
ACCEPT
Summary: positive regulation of ERK1 and ERK2 cascade is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0002588 positive regulation of antigen processing and presentation of peptide antigen via MHC class II
IDA
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
KEEP AS NON CORE
Summary: positive regulation of antigen processing and presentation of peptide antigen via MHC class II is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.
GO:0016020 membrane
IDA
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
ACCEPT
Summary: TREM2 localization to membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0050850 positive regulation of calcium-mediated signaling
IDA
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
ACCEPT
Summary: positive regulation of calcium-mediated signaling is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0070374 positive regulation of ERK1 and ERK2 cascade
IDA
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
ACCEPT
Summary: positive regulation of ERK1 and ERK2 cascade is a core downstream consequence of TREM2 receptor signaling.
Reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support calcium/ERK/PI3K-related activation and intracellular signaling.
Supporting Evidence:
PMID:11602640
TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
PMID:32514138
TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.
GO:0097028 dendritic cell differentiation
IDA
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
KEEP AS NON CORE
Summary: dendritic cell differentiation is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0097110 scaffold protein binding
IPI
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
KEEP AS NON CORE
Summary: scaffold protein binding is a specific interaction context for TREM2 but not the main molecular-function term.
Reason: Retain as non-core because the interaction may be real, while more informative core terms describe receptor activity and lipid/lipoprotein/aminophospholipid ligand binding.
GO:1903078 positive regulation of protein localization to plasma membrane
IDA
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
KEEP AS NON CORE
Summary: positive regulation of protein localization to plasma membrane is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:1903082 positive regulation of C-C chemokine receptor CCR7 signaling pathway
IDA
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
KEEP AS NON CORE
Summary: positive regulation of C-C chemokine receptor CCR7 signaling pathway is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:2000350 positive regulation of CD40 signaling pathway
IDA
PMID:11602640
A DAP12-mediated pathway regulates expression of CC chemokin...
KEEP AS NON CORE
Summary: positive regulation of CD40 signaling pathway is retained as context-specific TREM2-associated biology.
Reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the core TREM2 receptor-ligand signaling mechanism.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-210289
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-210300
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2395412
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2395801
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2396594
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2424480
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2424482
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2424484
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2424486
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-2424487
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-416725
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0005886 plasma membrane
TAS
Reactome:R-HSA-5696358
ACCEPT
Summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface receptor role.
Reason: Cell-surface/plasma-membrane localization is essential for ligand binding and DAP12/TYROBP-mediated signal transduction.
Supporting Evidence:
PMID:11602640
TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
PMID:24990881
missense mutations associated with FTD and FTD-like syndrome reduce TREM2 maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of TREM2-expressing cells.
PMID:27995897
Nasu-Hakola mutations impact protein stability and decrease folded TREM2 surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
GO:0006959 humoral immune response
TAS
PMID:10799849
Cutting edge: inflammatory responses can be triggered by TRE...
KEEP AS NON CORE
Summary: humoral immune response is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
Reason: TREM2 modulates myeloid and microglial immune responses, but specific cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
Supporting Evidence:
PMID:24990881
TREM2 is an innate immune receptor preferentially expressed by microglia and is involved in inflammation and phagocytosis.

Core Functions

TREM2 is a plasma-membrane myeloid receptor that associates with TYROBP/DAP12 and transduces ligand binding into intracellular kinase, PLCG2, calcium, ERK, and activation programs in microglia and other myeloid cells.

Supporting Evidence:
  • PMID:11602640
    TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is associated with DAP12.
  • PMID:11602640
    TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC maturation, and DC survival through activation of protein tyrosine kinases and extracellular signal-regulated kinase.
  • PMID:32514138
    TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing of neuronal debris, and lipid metabolism.

TREM2 recognizes lipid-rich damage and uptake ligands, including aminophospholipids on apoptotic cells, lipoprotein/apolipoprotein particles, sulfatides, and amyloid-beta-lipoprotein complexes, enabling microglial phagocytosis, activation, migration, and lipid handling.

Supporting Evidence:
  • PMID:27477018
    we identified a set of lipoprotein particles (including LDL) and apolipoproteins (including CLU/APOJ and APOE) as ligands
  • PMID:31101881
    TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
  • PMID:31902528
    TREM2 senses lipids and mediates myelin phagocytosis
  • PMID:33097708
    TREM2 deletion reduces microglial survival, impairs phagocytosis of key substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis

References

Manual transfer of experimentally-verified manual GO annotation data to orthologs by curator judgment of sequence similarity
Annotation inferences using phylogenetic trees
Gene Ontology annotation based on UniProtKB/Swiss-Prot Subcellular Location vocabulary mapping, accompanied by conservative changes to GO terms applied by UniProt
Automatic transfer of experimentally verified manual GO annotation data to orthologs using Ensembl Compara
Electronic Gene Ontology annotations created by ARBA machine learning models
Combined Automated Annotation using Multiple IEA Methods
Cutting edge: inflammatory responses can be triggered by TREM-1, a novel receptor expressed on neutrophils and monocytes.
A DAP12-mediated pathway regulates expression of CC chemokine receptor 7 and maturation of human dendritic cells.
DAP12/TREM2 deficiency results in impaired osteoclast differentiation and osteoporotic features.
Plexin-A1 and its interaction with DAP12 in immune responses and bone homeostasis.
OSCAR is a collagen receptor that costimulates osteoclastogenesis in DAP12-deficient humans and mice.
Sequential proteolytic processing of the triggering receptor expressed on myeloid cells-2 (TREM2) protein by ectodomain shedding and γ-secretase-dependent intramembranous cleavage.
TREM2 mutations implicated in neurodegeneration impair cell surface transport and phagocytosis.
Disease-Associated Mutations of TREM2 Alter the Processing of N-Linked Oligosaccharides in the Golgi Apparatus.
FRMD4A-cytohesin signaling modulates the cellular release of tau.
TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ, and Thereby Facilitates Uptake of Amyloid-Beta by Microglia.
Rare TREM2 variants associated with Alzheimer's disease display reduced cell surface expression.
Neurodegenerative disease mutations in TREM2 reveal a functional surface and distinct loss-of-function mechanisms.
A split-luciferase complementation, real-time reporting assay enables monitoring of the disease-associated transmembrane protein TREM2 in live cells.
Neurodegeneration-associated mutant TREM2 proteins abortively cycle between the ER and ER-Golgi intermediate compartment.
An Alzheimer-associated TREM2 variant occurs at the ADAM cleavage site and affects shedding and phagocytic function.
TREM2 shedding by cleavage at the H157-S158 bond is accelerated for the Alzheimer's disease-associated H157Y variant.
ADAM17 is the main sheddase for the generation of human triggering receptor expressed in myeloid cells (hTREM2) ectodomain and cleaves TREM2 after Histidine 157.
TREM2 Is a Receptor for β-Amyloid that Mediates Microglial Function.
Elevated TREM2 Gene Dosage Reprograms Microglia Responsivity and Ameliorates Pathological Phenotypes in Alzheimer's Disease Models.
Intracellular trafficking of TREM2 is regulated by presenilin 1.
The Microglial Innate Immune Receptor TREM2 Is Required for Synapse Elimination and Normal Brain Connectivity.
LILRB4 signalling in leukaemia cells mediates T cell suppression and tumour infiltration.
High-affinity interactions and signal transduction between Aβ oligomers and TREM2.
TREM2 acts as a tumor suppressor in hepatocellular carcinoma by targeting the PI3K/Akt/β-catenin pathway.
Aminophospholipids are signal-transducing TREM2 ligands on apoptotic cells.
TREM2 function impedes tau seeding in neuritic plaques.
The MS4A gene cluster is a key modulator of soluble TREM2 and Alzheimer's disease risk.
Increased soluble TREM2 in cerebrospinal fluid is associated with reduced cognitive and clinical decline in Alzheimer's disease.
TREM2 Regulates Microglial Cholesterol Metabolism upon Chronic Phagocytic Challenge.
Alzheimer's-associated PLCγ2 is a signaling node required for both TREM2 function and the inflammatory response in human microglia.
Gene expression and functional deficits underlie TREM2-knockout microglia responses in human models of Alzheimer's disease.
Reactome:R-HSA-210289
Recruitment of SYK to p-DAP12
Reactome:R-HSA-210300
Interaction of DAP12 and TREM2
Reactome:R-HSA-2395412
Phosphorylation of SYK
Reactome:R-HSA-2395801
Phosphorylation of LAT by p-SYK
Reactome:R-HSA-2396594
Phosphorylation of SLP-76 by p-SYK
Reactome:R-HSA-2424480
PI3K phosphorylates PIP2 to PIP3
Reactome:R-HSA-2424482
p85 regulatory unit of PI3K binds p-6Y-SYK
Reactome:R-HSA-2424484
Phosphorylation of BTK by p-SYK
Reactome:R-HSA-2424486
Phosphorylation and activation of VAV2/VAV3 by SYK
Reactome:R-HSA-2424487
Phosphorylation of PLC-gamma by p-BTK/p-SYK
Reactome:R-HSA-416725
SEMA6D binds to PLXNA1:TREM2:DAP12
Reactome:R-HSA-5696358
TREM,CD300 binds lipids

Suggested Questions for Experts

Q: Which TREM2 ligand classes are normal in-vivo microglial ligands across brain states, and which are disease-model or injury-specific substrates?

Suggested experts: microglial biology experts, GO immune receptor curators

Q: How should TREM2/TYROBP/DAP12 signaling be represented without over-curating every downstream cytokine, kinase, survival, or metabolic phenotype as a core function?

Suggested experts: innate immune signaling curators, microglial signaling experts

Q: Which amyloid-beta, synapse-pruning, and neuroinflammatory annotations reflect evolved TREM2 biology versus Alzheimer model-stage-specific phenotypes?

Suggested experts: Alzheimer microglia experts, GO neuroimmune curators

Suggested Experiments

Experiment: Compare endogenous human TREM2 knock-in and knockout microglia across defined ligands: apoptotic cells, purified aminophospholipids, APOE/CLU lipoprotein particles, myelin debris, synaptosomes, and amyloid-beta-lipoprotein complexes.

Hypothesis: TREM2 core ligand sensing can be decomposed into ligand classes with distinct requirements for TYROBP/DAP12, PLCG2, SYK, and calcium signaling.

Type: ligand-defined microglial uptake and signaling assays

Experiment: Use in-vivo baseline, injury, demyelination, and amyloid/tau model time courses to separate normal TREM2-dependent phagocytic functions from disease-stage-specific microglial states.

Hypothesis: Some TREM2 annotations, especially amyloid clearance and synapse pruning, are context-specific outputs of the same core receptor-phagocytosis program.

Type: time-course in-vivo microglial transcriptomics and functional assays

Experiment: Quantify cell-surface TREM2, soluble TREM2, and TYROBP-associated signaling after endogenous AD-risk variant knock-in under non-disease and challenge conditions.

Hypothesis: Disease-associated variants alter ligand binding, surface trafficking, or signaling amplitude without changing the core receptor function category.

Type: endogenous variant knock-in receptor trafficking and signaling

📚 Additional Documentation

Notes

(TREM2-notes.md)

TREM2 notes

Review status

  • First-pass review completed on 2026-06-19.
  • just fetch-gene-pmids human TREM2 completed successfully; all 31 PMID-backed publication caches were present after refresh.
  • Falcon deep research was attempted with timeout 180 just deep-research-falcon human TREM2 --fallback perplexity-lite, but the process timed out and no provider deep-research artifact was written. These notes therefore rely on the cached UniProt, GOA, and publication files.
  • just validate human TREM2 passes cleanly.

Functional synthesis

TREM2 is a plasma-membrane immunoreceptor expressed prominently in myeloid cells, including microglia. Its core function is ligand sensing through an extracellular immunoglobulin-like domain, with signaling through TYROBP/DAP12 and downstream kinase/Ca2+/ERK pathways. The original receptor characterization supports the surface receptor and DAP12 coupling model: PMID:11602640 The same study supports downstream signaling and survival/maturation responses: PMID:11602640

The best-supported ligand biology is lipid/apolipoprotein and damage-associated membrane sensing. TREM2 binds apolipoprotein and lipoprotein ligands, including APOE and CLU/APOJ: PMID:27477018 This ligand binding is functionally tied to uptake: PMID:27477018 and to microglial handling of amyloid-lipoprotein complexes: PMID:27477018

Independent ligand work supports aminophospholipids on apoptotic cells as signal-transducing TREM2 ligands: PMID:31101881 Myelin/lipid challenge studies further support lipid sensing as a microglial core function: PMID:31902528 and identify a role in cholesterol-related transcriptional adaptation after chronic phagocytosis: PMID:31902528

Human microglia data support the downstream functional consequences of TREM2 loss: PMID:33097708 and broader functional deficits in phagocytosis and migration. Earlier disease-variant work supports a microglial innate immune and phagocytic framing: PMID:24990881 Variant studies also distinguish Nasu-Hakola and Alzheimer-risk mechanisms, with many Alzheimer-associated variants affecting ligand binding rather than complete loss of surface expression: PMID:27995897

Annotation decisions

  • Accepted annotations for transmembrane signaling receptor activity, signaling receptor activity, lipid/apolipoprotein/phospholipid/amyloid ligand binding, TYROBP-linked signaling, phagocytosis/engulfment, apoptotic-cell clearance, microglial migration, microglial activation, lipid homeostasis, cholesterol transport/metabolism regulation, and plasma-membrane localization.
  • Kept non-core annotations for broader disease/pathology phenotypes, amyloid plaque association, synapse and neuronal downstream effects, cytokine/TLR/inflammatory responses, osteoclast and dendritic-cell biology, adaptive immune phenotypes, and general cellular stress or developmental readouts.
  • Marked generic protein binding and broad complex-binding annotations as over-annotated because they obscure the better-supported ligand and receptor-adapter biology.
  • No annotations were removed. Experimental disease-model and immune-cell annotations were retained as non-core when they appear biologically plausible but secondary to the receptor-ligand signaling role.

Final action distribution: 117 ACCEPT, 151 KEEP_AS_NON_CORE, 5 MARK_AS_OVER_ANNOTATED.

Knowledge gaps and experiments

  • The most useful GO-level refinement would be a specific term for TREM2-style damage-associated lipid/apolipoprotein immune receptor activity if the ontology does not already have an exact fit.
  • For Alzheimer-relevant mechanism, the key gap is how APOE isoform, lipidation state, amyloid-lipoprotein complexes, and myelin/apoptotic-cell ligands compete or cooperate in vivo.
  • Useful experiments would combine human iPSC microglia with isogenic TREM2 variants, defined APOE isoform/lipidation substrates, plaque or myelin debris models, and rescue with TYROBP/SYK/PLC signaling perturbations.
  • Spatial in-vivo work should separate direct TREM2 receptor functions from downstream plaque compaction, synapse remodeling, and inflammatory-state effects.

2026-06-20 second-pass audit

The second-pass audit added manual reference_review metadata for the main TREM2 receptor, DAP12 signaling, lipid/apolipoprotein ligand, amyloid-beta ligand, apoptotic-cell aminophospholipid ligand, myelin/cholesterol-response, and human microglia loss-of-function papers. No annotation action changes were needed: TREM2 remains curated as a myeloid/microglial immunoreceptor for damage-associated lipid, apolipoprotein, apoptotic-cell, and amyloid-associated ligands, with plaque, synaptic, and broad inflammatory phenotypes retained as non-core context where appropriate.

📄 View Raw YAML

id: Q9NZC2
gene_symbol: TREM2
product_type: PROTEIN
status: COMPLETE
taxon:
  id: NCBITaxon:9606
  label: Homo sapiens
description: 'TREM2 is a type I plasma-membrane immunoglobulin-superfamily receptor expressed mainly by
  microglia and other myeloid-lineage cells. Together with the TYROBP/DAP12 adaptor, TREM2 converts ligand
  binding at the cell surface into intracellular signaling through kinases and PLCG2-linked pathways that
  support cell survival, calcium/ERK signaling, phagocytosis, chemotaxis, microglial activation, and lipid
  handling. TREM2 recognizes lipid-rich and damage-associated ligands, including aminophospholipids exposed
  on apoptotic cells, lipoprotein particles and apolipoproteins, and amyloid-beta-lipoprotein complexes.
  Disease-model annotations involving amyloid plaques, synapse pruning, neuroinflammation, osteoclast
  biology, and dendritic cell maturation are important contexts but are secondary to the core receptor-ligand
  signaling and phagocytic myeloid-cell functions.'
alternative_products:
- name: '1'
  id: Q9NZC2-1
- name: 2 (TREM-2V)
  id: Q9NZC2-2
  sequence_note: VSP_010792
- name: '3'
  id: Q9NZC2-3
  sequence_note: VSP_010793
existing_annotations:
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: is_active_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0004888
    label: transmembrane signaling receptor activity
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: enables
  review:
    summary: TREM2 transmembrane signaling receptor activity is a core plasma-membrane receptor 
      function coupled to TYROBP/DAP12 signaling.
    action: ACCEPT
    reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives 
      intracellular signaling through DAP12/TYROBP-linked kinase pathways.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0007165
    label: signal transduction
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: involved_in
  review:
    summary: signal transduction is a core downstream consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0045088
    label: regulation of innate immune response
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: involved_in
  review:
    summary: regulation of innate immune response is a core downstream consequence of TREM2 receptor
      signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0050850
    label: positive regulation of calcium-mediated signaling
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: involved_in
  review:
    summary: positive regulation of calcium-mediated signaling is a core downstream consequence of 
      TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0060100
    label: positive regulation of phagocytosis, engulfment
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: involved_in
  review:
    summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0070374
    label: positive regulation of ERK1 and ERK2 cascade
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: involved_in
  review:
    summary: positive regulation of ERK1 and ERK2 cascade is a core downstream consequence of TREM2 
      receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0120035
    label: regulation of plasma membrane bounded cell projection organization
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: involved_in
  review:
    summary: regulation of plasma membrane bounded cell projection organization is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1903980
    label: positive regulation of microglial cell activation
  evidence_type: IBA
  original_reference_id: GO_REF:0000033
  qualifier: involved_in
  review:
    summary: positive regulation of microglial cell activation is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0004888
    label: transmembrane signaling receptor activity
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: enables
  review:
    summary: TREM2 transmembrane signaling receptor activity is a core plasma-membrane receptor 
      function coupled to TYROBP/DAP12 signaling.
    action: ACCEPT
    reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives 
      intracellular signaling through DAP12/TYROBP-linked kinase pathways.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0005576
    label: extracellular region
  evidence_type: IEA
  original_reference_id: GO_REF:0000044
  qualifier: located_in
  review:
    summary: The extracellular-region annotation likely reflects secreted or shed TREM2 
      isoforms/fragments rather than the core membrane receptor.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because soluble TREM2 biology is relevant, but the primary TREM2 
      function is cell-surface receptor signaling.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0006909
    label: phagocytosis
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: involved_in
  review:
    summary: phagocytosis is part of TREM2-mediated microglial/myeloid phagocytic or lipid-response 
      biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0008289
    label: lipid binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: enables
  review:
    summary: TREM2 lipid binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: TREM2 ligand recognition includes lipid-rich particles and supports microglial lipid 
      handling.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: we identified a set of lipoprotein particles (including LDL) and 
        apolipoproteins (including CLU/APOJ and APOE) as ligands
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
- term:
    id: GO:0035176
    label: social behavior
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: involved_in
  review:
    summary: social behavior is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0048468
    label: cell development
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: involved_in
  review:
    summary: cell development is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0050850
    label: positive regulation of calcium-mediated signaling
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: involved_in
  review:
    summary: positive regulation of calcium-mediated signaling is a core downstream consequence of 
      TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0071396
    label: cellular response to lipid
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: involved_in
  review:
    summary: cellular response to lipid is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0097367
    label: carbohydrate derivative binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: enables
  review:
    summary: carbohydrate derivative binding is retained as non-core TREM2-associated biology 
      pending deeper reference-specific adjudication.
    action: KEEP_AS_NON_CORE
    reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic 
      microglial function set, but the local evidence was not sufficient to remove it.
- term:
    id: GO:1900226
    label: negative regulation of NLRP3 inflammasome complex assembly
  evidence_type: IEA
  original_reference_id: GO_REF:0000117
  qualifier: involved_in
  review:
    summary: negative regulation of NLRP3 inflammasome complex assembly is retained as non-core 
      TREM2-associated biology pending deeper reference-specific adjudication.
    action: KEEP_AS_NON_CORE
    reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic 
      microglial function set, but the local evidence was not sufficient to remove it.
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:11602640
  qualifier: enables
  review:
    summary: Generic protein binding annotation for TREM2.
    action: MARK_AS_OVER_ANNOTATED
    reason: The interaction may be real, but this term is too generic. Specific ligand-binding, 
      receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:28490631
  qualifier: enables
  review:
    summary: Generic protein binding annotation for TREM2.
    action: MARK_AS_OVER_ANNOTATED
    reason: The interaction may be real, but this term is too generic. Specific ligand-binding, 
      receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:29611543
  qualifier: enables
  review:
    summary: Generic protein binding annotation for TREM2.
    action: MARK_AS_OVER_ANNOTATED
    reason: The interaction may be real, but this term is too generic. Specific ligand-binding, 
      receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
- term:
    id: GO:0005515
    label: protein binding
  evidence_type: IPI
  original_reference_id: PMID:30341064
  qualifier: enables
  review:
    summary: Generic protein binding annotation for TREM2.
    action: MARK_AS_OVER_ANNOTATED
    reason: The interaction may be real, but this term is too generic. Specific ligand-binding, 
      receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
- term:
    id: GO:0001530
    label: lipopolysaccharide binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: enables
  review:
    summary: lipopolysaccharide binding is electronically inferred pathogen-associated ligand or 
      defense biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0001774
    label: microglial cell activation
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0002282
    label: microglial cell activation involved in immune response
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: microglial cell activation involved in immune response is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0002291
    label: T cell activation via T cell receptor contact with antigen bound to MHC molecule on 
      antigen presenting cell
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: T cell activation via T cell receptor contact with antigen bound to MHC molecule on 
      antigen presenting cell is a plausible downstream immune/inflammatory consequence of TREM2 
      signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0002862
    label: negative regulation of inflammatory response to antigenic stimulus
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of inflammatory response to antigenic stimulus is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0002931
    label: response to ischemia
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: response to ischemia is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0010507
    label: negative regulation of autophagy
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of autophagy is retained as context-specific TREM2-associated 
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0010628
    label: positive regulation of gene expression
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: positive regulation of gene expression is retained as context-specific TREM2-associated
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0010875
    label: positive regulation of cholesterol efflux
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of cholesterol efflux is part of TREM2-mediated microglial/myeloid 
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0010887
    label: negative regulation of cholesterol storage
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of cholesterol storage is part of TREM2-mediated microglial/myeloid
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0010891
    label: negative regulation of triglyceride storage
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of triglyceride storage is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0010983
    label: positive regulation of high-density lipoprotein particle clearance
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of high-density lipoprotein particle clearance is part of 
      TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0017154
    label: semaphorin receptor activity
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: enables
  review:
    summary: semaphorin receptor activity is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0030215
    label: semaphorin receptor binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: enables
  review:
    summary: semaphorin receptor binding is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0032006
    label: regulation of TOR signaling
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: regulation of TOR signaling is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0032008
    label: positive regulation of TOR signaling
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of TOR signaling is retained as context-specific TREM2-associated 
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0032497
    label: detection of lipopolysaccharide
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: detection of lipopolysaccharide is electronically inferred pathogen-associated ligand 
      or defense biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0032499
    label: detection of peptidoglycan
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: detection of peptidoglycan is electronically inferred pathogen-associated ligand or 
      defense biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0032675
    label: regulation of interleukin-6 production
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: regulation of interleukin-6 production is a plausible downstream immune/inflammatory 
      consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0032691
    label: negative regulation of interleukin-1 beta production
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of interleukin-1 beta production is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0032720
    label: negative regulation of tumor necrosis factor production
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of tumor necrosis factor production is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0032733
    label: positive regulation of interleukin-10 production
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of interleukin-10 production is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0034136
    label: negative regulation of toll-like receptor 2 signaling pathway
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of toll-like receptor 2 signaling pathway is a plausible downstream
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0034144
    label: negative regulation of toll-like receptor 4 signaling pathway
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of toll-like receptor 4 signaling pathway is a plausible downstream
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0034151
    label: regulation of toll-like receptor 6 signaling pathway
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: regulation of toll-like receptor 6 signaling pathway is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0034241
    label: positive regulation of macrophage fusion
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of macrophage fusion is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0034351
    label: negative regulation of glial cell apoptotic process
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of glial cell apoptotic process is retained as non-core 
      TREM2-associated biology pending deeper reference-specific adjudication.
    action: KEEP_AS_NON_CORE
    reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic 
      microglial function set, but the local evidence was not sufficient to remove it.
- term:
    id: GO:0038023
    label: signaling receptor activity
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: enables
  review:
    summary: TREM2 signaling receptor activity is a core plasma-membrane receptor function coupled 
      to TYROBP/DAP12 signaling.
    action: ACCEPT
    reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives 
      intracellular signaling through DAP12/TYROBP-linked kinase pathways.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0042742
    label: defense response to bacterium
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: defense response to bacterium is electronically inferred pathogen-associated ligand or 
      defense biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0042834
    label: peptidoglycan binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: enables
  review:
    summary: peptidoglycan binding is electronically inferred pathogen-associated ligand or defense 
      biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0043066
    label: negative regulation of apoptotic process
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of apoptotic process is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0043124
    label: negative regulation of canonical NF-kappaB signal transduction
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of canonical NF-kappaB signal transduction is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0043268
    label: positive regulation of potassium ion transport
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of potassium ion transport is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0043277
    label: apoptotic cell clearance
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: apoptotic cell clearance is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0045088
    label: regulation of innate immune response
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: regulation of innate immune response is a core downstream consequence of TREM2 receptor
      signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0045672
    label: positive regulation of osteoclast differentiation
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of osteoclast differentiation is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
    supported_by:
    - reference_id: PMID:12925681
      supporting_text: loss of function mutations in DAP12 and TREM2 result in an inefficient and 
        delayed differentiation of osteoclasts
- term:
    id: GO:0045728
    label: respiratory burst after phagocytosis
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: respiratory burst after phagocytosis is a plausible downstream immune/inflammatory 
      consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0045960
    label: positive regulation of complement activation, classical pathway
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of complement activation, classical pathway is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0048143
    label: astrocyte activation
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: astrocyte activation is a plausible downstream immune/inflammatory consequence of TREM2
      signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0048678
    label: response to axon injury
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: response to axon injury is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0050766
    label: positive regulation of phagocytosis
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of phagocytosis is part of TREM2-mediated microglial/myeloid 
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0050866
    label: negative regulation of cell activation
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of cell activation is a plausible downstream immune/inflammatory 
      consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0050921
    label: positive regulation of chemotaxis
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of chemotaxis is part of TREM2-mediated microglial/myeloid 
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0051897
    label: positive regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of phosphatidylinositol 3-kinase/protein kinase B signal 
      transduction is a core downstream consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0051898
    label: negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal 
      transduction is a core downstream consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0055088
    label: lipid homeostasis
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: lipid homeostasis is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0060075
    label: regulation of resting membrane potential
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: regulation of resting membrane potential is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0060100
    label: positive regulation of phagocytosis, engulfment
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0061518
    label: microglial cell proliferation
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: microglial cell proliferation is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0061889
    label: negative regulation of astrocyte activation
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of astrocyte activation is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0070345
    label: negative regulation of fat cell proliferation
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of fat cell proliferation is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0070374
    label: positive regulation of ERK1 and ERK2 cascade
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: positive regulation of ERK1 and ERK2 cascade is a core downstream consequence of TREM2 
      receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0070392
    label: detection of lipoteichoic acid
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: detection of lipoteichoic acid is electronically inferred pathogen-associated ligand or
      defense biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0070891
    label: lipoteichoic acid binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: enables
  review:
    summary: lipoteichoic acid binding is electronically inferred pathogen-associated ligand or 
      defense biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0071223
    label: cellular response to lipoteichoic acid
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: cellular response to lipoteichoic acid is electronically inferred pathogen-associated 
      ligand or defense biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0071224
    label: cellular response to peptidoglycan
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: cellular response to peptidoglycan is electronically inferred pathogen-associated 
      ligand or defense biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0071333
    label: cellular response to glucose stimulus
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: cellular response to glucose stimulus is retained as context-specific TREM2-associated 
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0071456
    label: cellular response to hypoxia
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: cellular response to hypoxia is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0071526
    label: semaphorin-plexin signaling pathway
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: semaphorin-plexin signaling pathway is retained as context-specific TREM2-associated 
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0071640
    label: regulation of macrophage inflammatory protein 1 alpha production
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: regulation of macrophage inflammatory protein 1 alpha production is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0097062
    label: dendritic spine maintenance
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: dendritic spine maintenance is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0097242
    label: amyloid-beta clearance
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0098657
    label: import into cell
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: import into cell is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0110089
    label: regulation of hippocampal neuron apoptotic process
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: regulation of hippocampal neuron apoptotic process is Alzheimer-relevant TREM2 biology,
      but it is a context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0120035
    label: regulation of plasma membrane bounded cell projection organization
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: regulation of plasma membrane bounded cell projection organization is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0150062
    label: complement-mediated synapse pruning
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: complement-mediated synapse pruning is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0150076
    label: neuroinflammatory response
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: neuroinflammatory response is a plausible downstream immune/inflammatory consequence of
      TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0150078
    label: positive regulation of neuroinflammatory response
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of neuroinflammatory response is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0150079
    label: negative regulation of neuroinflammatory response
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: negative regulation of neuroinflammatory response is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0150094
    label: amyloid-beta clearance by cellular catabolic process
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: amyloid-beta clearance by cellular catabolic process is Alzheimer-relevant TREM2 
      biology, but it is a context-specific substrate/phenotype rather than the broad core receptor 
      function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1900015
    label: regulation of cytokine production involved in inflammatory response
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: regulation of cytokine production involved in inflammatory response is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1900016
    label: negative regulation of cytokine production involved in inflammatory response
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of cytokine production involved in inflammatory response is a 
      plausible downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1900223
    label: positive regulation of amyloid-beta clearance
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: positive regulation of amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but 
      it is a context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1901224
    label: positive regulation of non-canonical NF-kappaB signal transduction
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of non-canonical NF-kappaB signal transduction is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1901800
    label: positive regulation of proteasomal protein catabolic process
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of proteasomal protein catabolic process is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1902533
    label: positive regulation of intracellular signal transduction
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of intracellular signal transduction is a core downstream 
      consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:1903376
    label: regulation of oxidative stress-induced neuron intrinsic apoptotic signaling pathway
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: regulation of oxidative stress-induced neuron intrinsic apoptotic signaling pathway is 
      Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than
      the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1903753
    label: negative regulation of p38MAPK cascade
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of p38MAPK cascade is a plausible downstream immune/inflammatory 
      consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1903980
    label: positive regulation of microglial cell activation
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: positive regulation of microglial cell activation is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1904093
    label: negative regulation of autophagic cell death
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of autophagic cell death is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1904141
    label: positive regulation of microglial cell migration
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: positive regulation of microglial cell migration is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1904646
    label: cellular response to amyloid-beta
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: cellular response to amyloid-beta is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1904951
    label: positive regulation of establishment of protein localization
  evidence_type: IEA
  original_reference_id: GO_REF:0000120
  qualifier: involved_in
  review:
    summary: positive regulation of establishment of protein localization is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1905291
    label: positive regulation of CAMKK-AMPK signaling cascade
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of CAMKK-AMPK signaling cascade is a core downstream consequence of
      TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:1905581
    label: positive regulation of low-density lipoprotein particle clearance
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of low-density lipoprotein particle clearance is part of 
      TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:1905805
    label: excitatory synapse pruning
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: excitatory synapse pruning is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1905907
    label: negative regulation of amyloid fibril formation
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: negative regulation of amyloid fibril formation is Alzheimer-relevant TREM2 biology, 
      but it is a context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1990782
    label: protein tyrosine kinase binding
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: enables
  review:
    summary: protein tyrosine kinase binding is a specific interaction context for TREM2 but not the
      main molecular-function term.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the interaction may be real, while more informative core 
      terms describe receptor activity and lipid/lipoprotein/aminophospholipid ligand binding.
- term:
    id: GO:2001171
    label: positive regulation of ATP biosynthetic process
  evidence_type: IEA
  original_reference_id: GO_REF:0000107
  qualifier: involved_in
  review:
    summary: positive regulation of ATP biosynthetic process is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1902533
    label: positive regulation of intracellular signal transduction
  evidence_type: IDA
  original_reference_id: PMID:32514138
  qualifier: acts_upstream_of_or_within
  review:
    summary: positive regulation of intracellular signal transduction is a core downstream 
      consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: EXP
  original_reference_id: PMID:25615530
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: EXP
  original_reference_id: PMID:28768830
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: EXP
  original_reference_id: PMID:28923481
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0010468
    label: regulation of gene expression
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: regulation of gene expression is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0032675
    label: regulation of interleukin-6 production
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: regulation of interleukin-6 production is a plausible downstream immune/inflammatory 
      consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0043268
    label: positive regulation of potassium ion transport
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: positive regulation of potassium ion transport is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0071640
    label: regulation of macrophage inflammatory protein 1 alpha production
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: regulation of macrophage inflammatory protein 1 alpha production is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1902533
    label: positive regulation of intracellular signal transduction
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: positive regulation of intracellular signal transduction is a core downstream 
      consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:1902533
    label: positive regulation of intracellular signal transduction
  evidence_type: IDA
  original_reference_id: PMID:31902528
  qualifier: involved_in
  review:
    summary: positive regulation of intracellular signal transduction is a core downstream 
      consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0002116
    label: semaphorin receptor complex
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: part_of
  review:
    summary: semaphorin receptor complex is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0002291
    label: T cell activation via T cell receptor contact with antigen bound to MHC molecule on 
      antigen presenting cell
  evidence_type: IDA
  original_reference_id: PMID:16715077
  qualifier: involved_in
  review:
    summary: T cell activation via T cell receptor contact with antigen bound to MHC molecule on 
      antigen presenting cell is a plausible downstream immune/inflammatory consequence of TREM2 
      signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IDA
  original_reference_id: PMID:16715077
  qualifier: is_active_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0017154
    label: semaphorin receptor activity
  evidence_type: IDA
  original_reference_id: PMID:16715077
  qualifier: contributes_to
  review:
    summary: semaphorin receptor activity is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0071526
    label: semaphorin-plexin signaling pathway
  evidence_type: IDA
  original_reference_id: PMID:16715077
  qualifier: involved_in
  review:
    summary: semaphorin-plexin signaling pathway is retained as context-specific TREM2-associated 
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0004888
    label: transmembrane signaling receptor activity
  evidence_type: IDA
  original_reference_id: PMID:29518356
  qualifier: enables
  review:
    summary: TREM2 transmembrane signaling receptor activity is a core plasma-membrane receptor 
      function coupled to TYROBP/DAP12 signaling.
    action: ACCEPT
    reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives 
      intracellular signaling through DAP12/TYROBP-linked kinase pathways.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0004888
    label: transmembrane signaling receptor activity
  evidence_type: IDA
  original_reference_id: PMID:30333625
  qualifier: enables
  review:
    summary: TREM2 transmembrane signaling receptor activity is a core plasma-membrane receptor 
      function coupled to TYROBP/DAP12 signaling.
    action: ACCEPT
    reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives 
      intracellular signaling through DAP12/TYROBP-linked kinase pathways.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IDA
  original_reference_id: PMID:24078628
  qualifier: is_active_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IDA
  original_reference_id: PMID:27477018
  qualifier: is_active_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0071402
    label: cellular response to lipoprotein particle stimulus
  evidence_type: IDA
  original_reference_id: PMID:27477018
  qualifier: involved_in
  review:
    summary: cellular response to lipoprotein particle stimulus is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0097242
    label: amyloid-beta clearance
  evidence_type: IDA
  original_reference_id: PMID:27477018
  qualifier: involved_in
  review:
    summary: amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0008013
    label: beta-catenin binding
  evidence_type: IPI
  original_reference_id: PMID:30683932
  qualifier: enables
  review:
    summary: beta-catenin binding is a specific interaction context for TREM2 but not the main 
      molecular-function term.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the interaction may be real, while more informative core 
      terms describe receptor activity and lipid/lipoprotein/aminophospholipid ligand binding.
- term:
    id: GO:0051898
    label: negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
  evidence_type: IMP
  original_reference_id: PMID:30683932
  qualifier: involved_in
  review:
    summary: negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal 
      transduction is a core downstream consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0050921
    label: positive regulation of chemotaxis
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of chemotaxis is part of TREM2-mediated microglial/myeloid 
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0120035
    label: regulation of plasma membrane bounded cell projection organization
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: regulation of plasma membrane bounded cell projection organization is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1904141
    label: positive regulation of microglial cell migration
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of microglial cell migration is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0010507
    label: negative regulation of autophagy
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of autophagy is retained as context-specific TREM2-associated 
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0032006
    label: regulation of TOR signaling
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: regulation of TOR signaling is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0032008
    label: positive regulation of TOR signaling
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of TOR signaling is retained as context-specific TREM2-associated 
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1904093
    label: negative regulation of autophagic cell death
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of autophagic cell death is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1905291
    label: positive regulation of CAMKK-AMPK signaling cascade
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of CAMKK-AMPK signaling cascade is a core downstream consequence of
      TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:2001171
    label: positive regulation of ATP biosynthetic process
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of ATP biosynthetic process is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1900223
    label: positive regulation of amyloid-beta clearance
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but 
      it is a context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0006910
    label: phagocytosis, recognition
  evidence_type: IMP
  original_reference_id: PMID:32514138
  qualifier: involved_in
  review:
    summary: phagocytosis, recognition is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0010628
    label: positive regulation of gene expression
  evidence_type: IMP
  original_reference_id: PMID:32514138
  qualifier: acts_upstream_of_or_within
  review:
    summary: positive regulation of gene expression is retained as context-specific TREM2-associated
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0019216
    label: regulation of lipid metabolic process
  evidence_type: IMP
  original_reference_id: PMID:32514138
  qualifier: acts_upstream_of
  review:
    summary: regulation of lipid metabolic process is part of TREM2-mediated microglial/myeloid 
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0038023
    label: signaling receptor activity
  evidence_type: IMP
  original_reference_id: PMID:32514138
  qualifier: enables
  review:
    summary: TREM2 signaling receptor activity is a core plasma-membrane receptor function coupled 
      to TYROBP/DAP12 signaling.
    action: ACCEPT
    reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives 
      intracellular signaling through DAP12/TYROBP-linked kinase pathways.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0048678
    label: response to axon injury
  evidence_type: IMP
  original_reference_id: PMID:32514138
  qualifier: involved_in
  review:
    summary: response to axon injury is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0060100
    label: positive regulation of phagocytosis, engulfment
  evidence_type: IMP
  original_reference_id: PMID:32514138
  qualifier: acts_upstream_of
  review:
    summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0071396
    label: cellular response to lipid
  evidence_type: IMP
  original_reference_id: PMID:32514138
  qualifier: involved_in
  review:
    summary: cellular response to lipid is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0150079
    label: negative regulation of neuroinflammatory response
  evidence_type: IMP
  original_reference_id: PMID:32514138
  qualifier: involved_in
  review:
    summary: negative regulation of neuroinflammatory response is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1900226
    label: negative regulation of NLRP3 inflammasome complex assembly
  evidence_type: IMP
  original_reference_id: PMID:32514138
  qualifier: acts_upstream_of_or_within
  review:
    summary: negative regulation of NLRP3 inflammasome complex assembly is retained as non-core 
      TREM2-associated biology pending deeper reference-specific adjudication.
    action: KEEP_AS_NON_CORE
    reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic 
      microglial function set, but the local evidence was not sufficient to remove it.
- term:
    id: GO:0038160
    label: CXCL12-activated CXCR4 signaling pathway
  evidence_type: IMP
  original_reference_id: PMID:33097708
  qualifier: acts_upstream_of_or_within_positive_effect
  review:
    summary: CXCL12-activated CXCR4 signaling pathway is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0050829
    label: defense response to Gram-negative bacterium
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: acts_upstream_of_or_within_positive_effect
  review:
    summary: defense response to Gram-negative bacterium is electronically inferred 
      pathogen-associated ligand or defense biology.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because TREM-family receptors participate in myeloid immune sensing, 
      but the cached human TREM2 evidence reviewed here emphasizes lipids, apoptotic cells, and 
      microglial phagocytic ligands.
- term:
    id: GO:0070269
    label: pyroptotic inflammatory response
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: acts_upstream_of_or_within_negative_effect
  review:
    summary: pyroptotic inflammatory response is a plausible downstream immune/inflammatory 
      consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1900016
    label: negative regulation of cytokine production involved in inflammatory response
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of cytokine production involved in inflammatory response is a 
      plausible downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1900226
    label: negative regulation of NLRP3 inflammasome complex assembly
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: acts_upstream_of_or_within
  review:
    summary: negative regulation of NLRP3 inflammasome complex assembly is retained as non-core 
      TREM2-associated biology pending deeper reference-specific adjudication.
    action: KEEP_AS_NON_CORE
    reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic 
      microglial function set, but the local evidence was not sufficient to remove it.
- term:
    id: GO:0150079
    label: negative regulation of neuroinflammatory response
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of neuroinflammatory response is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1903753
    label: negative regulation of p38MAPK cascade
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of p38MAPK cascade is a plausible downstream immune/inflammatory 
      consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0034144
    label: negative regulation of toll-like receptor 4 signaling pathway
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of toll-like receptor 4 signaling pathway is a plausible downstream
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0050866
    label: negative regulation of cell activation
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of cell activation is a plausible downstream immune/inflammatory 
      consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0051897
    label: positive regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of phosphatidylinositol 3-kinase/protein kinase B signal 
      transduction is a core downstream consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0061889
    label: negative regulation of astrocyte activation
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of astrocyte activation is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0110089
    label: regulation of hippocampal neuron apoptotic process
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: regulation of hippocampal neuron apoptotic process is Alzheimer-relevant TREM2 biology,
      but it is a context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1903376
    label: regulation of oxidative stress-induced neuron intrinsic apoptotic signaling pathway
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: regulation of oxidative stress-induced neuron intrinsic apoptotic signaling pathway is 
      Alzheimer-relevant TREM2 biology, but it is a context-specific substrate/phenotype rather than
      the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0001786
    label: phosphatidylserine binding
  evidence_type: IDA
  original_reference_id: PMID:31101881
  qualifier: enables
  review:
    summary: TREM2 phosphatidylserine binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function 
      for apoptotic-cell and debris responses.
    supported_by:
    - reference_id: PMID:31101881
      supporting_text: TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is
        mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
- term:
    id: GO:0002282
    label: microglial cell activation involved in immune response
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: microglial cell activation involved in immune response is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0007613
    label: memory
  evidence_type: IDA
  original_reference_id: PMID:31462511
  qualifier: acts_upstream_of_positive_effect
  review:
    summary: memory is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0008429
    label: phosphatidylethanolamine binding
  evidence_type: IDA
  original_reference_id: PMID:31101881
  qualifier: enables
  review:
    summary: TREM2 phosphatidylethanolamine binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function 
      for apoptotic-cell and debris responses.
    supported_by:
    - reference_id: PMID:31101881
      supporting_text: TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is
        mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
- term:
    id: GO:0034136
    label: negative regulation of toll-like receptor 2 signaling pathway
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of toll-like receptor 2 signaling pathway is a plausible downstream
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0034151
    label: regulation of toll-like receptor 6 signaling pathway
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: regulation of toll-like receptor 6 signaling pathway is a plausible downstream 
      immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0097062
    label: dendritic spine maintenance
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: dendritic spine maintenance is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0150094
    label: amyloid-beta clearance by cellular catabolic process
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: amyloid-beta clearance by cellular catabolic process is Alzheimer-relevant TREM2 
      biology, but it is a context-specific substrate/phenotype rather than the broad core receptor 
      function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0150094
    label: amyloid-beta clearance by cellular catabolic process
  evidence_type: IMP
  original_reference_id: PMID:33097708
  qualifier: involved_in
  review:
    summary: amyloid-beta clearance by cellular catabolic process is Alzheimer-relevant TREM2 
      biology, but it is a context-specific substrate/phenotype rather than the broad core receptor 
      function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0001774
    label: microglial cell activation
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0002862
    label: negative regulation of inflammatory response to antigenic stimulus
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of inflammatory response to antigenic stimulus is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0007613
    label: memory
  evidence_type: IMP
  original_reference_id: PMID:29518357
  qualifier: acts_upstream_of_positive_effect
  review:
    summary: memory is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0034351
    label: negative regulation of glial cell apoptotic process
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of glial cell apoptotic process is retained as non-core 
      TREM2-associated biology pending deeper reference-specific adjudication.
    action: KEEP_AS_NON_CORE
    reason: The annotation is not part of the primary receptor-ligand signaling and phagocytic 
      microglial function set, but the local evidence was not sufficient to remove it.
- term:
    id: GO:0043124
    label: negative regulation of canonical NF-kappaB signal transduction
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of canonical NF-kappaB signal transduction is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0050766
    label: positive regulation of phagocytosis
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of phagocytosis is part of TREM2-mediated microglial/myeloid 
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0050766
    label: positive regulation of phagocytosis
  evidence_type: IMP
  original_reference_id: PMID:29518357
  qualifier: involved_in
  review:
    summary: positive regulation of phagocytosis is part of TREM2-mediated microglial/myeloid 
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0051898
    label: negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal transduction
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of phosphatidylinositol 3-kinase/protein kinase B signal 
      transduction is a core downstream consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:1900015
    label: regulation of cytokine production involved in inflammatory response
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: regulation of cytokine production involved in inflammatory response is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0060100
    label: positive regulation of phagocytosis, engulfment
  evidence_type: IMP
  original_reference_id: PMID:33097708
  qualifier: involved_in
  review:
    summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1902227
    label: negative regulation of macrophage colony-stimulating factor signaling pathway
  evidence_type: IMP
  original_reference_id: PMID:33097708
  qualifier: involved_in
  review:
    summary: negative regulation of macrophage colony-stimulating factor signaling pathway is a 
      plausible downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:1904141
    label: positive regulation of microglial cell migration
  evidence_type: IMP
  original_reference_id: PMID:33097708
  qualifier: involved_in
  review:
    summary: positive regulation of microglial cell migration is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1905808
    label: positive regulation of synapse pruning
  evidence_type: IMP
  original_reference_id: PMID:33097708
  qualifier: involved_in
  review:
    summary: positive regulation of synapse pruning is Alzheimer-relevant TREM2 biology, but it is a
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0035176
    label: social behavior
  evidence_type: IMP
  original_reference_id: PMID:29752066
  qualifier: acts_upstream_of_positive_effect
  review:
    summary: social behavior is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0045088
    label: regulation of innate immune response
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: regulation of innate immune response is a core downstream consequence of TREM2 receptor
      signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0045728
    label: respiratory burst after phagocytosis
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: respiratory burst after phagocytosis is a plausible downstream immune/inflammatory 
      consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0045960
    label: positive regulation of complement activation, classical pathway
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of complement activation, classical pathway is a plausible 
      downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0061518
    label: microglial cell proliferation
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: microglial cell proliferation is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0150062
    label: complement-mediated synapse pruning
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: complement-mediated synapse pruning is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0060100
    label: positive regulation of phagocytosis, engulfment
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1905805
    label: excitatory synapse pruning
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: excitatory synapse pruning is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0048678
    label: response to axon injury
  evidence_type: IMP
  original_reference_id: PMID:31235932
  qualifier: involved_in
  review:
    summary: response to axon injury is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1900223
    label: positive regulation of amyloid-beta clearance
  evidence_type: IMP
  original_reference_id: PMID:31235932
  qualifier: involved_in
  review:
    summary: positive regulation of amyloid-beta clearance is Alzheimer-relevant TREM2 biology, but 
      it is a context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0001786
    label: phosphatidylserine binding
  evidence_type: IDA
  original_reference_id: PMID:31902528
  qualifier: enables
  review:
    summary: TREM2 phosphatidylserine binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function 
      for apoptotic-cell and debris responses.
    supported_by:
    - reference_id: PMID:31101881
      supporting_text: TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is
        mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
- term:
    id: GO:0010875
    label: positive regulation of cholesterol efflux
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of cholesterol efflux is part of TREM2-mediated microglial/myeloid 
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0010887
    label: negative regulation of cholesterol storage
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of cholesterol storage is part of TREM2-mediated microglial/myeloid
      phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0010891
    label: negative regulation of triglyceride storage
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of triglyceride storage is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0048678
    label: response to axon injury
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: response to axon injury is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0120146
    label: sulfatide binding
  evidence_type: IDA
  original_reference_id: PMID:31902528
  qualifier: enables
  review:
    summary: TREM2 sulfatide binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function 
      for apoptotic-cell and debris responses.
    supported_by:
    - reference_id: PMID:31101881
      supporting_text: TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is
        mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
- term:
    id: GO:0140052
    label: cellular response to oxidised low-density lipoprotein particle stimulus
  evidence_type: IDA
  original_reference_id: PMID:31902528
  qualifier: involved_in
  review:
    summary: cellular response to oxidised low-density lipoprotein particle stimulus is part of 
      TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:1903980
    label: positive regulation of microglial cell activation
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of microglial cell activation is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1904951
    label: positive regulation of establishment of protein localization
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of establishment of protein localization is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1905907
    label: negative regulation of amyloid fibril formation
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of amyloid fibril formation is Alzheimer-relevant TREM2 biology, 
      but it is a context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0019209
    label: kinase activator activity
  evidence_type: IMP
  original_reference_id: PMID:31902528
  qualifier: enables
  review:
    summary: kinase activator activity is a core downstream consequence of TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0010983
    label: positive regulation of high-density lipoprotein particle clearance
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of high-density lipoprotein particle clearance is part of 
      TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0055088
    label: lipid homeostasis
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: lipid homeostasis is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0070345
    label: negative regulation of fat cell proliferation
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: negative regulation of fat cell proliferation is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1905581
    label: positive regulation of low-density lipoprotein particle clearance
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of low-density lipoprotein particle clearance is part of 
      TREM2-mediated microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 ligand sensing and phagocytic challenge regulate microglial lipid/cholesterol 
      handling.
    supported_by:
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
    - reference_id: PMID:31902528
      supporting_text: Our studies identify TREM2 as a key transcriptional regulator of cholesterol 
        transport and metabolism under conditions of chronic myelin phagocytic activity
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0044853
    label: plasma membrane raft
  evidence_type: IDA
  original_reference_id: PMID:31413141
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane raft is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0001774
    label: microglial cell activation
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: microglial cell activation is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0048143
    label: astrocyte activation
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: astrocyte activation is a plausible downstream immune/inflammatory consequence of TREM2
      signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IMP
  original_reference_id: PMID:24990881
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IMP
  original_reference_id: PMID:27589997
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IMP
  original_reference_id: PMID:28855300
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0060100
    label: positive regulation of phagocytosis, engulfment
  evidence_type: IMP
  original_reference_id: PMID:28855300
  qualifier: involved_in
  review:
    summary: positive regulation of phagocytosis, engulfment is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1901076
    label: positive regulation of engulfment of apoptotic cell
  evidence_type: IMP
  original_reference_id: PMID:24990881
  qualifier: involved_in
  review:
    summary: positive regulation of engulfment of apoptotic cell is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1903980
    label: positive regulation of microglial cell activation
  evidence_type: IMP
  original_reference_id: PMID:24990881
  qualifier: involved_in
  review:
    summary: positive regulation of microglial cell activation is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0030316
    label: osteoclast differentiation
  evidence_type: IMP
  original_reference_id: PMID:12925681
  qualifier: involved_in
  review:
    summary: osteoclast differentiation is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
    supported_by:
    - reference_id: PMID:12925681
      supporting_text: loss of function mutations in DAP12 and TREM2 result in an inefficient and 
        delayed differentiation of osteoclasts
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IMP
  original_reference_id: PMID:28855301
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0050714
    label: positive regulation of protein secretion
  evidence_type: IMP
  original_reference_id: PMID:27044754
  qualifier: involved_in
  review:
    summary: positive regulation of protein secretion is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0043277
    label: apoptotic cell clearance
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: apoptotic cell clearance is part of TREM2-mediated microglial/myeloid phagocytic or 
      lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:0008289
    label: lipid binding
  evidence_type: TAS
  original_reference_id: PMID:29518356
  qualifier: enables
  review:
    summary: TREM2 lipid binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: TREM2 ligand recognition includes lipid-rich particles and supports microglial lipid 
      handling.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: we identified a set of lipoprotein particles (including LDL) and 
        apolipoproteins (including CLU/APOJ and APOE) as ligands
    - reference_id: PMID:31902528
      supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
- term:
    id: GO:0038023
    label: signaling receptor activity
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: enables
  review:
    summary: TREM2 signaling receptor activity is a core plasma-membrane receptor function coupled 
      to TYROBP/DAP12 signaling.
    action: ACCEPT
    reason: TREM2 is a myeloid/microglial cell-surface receptor whose ligand engagement drives 
      intracellular signaling through DAP12/TYROBP-linked kinase pathways.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0060075
    label: regulation of resting membrane potential
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: regulation of resting membrane potential is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0120035
    label: regulation of plasma membrane bounded cell projection organization
  evidence_type: IGI
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: regulation of plasma membrane bounded cell projection organization is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1901800
    label: positive regulation of proteasomal protein catabolic process
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: positive regulation of proteasomal protein catabolic process is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1904141
    label: positive regulation of microglial cell migration
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: positive regulation of microglial cell migration is part of TREM2-mediated 
      microglial/myeloid phagocytic or lipid-response biology.
    action: ACCEPT
    reason: TREM2 is a core microglial/myeloid phagocytic receptor that supports engulfment, 
      activation, and migration responses.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1904646
    label: cellular response to amyloid-beta
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: involved_in
  review:
    summary: cellular response to amyloid-beta is Alzheimer-relevant TREM2 biology, but it is a 
      context-specific substrate/phenotype rather than the broad core receptor function.
    action: KEEP_AS_NON_CORE
    reason: Retain amyloid and synapse-pruning annotations as non-core contexts of TREM2 microglial 
      function; do not treat them as disease-progression hypotheses in the core function.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
    - reference_id: PMID:33097708
      supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
        substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
- term:
    id: GO:1990782
    label: protein tyrosine kinase binding
  evidence_type: ISS
  original_reference_id: PMID:29518356
  qualifier: enables
  review:
    summary: protein tyrosine kinase binding is a specific interaction context for TREM2 but not the
      main molecular-function term.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the interaction may be real, while more informative core 
      terms describe receptor activity and lipid/lipoprotein/aminophospholipid ligand binding.
- term:
    id: GO:0001540
    label: amyloid-beta binding
  evidence_type: IPI
  original_reference_id: PMID:29518356
  qualifier: enables
  review:
    summary: TREM2 amyloid-beta binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: Amyloid-beta binding is disease-relevant but mechanistically tied to TREM2 ligand uptake
      through lipoprotein-associated complexes; retain as part of ligand recognition.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: β-amyloid (Aβ) binds to lipoproteins and this complex is efficiently taken up
        by microglia in a TREM2-dependent fashion.
- term:
    id: GO:0044877
    label: protein-containing complex binding
  evidence_type: IPI
  original_reference_id: PMID:29518356
  qualifier: enables
  review:
    summary: Generic protein-containing complex binding annotation for TREM2.
    action: MARK_AS_OVER_ANNOTATED
    reason: The interaction may be real, but this term is too generic. Specific ligand-binding, 
      receptor-activity, TYROBP/DAP12 signaling, or phagocytosis annotations are more informative.
- term:
    id: GO:0010628
    label: positive regulation of gene expression
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of gene expression is retained as context-specific TREM2-associated
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0098657
    label: import into cell
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: import into cell is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0008035
    label: high-density lipoprotein particle binding
  evidence_type: IDA
  original_reference_id: PMID:27477018
  qualifier: enables
  review:
    summary: TREM2 high-density lipoprotein particle binding is part of its ligand-sensing receptor 
      biology.
    action: ACCEPT
    reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by 
      microglia or receptor-expressing cells.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: we identified a set of lipoprotein particles (including LDL) and 
        apolipoproteins (including CLU/APOJ and APOE) as ligands
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0030169
    label: low-density lipoprotein particle binding
  evidence_type: IDA
  original_reference_id: PMID:27477018
  qualifier: enables
  review:
    summary: TREM2 low-density lipoprotein particle binding is part of its ligand-sensing receptor 
      biology.
    action: ACCEPT
    reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by 
      microglia or receptor-expressing cells.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: we identified a set of lipoprotein particles (including LDL) and 
        apolipoproteins (including CLU/APOJ and APOE) as ligands
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0034185
    label: apolipoprotein binding
  evidence_type: IPI
  original_reference_id: PMID:27477018
  qualifier: enables
  review:
    summary: TREM2 apolipoprotein binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by 
      microglia or receptor-expressing cells.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: we identified a set of lipoprotein particles (including LDL) and 
        apolipoproteins (including CLU/APOJ and APOE) as ligands
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0034186
    label: apolipoprotein A-I binding
  evidence_type: IPI
  original_reference_id: PMID:27477018
  qualifier: enables
  review:
    summary: TREM2 apolipoprotein A-I binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by 
      microglia or receptor-expressing cells.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: we identified a set of lipoprotein particles (including LDL) and 
        apolipoproteins (including CLU/APOJ and APOE) as ligands
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0034189
    label: very-low-density lipoprotein particle binding
  evidence_type: IDA
  original_reference_id: PMID:27477018
  qualifier: enables
  review:
    summary: TREM2 very-low-density lipoprotein particle binding is part of its ligand-sensing 
      receptor biology.
    action: ACCEPT
    reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by 
      microglia or receptor-expressing cells.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: we identified a set of lipoprotein particles (including LDL) and 
        apolipoproteins (including CLU/APOJ and APOE) as ligands
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0071813
    label: lipoprotein particle binding
  evidence_type: IDA
  original_reference_id: PMID:27477018
  qualifier: enables
  review:
    summary: TREM2 lipoprotein particle binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: TREM2 recognizes apolipoproteins and lipoprotein particles and promotes their uptake by 
      microglia or receptor-expressing cells.
    supported_by:
    - reference_id: PMID:27477018
      supporting_text: we identified a set of lipoprotein particles (including LDL) and 
        apolipoproteins (including CLU/APOJ and APOE) as ligands
    - reference_id: PMID:27477018
      supporting_text: Overexpression of wild-type TREM2 was sufficient to enhance uptake of LDL, 
        CLU, and APOE in heterologous cells
- term:
    id: GO:0034241
    label: positive regulation of macrophage fusion
  evidence_type: ISS
  original_reference_id: GO_REF:0000024
  qualifier: involved_in
  review:
    summary: positive regulation of macrophage fusion is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0030316
    label: osteoclast differentiation
  evidence_type: IMP
  original_reference_id: PMID:21841309
  qualifier: acts_upstream_of_or_within
  review:
    summary: osteoclast differentiation is retained as context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
    supported_by:
    - reference_id: PMID:12925681
      supporting_text: loss of function mutations in DAP12 and TREM2 result in an inefficient and 
        delayed differentiation of osteoclasts
- term:
    id: GO:0005543
    label: phospholipid binding
  evidence_type: IDA
  original_reference_id: PMID:27995897
  qualifier: enables
  review:
    summary: TREM2 phospholipid binding is part of its ligand-sensing receptor biology.
    action: ACCEPT
    reason: Aminophospholipid and anionic lipid recognition is a core TREM2 ligand-sensing function 
      for apoptotic-cell and debris responses.
    supported_by:
    - reference_id: PMID:31101881
      supporting_text: TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is
        mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: IDA
  original_reference_id: PMID:27995897
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0070374
    label: positive regulation of ERK1 and ERK2 cascade
  evidence_type: IMP
  original_reference_id: PMID:27995897
  qualifier: involved_in
  review:
    summary: positive regulation of ERK1 and ERK2 cascade is a core downstream consequence of TREM2 
      receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0002588
    label: positive regulation of antigen processing and presentation of peptide antigen via MHC 
      class II
  evidence_type: IDA
  original_reference_id: PMID:11602640
  qualifier: involved_in
  review:
    summary: positive regulation of antigen processing and presentation of peptide antigen via MHC 
      class II is a plausible downstream immune/inflammatory consequence of TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
- term:
    id: GO:0016020
    label: membrane
  evidence_type: IDA
  original_reference_id: PMID:11602640
  qualifier: located_in
  review:
    summary: TREM2 localization to membrane is consistent with its type I cell-surface receptor 
      role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0050850
    label: positive regulation of calcium-mediated signaling
  evidence_type: IDA
  original_reference_id: PMID:11602640
  qualifier: involved_in
  review:
    summary: positive regulation of calcium-mediated signaling is a core downstream consequence of 
      TREM2 receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0070374
    label: positive regulation of ERK1 and ERK2 cascade
  evidence_type: IDA
  original_reference_id: PMID:11602640
  qualifier: involved_in
  review:
    summary: positive regulation of ERK1 and ERK2 cascade is a core downstream consequence of TREM2 
      receptor signaling.
    action: ACCEPT
    reason: TREM2 signals through DAP12/TYROBP and PLCG2-linked kinase pathways that support 
      calcium/ERK/PI3K-related activation and intracellular signaling.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
        maturation, and DC survival through activation of protein tyrosine kinases and extracellular
        signal-regulated kinase.
    - reference_id: PMID:32514138
      supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, 
        processing of neuronal debris, and lipid metabolism.
- term:
    id: GO:0097028
    label: dendritic cell differentiation
  evidence_type: IDA
  original_reference_id: PMID:11602640
  qualifier: involved_in
  review:
    summary: dendritic cell differentiation is retained as context-specific TREM2-associated 
      biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0097110
    label: scaffold protein binding
  evidence_type: IPI
  original_reference_id: PMID:11602640
  qualifier: enables
  review:
    summary: scaffold protein binding is a specific interaction context for TREM2 but not the main 
      molecular-function term.
    action: KEEP_AS_NON_CORE
    reason: Retain as non-core because the interaction may be real, while more informative core 
      terms describe receptor activity and lipid/lipoprotein/aminophospholipid ligand binding.
- term:
    id: GO:1903078
    label: positive regulation of protein localization to plasma membrane
  evidence_type: IDA
  original_reference_id: PMID:11602640
  qualifier: involved_in
  review:
    summary: positive regulation of protein localization to plasma membrane is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:1903082
    label: positive regulation of C-C chemokine receptor CCR7 signaling pathway
  evidence_type: IDA
  original_reference_id: PMID:11602640
  qualifier: involved_in
  review:
    summary: positive regulation of C-C chemokine receptor CCR7 signaling pathway is retained as 
      context-specific TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:2000350
    label: positive regulation of CD40 signaling pathway
  evidence_type: IDA
  original_reference_id: PMID:11602640
  qualifier: involved_in
  review:
    summary: positive regulation of CD40 signaling pathway is retained as context-specific 
      TREM2-associated biology.
    action: KEEP_AS_NON_CORE
    reason: This annotation reflects a cell-type-specific or downstream phenotype rather than the 
      core TREM2 receptor-ligand signaling mechanism.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-210289
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-210300
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-2395412
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-2395801
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-2396594
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-2424480
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-2424482
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-2424484
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-2424486
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-2424487
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-416725
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0005886
    label: plasma membrane
  evidence_type: TAS
  original_reference_id: Reactome:R-HSA-5696358
  qualifier: located_in
  review:
    summary: TREM2 localization to plasma membrane is consistent with its type I cell-surface 
      receptor role.
    action: ACCEPT
    reason: Cell-surface/plasma-membrane localization is essential for ligand binding and 
      DAP12/TYROBP-mediated signal transduction.
    supported_by:
    - reference_id: PMID:11602640
      supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
        associated with DAP12.
    - reference_id: PMID:24990881
      supporting_text: missense mutations associated with FTD and FTD-like syndrome reduce TREM2 
        maturation, abolish shedding by ADAM proteases, and impair the phagocytic activity of 
        TREM2-expressing cells.
    - reference_id: PMID:27995897
      supporting_text: Nasu-Hakola mutations impact protein stability and decrease folded TREM2 
        surface expression, whereas Alzheimer's risk variants impact binding to a TREM2 ligand.
- term:
    id: GO:0006959
    label: humoral immune response
  evidence_type: TAS
  original_reference_id: PMID:10799849
  qualifier: involved_in
  review:
    summary: humoral immune response is a plausible downstream immune/inflammatory consequence of 
      TREM2 signaling.
    action: KEEP_AS_NON_CORE
    reason: TREM2 modulates myeloid and microglial immune responses, but specific 
      cytokine/TLR/neuroinflammatory outputs are context-dependent and should remain non-core.
    supported_by:
    - reference_id: PMID:24990881
      supporting_text: TREM2 is an innate immune receptor preferentially expressed by microglia and 
        is involved in inflammation and phagocytosis.
references:
- id: GO_REF:0000024
  title: Manual transfer of experimentally-verified manual GO annotation data to orthologs by 
    curator judgment of sequence similarity
  findings: []
- id: GO_REF:0000033
  title: Annotation inferences using phylogenetic trees
  findings: []
- id: GO_REF:0000044
  title: Gene Ontology annotation based on UniProtKB/Swiss-Prot Subcellular Location vocabulary 
    mapping, accompanied by conservative changes to GO terms applied by UniProt
  findings: []
- id: GO_REF:0000107
  title: Automatic transfer of experimentally verified manual GO annotation data to orthologs using 
    Ensembl Compara
  findings: []
- id: GO_REF:0000117
  title: Electronic Gene Ontology annotations created by ARBA machine learning models
  findings: []
- id: GO_REF:0000120
  title: Combined Automated Annotation using Multiple IEA Methods
  findings: []
- id: PMID:10799849
  title: 'Cutting edge: inflammatory responses can be triggered by TREM-1, a novel receptor expressed
    on neutrophils and monocytes.'
  findings: []
- id: PMID:11602640
  title: A DAP12-mediated pathway regulates expression of CC chemokine receptor 7 and maturation of 
    human dendritic cells.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Cached full-text record directly establishes human TREM2 as a
      DAP12-associated cell-surface receptor and supports downstream kinase/ERK
      signaling.
- id: PMID:12925681
  title: DAP12/TREM2 deficiency results in impaired osteoclast differentiation and osteoporotic 
    features.
  findings: []
- id: PMID:16715077
  title: Plexin-A1 and its interaction with DAP12 in immune responses and bone homeostasis.
  findings: []
- id: PMID:21841309
  title: OSCAR is a collagen receptor that costimulates osteoclastogenesis in DAP12-deficient humans
    and mice.
  findings: []
- id: PMID:24078628
  title: Sequential proteolytic processing of the triggering receptor expressed on myeloid cells-2 
    (TREM2) protein by ectodomain shedding and γ-secretase-dependent intramembranous cleavage.
  findings: []
- id: PMID:24990881
  title: TREM2 mutations implicated in neurodegeneration impair cell surface transport and 
    phagocytosis.
  findings: []
- id: PMID:25615530
  title: Disease-Associated Mutations of TREM2 Alter the Processing of N-Linked Oligosaccharides in 
    the Golgi Apparatus.
  findings: []
- id: PMID:27044754
  title: FRMD4A-cytohesin signaling modulates the cellular release of tau.
  findings: []
- id: PMID:27477018
  title: TREM2 Binds to Apolipoproteins, Including APOE and CLU/APOJ, and Thereby Facilitates Uptake
    of Amyloid-Beta by Microglia.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Cached abstract directly supports TREM2 binding to 
      apolipoproteins/lipoprotein particles, including APOE and CLU, and 
      TREM2-dependent uptake of amyloid-lipoprotein complexes.
- id: PMID:27589997
  title: Rare TREM2 variants associated with Alzheimer's disease display reduced cell surface 
    expression.
  findings: []
- id: PMID:27995897
  title: Neurodegenerative disease mutations in TREM2 reveal a functional surface and distinct 
    loss-of-function mechanisms.
  findings: []
- id: PMID:28490631
  title: A split-luciferase complementation, real-time reporting assay enables monitoring of the 
    disease-associated transmembrane protein TREM2 in live cells.
  findings: []
- id: PMID:28768830
  title: Neurodegeneration-associated mutant TREM2 proteins abortively cycle between the ER and 
    ER-Golgi intermediate compartment.
  findings: []
- id: PMID:28855300
  title: An Alzheimer-associated TREM2 variant occurs at the ADAM cleavage site and affects shedding
    and phagocytic function.
  findings: []
- id: PMID:28855301
  title: TREM2 shedding by cleavage at the H157-S158 bond is accelerated for the Alzheimer's 
    disease-associated H157Y variant.
  findings: []
- id: PMID:28923481
  title: ADAM17 is the main sheddase for the generation of human triggering receptor expressed in 
    myeloid cells (hTREM2) ectodomain and cleaves TREM2 after Histidine 157.
  findings: []
- id: PMID:29518356
  title: TREM2 Is a Receptor for β-Amyloid that Mediates Microglial Function.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Cached record supports TREM2 amyloid-beta receptor activity
      and microglial functional consequences; used as Alzheimer-context ligand
      evidence.
- id: PMID:29518357
  title: Elevated TREM2 Gene Dosage Reprograms Microglia Responsivity and Ameliorates Pathological 
    Phenotypes in Alzheimer's Disease Models.
  findings: []
- id: PMID:29611543
  title: Intracellular trafficking of TREM2 is regulated by presenilin 1.
  findings: []
- id: PMID:29752066
  title: The Microglial Innate Immune Receptor TREM2 Is Required for Synapse Elimination and Normal 
    Brain Connectivity.
  findings: []
- id: PMID:30333625
  title: LILRB4 signalling in leukaemia cells mediates T cell suppression and tumour infiltration.
  findings: []
- id: PMID:30341064
  title: High-affinity interactions and signal transduction between Aβ oligomers and TREM2.
  findings: []
- id: PMID:30683932
  title: TREM2 acts as a tumor suppressor in hepatocellular carcinoma by targeting the 
    PI3K/Akt/β-catenin pathway.
  findings: []
- id: PMID:31101881
  title: Aminophospholipids are signal-transducing TREM2 ligands on apoptotic cells.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Cached abstract supports phosphatidylserine and 
      phosphatidylethanolamine as signal-transducing TREM2 ligands on apoptotic
      cells.
- id: PMID:31235932
  title: TREM2 function impedes tau seeding in neuritic plaques.
  findings: []
- id: PMID:31413141
  title: The MS4A gene cluster is a key modulator of soluble TREM2 and Alzheimer's disease risk.
  findings: []
- id: PMID:31462511
  title: Increased soluble TREM2 in cerebrospinal fluid is associated with reduced cognitive and 
    clinical decline in Alzheimer's disease.
  findings: []
- id: PMID:31902528
  title: TREM2 Regulates Microglial Cholesterol Metabolism upon Chronic Phagocytic Challenge.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Cached abstract supports TREM2 lipid/myelin sensing, 
      microglial phagocytosis, and cholesterol-metabolism adaptation after 
      chronic phagocytic challenge.
- id: PMID:32514138
  title: Alzheimer's-associated PLCγ2 is a signaling node required for both TREM2 function and the 
    inflammatory response in human microglia.
  findings: []
- id: PMID:33097708
  title: Gene expression and functional deficits underlie TREM2-knockout microglia responses in 
    human models of Alzheimer's disease.
  findings: []
  reference_review:
    relevance: HIGH
    correctness: VERIFIED
    review_notes: Cached abstract supports human microglial TREM2 loss-of-
      function effects on survival, phagocytosis, migration, metabolism, and 
      inflammatory responses.
- id: Reactome:R-HSA-210289
  title: Recruitment of SYK to p-DAP12
  findings: []
- id: Reactome:R-HSA-210300
  title: Interaction of DAP12 and TREM2
  findings: []
- id: Reactome:R-HSA-2395412
  title: Phosphorylation of SYK
  findings: []
- id: Reactome:R-HSA-2395801
  title: Phosphorylation of LAT by p-SYK
  findings: []
- id: Reactome:R-HSA-2396594
  title: Phosphorylation of SLP-76 by p-SYK
  findings: []
- id: Reactome:R-HSA-2424480
  title: PI3K phosphorylates PIP2 to PIP3
  findings: []
- id: Reactome:R-HSA-2424482
  title: p85 regulatory unit of PI3K binds p-6Y-SYK
  findings: []
- id: Reactome:R-HSA-2424484
  title: Phosphorylation of BTK by p-SYK
  findings: []
- id: Reactome:R-HSA-2424486
  title: Phosphorylation and activation of VAV2/VAV3 by SYK
  findings: []
- id: Reactome:R-HSA-2424487
  title: Phosphorylation of PLC-gamma by p-BTK/p-SYK
  findings: []
- id: Reactome:R-HSA-416725
  title: SEMA6D binds to PLXNA1:TREM2:DAP12
  findings: []
- id: Reactome:R-HSA-5696358
  title: TREM,CD300 binds lipids
  findings: []
core_functions:
- molecular_function:
    id: GO:0004888
    label: transmembrane signaling receptor activity
  description: TREM2 is a plasma-membrane myeloid receptor that associates with TYROBP/DAP12 and 
    transduces ligand binding into intracellular kinase, PLCG2, calcium, ERK, and activation 
    programs in microglia and other myeloid cells.
  directly_involved_in:
  - id: GO:0007165
    label: signal transduction
  - id: GO:0045088
    label: regulation of innate immune response
  - id: GO:1902533
    label: positive regulation of intracellular signal transduction
  - id: GO:0050850
    label: positive regulation of calcium-mediated signaling
  - id: GO:0070374
    label: positive regulation of ERK1 and ERK2 cascade
  locations:
  - id: GO:0005886
    label: plasma membrane
  supported_by:
  - reference_id: PMID:11602640
    supporting_text: TREM-2 is a cell surface receptor on human monocyte-derived DCs, which is 
      associated with DAP12.
  - reference_id: PMID:11602640
    supporting_text: TREM-2/DAP12 promotes upregulation of CC chemokine receptor 7, partial DC 
      maturation, and DC survival through activation of protein tyrosine kinases and extracellular 
      signal-regulated kinase.
  - reference_id: PMID:32514138
    supporting_text: TREM2 signals through PLCγ2 to mediate cell survival, phagocytosis, processing 
      of neuronal debris, and lipid metabolism.
- molecular_function:
    id: GO:0008289
    label: lipid binding
  description: TREM2 recognizes lipid-rich damage and uptake ligands, including aminophospholipids 
    on apoptotic cells, lipoprotein/apolipoprotein particles, sulfatides, and 
    amyloid-beta-lipoprotein complexes, enabling microglial phagocytosis, activation, migration, and
    lipid handling.
  directly_involved_in:
  - id: GO:0060100
    label: positive regulation of phagocytosis, engulfment
  - id: GO:0043277
    label: apoptotic cell clearance
  - id: GO:1903980
    label: positive regulation of microglial cell activation
  - id: GO:1904141
    label: positive regulation of microglial cell migration
  - id: GO:0055088
    label: lipid homeostasis
  locations:
  - id: GO:0005886
    label: plasma membrane
  supported_by:
  - reference_id: PMID:27477018
    supporting_text: we identified a set of lipoprotein particles (including LDL) and 
      apolipoproteins (including CLU/APOJ and APOE) as ligands
  - reference_id: PMID:31101881
    supporting_text: TREM2-dependent signal transduction in response to apoptotic Neuro2a cells is 
      mediated by aminophospholipid ligands, phosphatidylserine and phosphatidylethanolamine
  - reference_id: PMID:31902528
    supporting_text: TREM2 senses lipids and mediates myelin phagocytosis
  - reference_id: PMID:33097708
    supporting_text: TREM2 deletion reduces microglial survival, impairs phagocytosis of key 
      substrates including APOE, and inhibits SDF-1α/CXCR4-mediated chemotaxis
proposed_new_terms: []
suggested_questions:
- question: Which TREM2 ligand classes are normal in-vivo microglial ligands across brain states, 
    and which are disease-model or injury-specific substrates?
  experts:
  - microglial biology experts
  - GO immune receptor curators
- question: How should TREM2/TYROBP/DAP12 signaling be represented without over-curating every 
    downstream cytokine, kinase, survival, or metabolic phenotype as a core function?
  experts:
  - innate immune signaling curators
  - microglial signaling experts
- question: Which amyloid-beta, synapse-pruning, and neuroinflammatory annotations reflect evolved 
    TREM2 biology versus Alzheimer model-stage-specific phenotypes?
  experts:
  - Alzheimer microglia experts
  - GO neuroimmune curators
suggested_experiments:
- description: 'Compare endogenous human TREM2 knock-in and knockout microglia across defined ligands:
    apoptotic cells, purified aminophospholipids, APOE/CLU lipoprotein particles, myelin debris, synaptosomes,
    and amyloid-beta-lipoprotein complexes.'
  hypothesis: TREM2 core ligand sensing can be decomposed into ligand classes with distinct 
    requirements for TYROBP/DAP12, PLCG2, SYK, and calcium signaling.
  experiment_type: ligand-defined microglial uptake and signaling assays
- description: Use in-vivo baseline, injury, demyelination, and amyloid/tau model time courses to 
    separate normal TREM2-dependent phagocytic functions from disease-stage-specific microglial 
    states.
  hypothesis: Some TREM2 annotations, especially amyloid clearance and synapse pruning, are 
    context-specific outputs of the same core receptor-phagocytosis program.
  experiment_type: time-course in-vivo microglial transcriptomics and functional assays
- description: Quantify cell-surface TREM2, soluble TREM2, and TYROBP-associated signaling after 
    endogenous AD-risk variant knock-in under non-disease and challenge conditions.
  hypothesis: Disease-associated variants alter ligand binding, surface trafficking, or signaling 
    amplitude without changing the core receptor function category.
  experiment_type: endogenous variant knock-in receptor trafficking and signaling