Hspa8

UniProt ID: P63018
Organism: Rattus norvegicus
Review Status: DRAFT
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Gene Description

Hspa8 (UniProt P63018; HSP7C_RAT) encodes the rat Heat shock cognate 71 kDa protein, the constitutively expressed cytosolic HSP70-family chaperone known as HSC70/HSC73. Unlike the stress-inducible Hspa1a/Hspa1b (HSP70) paralogs, HSC70 is expressed under basal conditions and is the housekeeping HSP70. Its core molecular function is an ATP-dependent (EC 3.6.4.10) protein-folding chaperone activity built on the canonical HSP70 architecture: an N-terminal nucleotide-binding/ATPase domain and a C-terminal substrate-binding domain with a helical lid, whose nucleotide state (ATP = low affinity/fast client exchange; ADP = high affinity) governs cyclic binding and release of unfolded/misfolded client polypeptides. The ATPase cycle is driven by J-domain (HSP40/DNAJ) co-chaperones that stimulate ATP hydrolysis and reset by nucleotide-exchange factors. Three best-supported, mechanistically distinct roles emerge: (1) general proteostasis - protein folding, refolding, unfolding and disassembly of protein complexes; (2) clathrin-coated vesicle uncoating - the clathrin-uncoating ATPase activity in which the J-domain co-chaperone auxilin (DNAJC6) recruits HSC70 to clathrin-coated vesicles and stimulates its ATPase to disassemble clathrin coats, central to clathrin-mediated endocytosis and synaptic vesicle recycling; and (3) chaperone-mediated autophagy (CMA) - cytosolic HSC70 recognizes KFERQ-like pentapeptide motifs in soluble client proteins and delivers them to LAMP-2A at the lysosomal membrane, with lysosomal HSC70 supporting translocation into the lysosomal lumen for degradation. HSC70 acts predominantly in the cytosol, with additional functional pools at the lysosomal membrane/lumen and at endocytic/synaptic trafficking structures. It is a highly pleiotropic protein; many of its annotated processes and locations reflect this pleiotropy rather than its evolved core function.

Existing Annotations Review

GO Term Evidence Action Reason
GO:0005634 nucleus
IBA
GO_REF:0000033
KEEP AS NON CORE
Summary: Manual review: HSC70 has documented nuclear/nucleolar pools (e.g. nuclear import roles), but the nucleus is not the site of its core chaperone, clathrin-uncoating, or CMA functions.
Reason: Valid localization but peripheral to the evolved cytosolic chaperone function.
GO:0005737 cytoplasm
IBA
GO_REF:0000033
ACCEPT
Summary: Manual review: cytoplasm is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005886 plasma membrane
IBA
GO_REF:0000033
MARK AS OVER ANNOTATED
Summary: Manual review: plasma membrane is not the site of the core chaperone function of HSC70, which acts in the cytosol; surface/extracellular pools are context-specific (e.g. exosomal/cell-surface) and do not represent the evolved function.
Reason: Localization over-annotation; the core HSC70 function is cytosolic, not at the plasma membrane.
Propagation Review
Root cause: PROPAGATION BAD
Failure modes: COMPARTMENT OR COMPLEX MISMATCH CONTEXT OR TISSUE MISMATCH
Sources checked:
PANTHER:PTN002500132 · PANTHER:PTN002500132 SOURCE STALE OR MISSING
The GOA WITH/FROM names this node with mostly fungal (Candida) cell-surface members as seeds, but no IBD for GO:0005886 remains anywhere in the current PTHR19375 PAINT slice (snapshot rows dated through 2026-08-28); the node now carries only nucleus (GO:0005634) and cytosol (GO:0005829). PAINT has therefore withdrawn or re-scoped the plasma-membrane assertion, which confirms the corrective call here: the compartment reflected fungal cell-surface and moonlighting contexts, whereas the core HSC70 chaperone cycle is cytosolic.
GO:0016887 ATP hydrolysis activity
IBA
GO_REF:0000033
ACCEPT
Summary: Manual review: ATP hydrolysis activity underpins the ATP-dependent chaperone cycle of Hspa8/HSC70, in which the nucleotide state controls client affinity and is regulated by J-domain co-chaperones.
Reason: Retained as supporting the core ATP-driven chaperone cycle, corroborated by falcon deep research.
Supporting Evidence:
file:rat/Hspa8/Hspa8-deep-research-falcon.md
nucleotide state controls substrate affinity (ATP-bound low affinity/fast exchange; ADP-bound high affinity/slow exchange)
GO:0031072 heat shock protein binding
IBA
GO_REF:0000033
ACCEPT
Summary: Manual review: heat shock protein binding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0044183 protein folding chaperone
IBA
GO_REF:0000033
ACCEPT
Summary: Manual review: protein folding chaperone is the core molecular function of Hspa8/HSC70; the constitutive HSP70-family ATP-dependent chaperone that assists folding, refolding, unfolding and complex disassembly of client proteins.
Reason: Retained as the core molecular function, corroborated by falcon deep research.
Supporting Evidence:
file:rat/Hspa8/Hspa8-deep-research-falcon.md
Hspa8/HSC70 is a **constitutive** HSP70-family **ATPase chaperone** that supports proteostasis by assisting **protein folding/refolding**, **unfolding**, and **disassembly of protein complexes**
GO:0005829 cytosol
IBA
GO_REF:0000033
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0072318 clathrin coat disassembly
IBA
GO_REF:0000033
ACCEPT
Summary: Manual review: clathrin coat disassembly is a core process for Hspa8/HSC70; the auxilin (DNAJC6)-recruited HSC70 ATPase disassembles clathrin coats from clathrin-coated vesicles.
Reason: Retained as a core process, corroborated by falcon deep research.
Supporting Evidence:
file:rat/Hspa8/Hspa8-deep-research-falcon.md
the **J-domain co-chaperone auxilin (DNAJC6)** recruits HSC70 to CCVs and **stimulates HSC70 ATPase activity** to disassemble clathrin coats
GO:0042026 protein refolding
IBA
GO_REF:0000033
ACCEPT
Summary: Manual review: protein refolding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0007165 signal transduction
IEA
GO_REF:0000108
MARK AS OVER ANNOTATED
Summary: Manual review: signal transduction is an over-broad inter-ontology IEA inference; HSC70 is a chaperone and does not act as a signal transducer. This generic term over-annotates the gene.
Reason: Over-broad IEA inference that does not reflect the evolved chaperone function of HSC70.
GO:0000166 nucleotide binding
IEA
GO_REF:0000043
ACCEPT
Summary: Manual review: nucleotide binding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0001916 positive regulation of T cell mediated cytotoxicity
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: Manual review: positive regulation of T cell mediated cytotoxicity may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005524 ATP binding
IEA
GO_REF:0000120
ACCEPT
Summary: Manual review: ATP binding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005681 spliceosomal complex
IEA
GO_REF:0000043
KEEP AS NON CORE
Summary: Manual review: spliceosomal complex may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005730 nucleolus
IEA
GO_REF:0000044
KEEP AS NON CORE
Summary: Manual review: nucleolus may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005737 cytoplasm
IEA
GO_REF:0000120
ACCEPT
Summary: Manual review: cytoplasm is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005765 lysosomal membrane
IEA
GO_REF:0000044
ACCEPT
Summary: Manual review: the lysosomal membrane is a functional location of the core chaperone-mediated autophagy role of HSC70, where the chaperone-substrate complex engages LAMP-2A; this electronic annotation is consistent with the experimentally supported IDA annotations to the same term.
Reason: Lysosomal membrane is a bona fide functional location for the core CMA role of HSC70 (experimentally supported by PMID:18644871 and PMID:9038169); accepted for consistency with those IDA annotations.
GO:0005886 plasma membrane
IEA
GO_REF:0000044
MARK AS OVER ANNOTATED
Summary: Manual review: plasma membrane is not the site of the core chaperone function of HSC70; surface/extracellular pools are context-specific and do not represent the evolved function.
Reason: Localization over-annotation; the core HSC70 function is cytosolic, not at the plasma membrane.
GO:0006397 mRNA processing
IEA
GO_REF:0000043
KEEP AS NON CORE
Summary: Manual review: mRNA processing may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0006914 autophagy
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: Manual review: autophagy may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0008289 lipid binding
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: Manual review: lipid binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0008380 RNA splicing
IEA
GO_REF:0000043
KEEP AS NON CORE
Summary: Manual review: RNA splicing may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0016787 hydrolase activity
IEA
GO_REF:0000043
MARK AS OVER ANNOTATED
Summary: Manual review: HSC70 is indeed a hydrolase (an ATPase, EC 3.6.4.10), so GO:0016787 is not factually wrong, but it is an over-broad parent term. The more specific GO:0016887 (ATP hydrolysis activity) is already ACCEPTed and captures the actual catalytic function.
Reason: GO:0016887 (ATP hydrolysis activity) is already ACCEPTed and is more specific; GO:0016787 (hydrolase activity) is the over-broad parent and is redundant.
GO:0016887 ATP hydrolysis activity
IEA
GO_REF:0000120
ACCEPT
Summary: Manual review: ATP hydrolysis activity is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0042470 melanosome
IEA
GO_REF:0000044
KEEP AS NON CORE
Summary: Manual review: melanosome may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043066 negative regulation of apoptotic process
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: Manual review: negative regulation of apoptotic process may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043168 anion binding
IEA
GO_REF:0000117
KEEP AS NON CORE
Summary: Manual review: anion binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0001786 phosphatidylserine binding
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: phosphatidylserine binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005770 late endosome
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: late endosome may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005829 cytosol
IEA
GO_REF:0000107
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0030674 protein-macromolecule adaptor activity
IEA
GO_REF:0000107
ACCEPT
Summary: Manual review: protein-macromolecule adaptor activity is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0035651 AP-3 adaptor complex binding
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: AP-3 adaptor complex binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0044788 host-mediated perturbation of viral process
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: host-mediated perturbation of viral process may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0044829 host-mediated activation of viral genome replication
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: host-mediated activation of viral genome replication may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0048471 perinuclear region of cytoplasm
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: perinuclear region of cytoplasm may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0051082 unfolded protein binding
IEA
GO_REF:0000120
MODIFY
Summary: Manual review: unfolded protein binding is better represented by a more specific replacement term for Hspa8.
Reason: Modified to align with current curation guidance and improve term specificity.
Proposed replacements: protein folding chaperone
GO:0061684 chaperone-mediated autophagy
IEA
GO_REF:0000107
ACCEPT
Summary: Manual review: chaperone-mediated autophagy is a core process for Hspa8/HSC70, which selects KFERQ-motif clients and delivers them to LAMP-2A for lysosomal degradation.
Reason: Retained as a core process, corroborated by falcon deep research.
Supporting Evidence:
file:rat/Hspa8/Hspa8-deep-research-falcon.md
recognition of KFERQ-like pentapeptide motifs** in client proteins targeted for **chaperone-mediated autophagy (CMA)**
GO:0061738 late endosomal microautophagy
IEA
GO_REF:0000107
ACCEPT
Summary: Manual review: late endosomal microautophagy is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0070062 extracellular exosome
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: extracellular exosome may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0072318 clathrin coat disassembly
IEA
GO_REF:0000120
ACCEPT
Summary: Manual review: clathrin coat disassembly is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0098684 photoreceptor ribbon synapse
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: photoreceptor ribbon synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098690 glycinergic synapse
IEA
GO_REF:0000120
KEEP AS NON CORE
Summary: Manual review: glycinergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098793 presynapse
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: presynapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098978 glutamatergic synapse
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: glutamatergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0099175 regulation of postsynapse organization
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: regulation of postsynapse organization may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0099634 postsynaptic specialization membrane
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: postsynaptic specialization membrane may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0160020 positive regulation of ferroptosis
IEA
GO_REF:0000107
KEEP AS NON CORE
Summary: Manual review: positive regulation of ferroptosis may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0030674 protein-macromolecule adaptor activity
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: protein-macromolecule adaptor activity is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0061684 chaperone-mediated autophagy
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: chaperone-mediated autophagy is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0160020 positive regulation of ferroptosis
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: positive regulation of ferroptosis may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005737 cytoplasm
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: cytoplasm is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0101031 protein folding chaperone complex
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: protein folding chaperone complex is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005615 extracellular space
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: extracellular space may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0030335 positive regulation of cell migration
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: positive regulation of cell migration may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0048018 receptor ligand activity
ISO
GO_REF:0000121
REMOVE
Summary: Manual review: receptor ligand activity is not sufficiently supported as a direct function of Hspa8.
Reason: Removed due weak mechanistic support or likely misannotation for this gene.
Propagation Review
Root cause: SOURCE WEAK OR INFERRED
Failure modes: SOURCE EVIDENCE WEAK
Sources checked:
UniProtKB:P11142 · human HSPA8 SOURCE WEAK OR INFERRED
Human HSPA8 donor claim of receptor-ligand activity has weak mechanistic support and is a likely misannotation; nothing robust transfers to rat.
GO:0006457 protein folding
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: protein folding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0014069 postsynaptic density
IEP
PMID:11230095
Presence of both constitutive and inducible forms of heat sh...
KEEP AS NON CORE
Summary: Manual review: postsynaptic density may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0014069 postsynaptic density
IDA
PMID:11230095
Presence of both constitutive and inducible forms of heat sh...
KEEP AS NON CORE
Summary: Manual review: postsynaptic density may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0014069 postsynaptic density
IDA
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: postsynaptic density may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098684 photoreceptor ribbon synapse
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: photoreceptor ribbon synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098690 glycinergic synapse
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: glycinergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098690 glycinergic synapse
IDA
PMID:21209184
Heat shock cognate protein 70 regulates gephyrin clustering.
KEEP AS NON CORE
Summary: Manual review: glycinergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098690 glycinergic synapse
EXP
PMID:21209184
Heat shock cognate protein 70 regulates gephyrin clustering.
KEEP AS NON CORE
Summary: Manual review: glycinergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098690 glycinergic synapse
IMP
PMID:21209184
Heat shock cognate protein 70 regulates gephyrin clustering.
KEEP AS NON CORE
Summary: Manual review: glycinergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098793 presynapse
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: presynapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098793 presynapse
IEP
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: presynapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098793 presynapse
IDA
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: presynapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098794 postsynapse
IEP
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: postsynapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098794 postsynapse
IDA
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: postsynapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098880 maintenance of postsynaptic specialization structure
IDA
PMID:21209184
Heat shock cognate protein 70 regulates gephyrin clustering.
KEEP AS NON CORE
Summary: Manual review: maintenance of postsynaptic specialization structure may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098880 maintenance of postsynaptic specialization structure
EXP
PMID:21209184
Heat shock cognate protein 70 regulates gephyrin clustering.
KEEP AS NON CORE
Summary: Manual review: maintenance of postsynaptic specialization structure may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098880 maintenance of postsynaptic specialization structure
IMP
PMID:21209184
Heat shock cognate protein 70 regulates gephyrin clustering.
KEEP AS NON CORE
Summary: Manual review: maintenance of postsynaptic specialization structure may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098978 glutamatergic synapse
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: glutamatergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098978 glutamatergic synapse
IEP
PMID:11230095
Presence of both constitutive and inducible forms of heat sh...
KEEP AS NON CORE
Summary: Manual review: glutamatergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098978 glutamatergic synapse
IDA
PMID:11230095
Presence of both constitutive and inducible forms of heat sh...
KEEP AS NON CORE
Summary: Manual review: glutamatergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098978 glutamatergic synapse
IEP
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: glutamatergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0098978 glutamatergic synapse
IDA
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: glutamatergic synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0099175 regulation of postsynapse organization
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: regulation of postsynapse organization may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0099634 postsynaptic specialization membrane
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: postsynaptic specialization membrane may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0072318 clathrin coat disassembly
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: clathrin coat disassembly is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0072318 clathrin coat disassembly
IDA
PMID:8524399
Role of auxilin in uncoating clathrin-coated vesicles.
ACCEPT
Summary: Manual review: clathrin coat disassembly is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005765 lysosomal membrane
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: the lysosomal membrane is a functional location of the core chaperone-mediated autophagy role of HSC70; this orthology-based annotation is consistent with the experimentally supported IDA annotations to the same term.
Reason: Lysosomal membrane is a bona fide functional location for the core CMA role of HSC70 (experimentally supported by PMID:18644871 and PMID:9038169); accepted for consistency with those IDA annotations.
GO:0030674 protein-macromolecule adaptor activity
ISS
GO_REF:0000024
ACCEPT
Summary: Manual review: protein-macromolecule adaptor activity is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0061740 protein targeting to lysosome involved in chaperone-mediated autophagy
ISS
GO_REF:0000024
ACCEPT
Summary: Manual review: protein targeting to lysosome in CMA is a core process for Hspa8/HSC70; cytosolic HSC70 recognizes KFERQ-like motifs and delivers clients to LAMP-2A, and lysosomal HSC70 supports translocation into the lysosomal lumen.
Reason: Retained as a core CMA process, corroborated by falcon deep research.
Supporting Evidence:
file:rat/Hspa8/Hspa8-deep-research-falcon.md
recognition of KFERQ-like pentapeptide motifs** in client proteins targeted for **chaperone-mediated autophagy (CMA)**
file:rat/Hspa8/Hspa8-deep-research-falcon.md
Lysosomal HSC70 (Lys‑HSC70)** facilitates **substrate translocation** into the lysosome lumen
GO:1900226 negative regulation of NLRP3 inflammasome complex assembly
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: negative regulation of NLRP3 inflammasome complex assembly may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0001786 phosphatidylserine binding
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: phosphatidylserine binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0061635 regulation of protein complex stability
IDA
PMID:18644871
The chaperone-mediated autophagy receptor organizes in dynam...
KEEP AS NON CORE
Summary: Manual review: regulation of protein complex stability may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0140545 ATP-dependent protein disaggregase activity
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: ATP-dependent protein disaggregase activity is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0061684 chaperone-mediated autophagy
IGI
PMID:18644871
The chaperone-mediated autophagy receptor organizes in dynam...
ACCEPT
Summary: Manual review: chaperone-mediated autophagy is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0043202 lysosomal lumen
TAS
Reactome:R-RNO-9625160
KEEP AS NON CORE
Summary: Manual review: lysosomal lumen may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043202 lysosomal lumen
TAS
Reactome:R-RNO-9625173
KEEP AS NON CORE
Summary: Manual review: lysosomal lumen may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043202 lysosomal lumen
TAS
Reactome:R-RNO-9626028
KEEP AS NON CORE
Summary: Manual review: lysosomal lumen may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0048471 perinuclear region of cytoplasm
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: perinuclear region of cytoplasm may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005829 cytosol
TAS
Reactome:R-RNO-9613356
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005829 cytosol
TAS
Reactome:R-RNO-9615712
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005829 cytosol
TAS
Reactome:R-RNO-9620205
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005829 cytosol
TAS
Reactome:R-RNO-9622832
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005829 cytosol
TAS
Reactome:R-RNO-9626236
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005829 cytosol
TAS
Reactome:R-RNO-9626239
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005829 cytosol
TAS
Reactome:R-RNO-9626254
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0043195 terminal bouton
HDA
PMID:24876496
Composition of isolated synaptic boutons reveals the amounts...
KEEP AS NON CORE
Summary: Manual review: terminal bouton may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0031625 ubiquitin protein ligase binding
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: ubiquitin protein ligase binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0051087 protein-folding chaperone binding
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: protein-folding chaperone binding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0005515 protein binding
IPI
PMID:15708368
Small glutamine-rich tetratricopeptide repeat-containing pro...
MARK AS OVER ANNOTATED
Summary: Manual review: protein binding is too generic or over-extended for Hspa8.
Reason: Marked over-annotated because more specific terms capture the biology more accurately.
GO:0030425 dendrite
IDA
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: dendrite may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0044309 neuron spine
IDA
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: neuron spine may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0061740 protein targeting to lysosome involved in chaperone-mediated autophagy
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: protein targeting to lysosome involved in chaperone-mediated autophagy is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0044788 host-mediated perturbation of viral process
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: host-mediated perturbation of viral process may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0044829 host-mediated activation of viral genome replication
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: host-mediated activation of viral genome replication may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005765 lysosomal membrane
IDA
PMID:18644871
The chaperone-mediated autophagy receptor organizes in dynam...
ACCEPT
Summary: Manual review: this IDA derives from a key CMA mechanistic paper showing HSC70 organizes in dynamic protein complexes with LAMP-2A at the lysosomal membrane. The lysosomal membrane is a functional location of the core CMA role of HSC70 (also listed in the CMA core_function), so this experimental annotation is accepted as core.
Reason: The lysosomal membrane is a bona fide functional location for the core chaperone-mediated autophagy role of HSC70, directly demonstrated experimentally in PMID:18644871. Falcon corroboration is captured in the references findings block.
GO:0032984 protein-containing complex disassembly
IMP
PMID:18644871
The chaperone-mediated autophagy receptor organizes in dynam...
KEEP AS NON CORE
Summary: Manual review: protein-containing complex disassembly may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:1990836 lysosomal matrix
IDA
PMID:18644871
The chaperone-mediated autophagy receptor organizes in dynam...
KEEP AS NON CORE
Summary: Manual review: lysosomal matrix may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005770 late endosome
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: late endosome may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0061738 late endosomal microautophagy
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: late endosomal microautophagy is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0016887 ATP hydrolysis activity
IDA
PMID:20060297
Chaperone-assisted selective autophagy is essential for musc...
ACCEPT
Summary: Manual review: ATP hydrolysis activity is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:1904764 chaperone-mediated autophagy translocation complex disassembly
IDA
PMID:18644871
The chaperone-mediated autophagy receptor organizes in dynam...
KEEP AS NON CORE
Summary: Manual review: chaperone-mediated autophagy translocation complex disassembly may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0042026 protein refolding
ISS
GO_REF:0000024
ACCEPT
Summary: Manual review: protein refolding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0061684 chaperone-mediated autophagy
IDA
PMID:18644871
The chaperone-mediated autophagy receptor organizes in dynam...
ACCEPT
Summary: Manual review: chaperone-mediated autophagy is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0031647 regulation of protein stability
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: regulation of protein stability may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0097214 positive regulation of lysosomal membrane permeability
IMP
PMID:9038169
A population of rat liver lysosomes responsible for the sele...
KEEP AS NON CORE
Summary: Manual review: positive regulation of lysosomal membrane permeability may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:1990832 slow axonal transport
IEP
PMID:11933046
Slow axonal transport of the cytosolic chaperonin CCT with H...
KEEP AS NON CORE
Summary: Manual review: slow axonal transport may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:1990833 clathrin-uncoating ATPase activity
IDA
PMID:8363588
High-level expression of soluble rat hsc70 in Escherichia co...
ACCEPT
Summary: Manual review: clathrin-uncoating ATPase activity is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:1990834 response to odorant
IEP
PMID:11268007
Odorants as cell-type specific activators of a heat shock re...
KEEP AS NON CORE
Summary: Manual review: response to odorant may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0000082 G1/S transition of mitotic cell cycle
IEP
PMID:10198213
73-kDa heat shock cognate protein interacts directly with P2...
KEEP AS NON CORE
Summary: Manual review: G1/S transition of mitotic cell cycle may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0070301 cellular response to hydrogen peroxide
IEP
PMID:10329212
A physiologically relevant hyperthermia selectively activate...
KEEP AS NON CORE
Summary: Manual review: cellular response to hydrogen peroxide may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:1904593 prostaglandin binding
IPI
PMID:21445266
Proteomic identification of protein targets for 15-deoxy-Δ(1...
REMOVE
Summary: Manual review: prostaglandin binding is not sufficiently supported as a direct function of Hspa8.
Reason: Removed due weak mechanistic support or likely misannotation for this gene.
GO:0001916 positive regulation of T cell mediated cytotoxicity
IMP
PMID:8871636
Involvement of peptide antigens in the cytotoxicity between ...
KEEP AS NON CORE
Summary: Manual review: positive regulation of T cell mediated cytotoxicity may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005776 autophagosome
IDA
PMID:20392947
Essential role of the redox-sensitive kinase p66shc in deter...
KEEP AS NON CORE
Summary: Manual review: autophagosome may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005874 microtubule
IDA
PMID:2527848
Beta-internexin is a microtubule-associated protein identica...
KEEP AS NON CORE
Summary: Manual review: microtubule may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005882 intermediate filament
IDA
PMID:7721954
Integrity of intermediate filaments is associated with the d...
KEEP AS NON CORE
Summary: Manual review: intermediate filament may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0009986 cell surface
IDA
PMID:8228242
70 kDa heat shock cognate protein is a transformation-associ...
KEEP AS NON CORE
Summary: Manual review: cell surface may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0010628 positive regulation of gene expression
IMP
PMID:8734434
Antisense oligonucleotide to the 70-kDa heat shock cognate p...
KEEP AS NON CORE
Summary: Manual review: positive regulation of gene expression may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0032279 asymmetric synapse
IDA
PMID:11230095
Presence of both constitutive and inducible forms of heat sh...
KEEP AS NON CORE
Summary: Manual review: asymmetric synapse may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043197 dendritic spine
IDA
PMID:11230095
Presence of both constitutive and inducible forms of heat sh...
KEEP AS NON CORE
Summary: Manual review: dendritic spine may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043198 dendritic shaft
IDA
PMID:11230095
Presence of both constitutive and inducible forms of heat sh...
KEEP AS NON CORE
Summary: Manual review: dendritic shaft may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043204 perikaryon
IDA
PMID:11230095
Presence of both constitutive and inducible forms of heat sh...
KEEP AS NON CORE
Summary: Manual review: perikaryon may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0045471 response to ethanol
IEP
PMID:12170112
Stress response caused by chronic alcohol intake in aged rat...
KEEP AS NON CORE
Summary: Manual review: response to ethanol may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0050766 positive regulation of phagocytosis
IMP
PMID:9886410
Clathrin-coated pit-associated proteins are required for alv...
KEEP AS NON CORE
Summary: Manual review: positive regulation of phagocytosis may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:1904592 positive regulation of protein refolding
IDA
PMID:11360998
The effect of mutating arginine-469 on the substrate binding...
KEEP AS NON CORE
Summary: Manual review: positive regulation of protein refolding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0001822 kidney development
IEP
PMID:9764759
Constitutive expression of heat shock proteins Hsp90, Hsc70,...
KEEP AS NON CORE
Summary: Manual review: kidney development may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0001917 photoreceptor inner segment
IDA
PMID:10549657
Constitutive and inducible heat shock protein 70 immunoreact...
KEEP AS NON CORE
Summary: Manual review: photoreceptor inner segment may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005765 lysosomal membrane
IDA
PMID:9038169
A population of rat liver lysosomes responsible for the sele...
ACCEPT
Summary: Manual review: PMID:9038169 (Cuervo, Dice & Knecht 1997) is a foundational CMA paper showing lysosomal hsc73 mediates selective uptake/degradation of cytosolic substrates in rat liver lysosomes. The lysosomal membrane is a functional location of the core CMA role of HSC70.
Reason: The lysosomal membrane is a bona fide functional location for the core chaperone-mediated autophagy role of HSC70, directly demonstrated experimentally in PMID:9038169.
GO:0006606 protein import into nucleus
IMP
PMID:8892974
Evidence for the existence of a novel mechanism for the nucl...
KEEP AS NON CORE
Summary: Manual review: protein import into nucleus may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0007519 skeletal muscle tissue development
IEP
PMID:12909603
Endogenous expression and developmental changes of HSP72 in ...
KEEP AS NON CORE
Summary: Manual review: skeletal muscle tissue development may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0008088 axo-dendritic transport
IEP
PMID:11933046
Slow axonal transport of the cytosolic chaperonin CCT with H...
KEEP AS NON CORE
Summary: Manual review: axo-dendritic transport may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0009410 response to xenobiotic stimulus
IEP
PMID:10425221
Identification of a novel splice variant of heat shock cogna...
KEEP AS NON CORE
Summary: Manual review: response to xenobiotic stimulus may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0014823 response to activity
IEP
PMID:10606824
The effects of exercise duration on adrenal HSP72/73 inducti...
KEEP AS NON CORE
Summary: Manual review: response to activity may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0021549 cerebellum development
IEP
PMID:9764759
Constitutive expression of heat shock proteins Hsp90, Hsc70,...
KEEP AS NON CORE
Summary: Manual review: cerebellum development may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0034605 cellular response to heat
IEP
PMID:8955942
Differential expression of heat shock 70 proteins in primary...
KEEP AS NON CORE
Summary: Manual review: cellular response to heat may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0042277 peptide binding
IDA
PMID:8363588
High-level expression of soluble rat hsc70 in Escherichia co...
KEEP AS NON CORE
Summary: Manual review: peptide binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0042594 response to starvation
IEP
PMID:9038169
A population of rat liver lysosomes responsible for the sele...
KEEP AS NON CORE
Summary: Manual review: response to starvation may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0044743 protein transmembrane import into intracellular organelle
IEP
PMID:9038169
A population of rat liver lysosomes responsible for the sele...
KEEP AS NON CORE
Summary: Manual review: protein transmembrane import into intracellular organelle may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0045862 positive regulation of proteolysis
IDA
PMID:9987077
Direct lysosomal uptake of alpha 2-microglobulin contributes...
KEEP AS NON CORE
Summary: Manual review: positive regulation of proteolysis may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0046777 protein autophosphorylation
IDA
PMID:8420978
Aspartyl residue 10 is essential for ATPase activity of rat ...
REMOVE
Summary: Manual review: PMID:8420978 (Huang et al. 1993, "Aspartyl residue 10 is essential for ATPase activity of rat hsc70") studies the ATPase activity of HSC70, not protein autophosphorylation. The annotation appears to be a database error.
Reason: Misannotation: PMID:8420978 reports ATPase activity (the catalytic mechanism of HSC70), not protein autophosphorylation; there is no evidence in this reference that HSC70 autophosphorylates.
GO:0048471 perinuclear region of cytoplasm
IDA
PMID:10549657
Constitutive and inducible heat shock protein 70 immunoreact...
KEEP AS NON CORE
Summary: Manual review: perinuclear region of cytoplasm may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0071276 cellular response to cadmium ion
IEP
PMID:9736451
Involvement of heat shock elements and basal transcription e...
KEEP AS NON CORE
Summary: Manual review: cellular response to cadmium ion may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0072318 clathrin coat disassembly
IDA
PMID:8363588
High-level expression of soluble rat hsc70 in Escherichia co...
ACCEPT
Summary: Manual review: clathrin coat disassembly is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:1990836 lysosomal matrix
IDA
PMID:9038169
A population of rat liver lysosomes responsible for the sele...
KEEP AS NON CORE
Summary: Manual review: lysosomal matrix may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0010045 response to nickel cation
IEP
PMID:15736156
Effects of nickel poisoning on expression pattern of the 72/...
KEEP AS NON CORE
Summary: Manual review: response to nickel cation may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0010667 negative regulation of cardiac muscle cell apoptotic process
IMP
PMID:21958194
Constitutive heat shock protein 70 interacts with α-enolase ...
KEEP AS NON CORE
Summary: Manual review: negative regulation of cardiac muscle cell apoptotic process may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0019899 enzyme binding
IPI
PMID:21958194
Constitutive heat shock protein 70 interacts with α-enolase ...
KEEP AS NON CORE
Summary: Manual review: enzyme binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
IDA
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: axon may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0030900 forebrain development
IEP
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: forebrain development may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0031686 A1 adenosine receptor binding
IPI
PMID:10866672
The heat shock cognate protein hsc73 assembles with A(1) ade...
KEEP AS NON CORE
Summary: Manual review: PMID:10866672 documents a direct, specific rat hsc73-A1 adenosine receptor interaction (Kd ~0.5 nM by surface plasmon resonance, plus affinity chromatography, co-immunoprecipitation and confocal colocalization). This is a genuine documented interaction but is peripheral to the evolved core chaperone function.
Reason: Well-supported experimental rat interaction (PMID:10866672) but peripheral to the core HSC70 chaperone, clathrin-uncoating, and CMA functions; retained as non-core rather than removed.
GO:0032355 response to estradiol
IEP
PMID:9990085
The 70-kDa heat shock cognate protein (Hsc73) gene is enhanc...
KEEP AS NON CORE
Summary: Manual review: response to estradiol may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0032570 response to progesterone
IEP
PMID:9990085
The 70-kDa heat shock cognate protein (Hsc73) gene is enhanc...
KEEP AS NON CORE
Summary: Manual review: response to progesterone may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043025 neuronal cell body
IDA
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: neuronal cell body may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043085 positive regulation of catalytic activity
IMP
PMID:21958194
Constitutive heat shock protein 70 interacts with α-enolase ...
KEEP AS NON CORE
Summary: Manual review: positive regulation of catalytic activity may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043195 terminal bouton
IDA
PMID:9878698
Presence of molecular chaperones, heat shock cognate (Hsc) 7...
KEEP AS NON CORE
Summary: Manual review: terminal bouton may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0044849 estrous cycle
IEP
PMID:9990085
The 70-kDa heat shock cognate protein (Hsc73) gene is enhanc...
KEEP AS NON CORE
Summary: Manual review: estrous cycle may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0032991 protein-containing complex
IDA
PMID:20596529
Molecular composition of staufen2-containing ribonucleoprote...
KEEP AS NON CORE
Summary: Manual review: protein-containing complex may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0003723 RNA binding
IDA
PMID:23159318
Identification in the rat brain of a set of nuclear proteins...
KEEP AS NON CORE
Summary: Manual review: RNA binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005515 protein binding
IPI
PMID:23159318
Identification in the rat brain of a set of nuclear proteins...
MARK AS OVER ANNOTATED
Summary: Manual review: protein binding is too generic or over-extended for Hspa8.
Reason: Marked over-annotated because more specific terms capture the biology more accurately.
GO:0046034 ATP metabolic process
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: ATP metabolic process may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0055131 C3HC4-type RING finger domain binding
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: C3HC4-type RING finger domain binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0048026 positive regulation of mRNA splicing, via spliceosome
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: positive regulation of mRNA splicing, via spliceosome may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0001664 G protein-coupled receptor binding
ISO
GO_REF:0000121
REMOVE
Summary: Manual review: G protein-coupled receptor binding is not sufficiently supported as a direct function of Hspa8.
Reason: Removed due weak mechanistic support or likely misannotation for this gene.
Propagation Review
Root cause: SOURCE WEAK OR INFERRED
Failure modes: SOURCE EVIDENCE WEAK
Sources checked:
UniProtKB:P11142 · human HSPA8 SOURCE WEAK OR INFERRED
Human HSPA8 donor claim of G protein-coupled receptor binding has weak mechanistic support (likely misannotation) and does not warrant transfer.
GO:0070062 extracellular exosome
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: extracellular exosome may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005515 protein binding
IPI
PMID:18307834
Cathepsin C propeptide interacts with intestinal alkaline ph...
MARK AS OVER ANNOTATED
Summary: Manual review: protein binding is too generic or over-extended for Hspa8.
Reason: Marked over-annotated because more specific terms capture the biology more accurately.
GO:0005524 ATP binding
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: ATP binding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0019899 enzyme binding
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: enzyme binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0031072 heat shock protein binding
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: heat shock protein binding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:1902904 negative regulation of supramolecular fiber organization
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: negative regulation of supramolecular fiber organization may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005829 cytosol
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: cytosol is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0042026 protein refolding
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: protein refolding is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0051082 unfolded protein binding
ISO
GO_REF:0000121
MODIFY
Summary: Manual review: unfolded protein binding is better represented by a more specific replacement term for Hspa8.
Reason: Modified to align with current curation guidance and improve term specificity.
Propagation Review
Root cause: TERM SCOPING PROBLEM
Failure modes: GRANULARITY MISMATCH
Sources checked:
UniProtKB:P11142 · human HSPA8 SUPPORTS SOURCE BUT NOT TARGET
Human donor evidence for unfolded-protein binding is sound; the target-side issue is term choice -- protein folding chaperone (GO:0044183) better captures the Hsc70 activity.
MGI:MGI:105384 · mouse Hspa8 SUPPORTS SOURCE BUT NOT TARGET
Mouse Hspa8 donor likewise supports client-protein binding; folded into the same MODIFY to GO:0044183 protein folding chaperone.
Proposed replacements: protein folding chaperone
GO:0000974 Prp19 complex
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: Manual review: Prp19 complex may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0000974 Prp19 complex
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: Prp19 complex may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005634 nucleus
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: Manual review: HSC70 has documented nuclear/nucleolar pools, but the nucleus is not the site of its core chaperone, clathrin-uncoating, or CMA functions.
Reason: Valid localization but peripheral to the evolved cytosolic chaperone function.
GO:0005634 nucleus
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: HSC70 has documented nuclear/nucleolar pools, but the nucleus is not the site of its core chaperone, clathrin-uncoating, or CMA functions.
Reason: Valid localization but peripheral to the evolved cytosolic chaperone function.
GO:0045892 negative regulation of DNA-templated transcription
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: Manual review: negative regulation of DNA-templated transcription may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005515 protein binding
IPI
PMID:19457116
The molecular chaperone Hsc70 interacts with the vesicular m...
MARK AS OVER ANNOTATED
Summary: Manual review: protein binding is too generic or over-extended for Hspa8.
Reason: Marked over-annotated because more specific terms capture the biology more accurately.
GO:0008021 synaptic vesicle
IDA
PMID:19457116
The molecular chaperone Hsc70 interacts with the vesicular m...
KEEP AS NON CORE
Summary: Manual review: synaptic vesicle may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0043025 neuronal cell body
IDA
PMID:19457116
The molecular chaperone Hsc70 interacts with the vesicular m...
KEEP AS NON CORE
Summary: Manual review: neuronal cell body may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0045892 negative regulation of DNA-templated transcription
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: negative regulation of DNA-templated transcription may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005515 protein binding
IPI
PMID:17877381
Increased striatal mRNA and protein levels of the immunophil...
MARK AS OVER ANNOTATED
Summary: Manual review: protein binding is too generic or over-extended for Hspa8.
Reason: Marked over-annotated because more specific terms capture the biology more accurately.
GO:0005515 protein binding
IPI
PMID:9528774
The carboxy-terminal domain of Hsc70 provides binding sites ...
MARK AS OVER ANNOTATED
Summary: Manual review: protein binding is too generic or over-extended for Hspa8.
Reason: Marked over-annotated because more specific terms capture the biology more accurately.
GO:0016887 ATP hydrolysis activity
IDA
PMID:9528774
The carboxy-terminal domain of Hsc70 provides binding sites ...
ACCEPT
Summary: Manual review: ATP hydrolysis activity is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0032991 protein-containing complex
IDA
PMID:9528774
The carboxy-terminal domain of Hsc70 provides binding sites ...
KEEP AS NON CORE
Summary: Manual review: protein-containing complex may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005102 signaling receptor binding
IPI
PMID:11133993
Characteristics of the interaction between Hsc70 and the tra...
KEEP AS NON CORE
Summary: Manual review: signaling receptor binding may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005515 protein binding
IPI
PMID:10198213
73-kDa heat shock cognate protein interacts directly with P2...
MARK AS OVER ANNOTATED
Summary: Manual review: protein binding is too generic or over-extended for Hspa8.
Reason: Marked over-annotated because more specific terms capture the biology more accurately.
GO:0032991 protein-containing complex
IDA
PMID:10198213
73-kDa heat shock cognate protein interacts directly with P2...
KEEP AS NON CORE
Summary: Manual review: protein-containing complex may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0051082 unfolded protein binding
IDA
PMID:11133993
Characteristics of the interaction between Hsc70 and the tra...
MODIFY
Summary: Manual review: unfolded protein binding is better represented by a more specific replacement term for Hspa8.
Reason: Modified to align with current curation guidance and improve term specificity.
Proposed replacements: protein folding chaperone
GO:1990904 ribonucleoprotein complex
ISS
GO_REF:0000024
KEEP AS NON CORE
Summary: Manual review: ribonucleoprotein complex may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:1990904 ribonucleoprotein complex
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: ribonucleoprotein complex may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.
GO:0005515 protein binding
IPI
PMID:12514190
Cloning and characterization of a novel GRP78-binding protei...
MARK AS OVER ANNOTATED
Summary: Manual review: protein binding is too generic or over-extended for Hspa8.
Reason: Marked over-annotated because more specific terms capture the biology more accurately.
GO:0016887 ATP hydrolysis activity
ISO
GO_REF:0000121
ACCEPT
Summary: Manual review: ATP hydrolysis activity is consistent with known biology of Hspa8.
Reason: Retained as supported or plausible for this gene and evidence context.
GO:0051726 regulation of cell cycle
ISO
GO_REF:0000121
KEEP AS NON CORE
Summary: Manual review: regulation of cell cycle may be context-dependent or peripheral for Hspa8.
Reason: Kept as non-core to preserve potentially valid context-specific annotation without elevating it to core function.

Core Functions

ATP-dependent (EC 3.6.4.10) protein-folding chaperone activity: cyclic, nucleotide-state-dependent binding and release of unfolded/misfolded client polypeptides to assist protein folding, refolding, unfolding and disassembly of protein complexes, with the ATPase cycle regulated by J-domain (HSP40/DNAJ) co-chaperones and nucleotide-exchange factors.

Molecular Function:
protein folding chaperone
Cellular Locations:
Supporting Evidence:
  • file:rat/Hspa8/Hspa8-deep-research-falcon.md
    Hspa8/HSC70 is a **constitutive** HSP70-family **ATPase chaperone** that supports proteostasis by assisting **protein folding/refolding**, **unfolding**, and **disassembly of protein complexes**
  • file:rat/Hspa8/Hspa8-deep-research-falcon.md
    nucleotide state controls substrate affinity (ATP-bound low affinity/fast exchange; ADP-bound high affinity/slow exchange)
  • file:rat/Hspa8/Hspa8-deep-research-falcon.md
    J-domain proteins (HSP40/DNAJ)** stimulate ATP hydrolysis through engagement of the nucleotide-binding domain, and nucleotide exchange factors reset the cycle by promoting ADP→ATP exchange

Clathrin-uncoating ATPase activity: HSC70, recruited by the J-domain co-chaperone auxilin (DNAJC6), hydrolyzes ATP to disassemble clathrin coats from clathrin-coated vesicles, recycling the pool of free clathrin in clathrin-mediated endocytosis and synaptic vesicle recycling.

Directly Involved In:
Cellular Locations:
Supporting Evidence:
  • file:rat/Hspa8/Hspa8-deep-research-falcon.md
    the **J-domain co-chaperone auxilin (DNAJC6)** recruits HSC70 to CCVs and **stimulates HSC70 ATPase activity** to disassemble clathrin coats

Chaperone-mediated autophagy: cytosolic HSC70 recognizes KFERQ-like pentapeptide motifs in soluble client proteins and delivers them to LAMP-2A at the lysosomal membrane; lysosomal HSC70 supports substrate translocation into the lysosomal lumen for degradation. In this role HSC70 acts as a cargo selector/adaptor, bringing the client protein together with the LAMP-2A receptor.

Supporting Evidence:
  • file:rat/Hspa8/Hspa8-deep-research-falcon.md
    recognition of KFERQ-like pentapeptide motifs** in client proteins targeted for **chaperone-mediated autophagy (CMA)**
  • file:rat/Hspa8/Hspa8-deep-research-falcon.md
    Lysosomal HSC70 (Lys‑HSC70)** facilitates **substrate translocation** into the lysosome lumen

References

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Deep Research

Falcon

(Hspa8-deep-research-falcon.md)

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📚 Additional Documentation

Bioreason Rl Predictions

(Hspa8-bioreason-rl-predictions.md)

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Bioreason Rl Review

(Hspa8-bioreason-rl-review.md)

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📄 View Raw YAML

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