Ligand gap check: results

Generated by scripts/ligand_gap_check.py and scripts/triage.py; do not edit by hand.
GO version: releases/2026-07-26. GAFs: current.geneontology.org at run time.

Pathway terms per organism

Organism	Pathway terms used	No ligand gene	Ligand-activity genes	Ligand genes in no signaling term
human	237	78	571	22
mouse	234	86	602	48
rat	231	79	547	46
zebrafish	145	91	431	62
fly	74	39	117	1
yeast	6	6	4	0
pombe	4	4	4	0

Failures by triage category

Category	human	mouse	rat	zebrafish	fly	yeast	pombe
A: ligand counterpart not on the pathway	11	10	13	29	5	0	0
B: ligand counterpart only on a related pathway term	8	9	9	7	1	0	0
C: ligand counterpart on the pathway but lacks a ligand-activity MF	4	4	4	8	2	0	0
N: no same-symbol ligand counterpart annotated	0	5	1	7	0	0	0
U: no donor ligand available	3	7	7	1	4	6	2
X: no protein ligand expected (small molecule, light, microbial)	37	40	34	29	17	0	1
Y: antigen, contact or ligand-independent	11	11	11	10	4	0	0
O: suspect over-annotation	0	0	0	0	6	0	0
G: misapplied or generic term	4	0	0	0	0	0	1

Curation candidates (categories A, B, C)

Organism	Term	Category	Ligand lookups
human	netrin-activated signaling pathway (GO:0038007)	A	NTN1: A; NTN3: A
human	apolipoprotein A-I-mediated signaling pathway (GO:0038027)	A	APOA1: A (ligand MF)
human	interleukin-22-mediated signaling pathway (GO:0140865)	A	IL22: A (ligand MF)
human	complement component C5a signaling pathway (GO:0038178)	A	C5: A (ligand MF)
human	Fas signaling pathway (GO:0036337)	A	FASLG: A (ligand MF)
human	thyroid-stimulating hormone signaling pathway (GO:0038194)	A	CGA: A (ligand MF); TSHB: A (ligand MF)
human	interleukin-21-mediated signaling pathway (GO:0038114)	A	IL21: A
human	Kit signaling pathway (GO:0038109)	A	KITLG: A (ligand MF)
human	interleukin-8-mediated signaling pathway (GO:0038112)	A	CXCL8: A (ligand MF)
human	bombesin receptor signaling pathway (GO:0031989)	A	GRP: A (ligand MF); NMB: A (ligand MF)
human	macrophage migration inhibitory factor signaling pathway (GO:0035691)	A	MIF: A (ligand MF)
human	phospholipase C-activating angiotensin-activated signaling pathway (GO:0086097)	B	AGT: B (ligand MF) on angiotensin-activated signaling pathway
human	galanin-activated signaling pathway (GO:0090663)	B	GAL: B (ligand MF) on neuropeptide signaling pathway; GALP: A (ligand MF)
human	vascular endothelial growth factor receptor-1 signaling pathway (GO:0036323)	B	PGF: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway; VEGFA: B (ligand MF) on positive regulation of endothelial cell chemotaxis by VEGF-activated vascular endothelial growth factor receptor signaling pathway; VEGFB: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway
human	Notch signaling involved in heart development (GO:0061314)	B	DLL4: B (ligand MF) on Notch signaling pathway; JAG1: B (ligand MF) on Notch signaling pathway
human	fibroblast growth factor receptor signaling pathway involved in negative regulation of apoptotic process in bone marrow cell (GO:0035602)	B	FGF10: B (ligand MF) on fibroblast growth factor receptor signaling pathway; FGF2: B (ligand MF) on fibroblast growth factor receptor signaling pathway; FGF8: B (ligand MF) on fibroblast growth factor receptor signaling pathway
human	adenylate cyclase-inhibiting opioid receptor signaling pathway (GO:0031635)	B	PDYN: A (ligand MF); PENK: B (ligand MF) on G protein-coupled opioid receptor signaling pathway; POMC: B (ligand MF) on adenylate cyclase-activating G protein-coupled receptor signaling pathway
human	vascular endothelial growth factor receptor-3 signaling pathway (GO:0036325)	B	VEGFC: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway; VEGFD: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway
human	phospholipase C-activating endothelin receptor signaling pathway (GO:0160135)	B	EDN1: B (ligand MF) on endothelin receptor signaling pathway; EDN2: B (ligand MF) on endothelin receptor signaling pathway; EDN3: A (ligand MF)
human	collagen-activated signaling pathway (GO:0038065)	C	COL1A1: A; COL4A2: C
human	collagen-activated tyrosine kinase receptor signaling pathway (GO:0038063)	C	COL1A1: A; COL4A2: C
human	interleukin-34-mediated signaling pathway (GO:0061514)	C	IL34: C
human	fibroblast growth factor receptor apoptotic signaling pathway (GO:1902178)	C	FGF2: B (ligand MF) on fibroblast growth factor receptor signaling pathway; FGF8: B (ligand MF) on fibroblast growth factor receptor signaling pathway; FGFR3: C
mouse	G protein-coupled receptor signaling pathway, coupled to cyclic nucleotide second messenger (GO:0007187)	A	Agt: A (ligand MF); Ccl2: A (ligand MF); Npy: A (ligand MF)
mouse	G protein-coupled acetylcholine receptor signaling pathway (GO:0007213)	A	Agrn: A (ligand MF)
mouse	netrin-activated signaling pathway (GO:0038007)	A	Ntn1: A; Ntn3: A
mouse	apolipoprotein A-I-mediated signaling pathway (GO:0038027)	A	Apoa1: A (ligand MF)
mouse	brain-derived neurotrophic factor receptor signaling pathway (GO:0031547)	A	Bdnf: A (ligand MF)
mouse	bombesin receptor signaling pathway (GO:0031989)	A	Grp: A (ligand MF); Nmb: A (ligand MF)
mouse	G protein-coupled receptor signaling pathway coupled to cGMP nucleotide second messenger (GO:0007199)	A	Agt: A (ligand MF)
mouse	interleukin-22-mediated signaling pathway (GO:0140865)	A	Il22: A (ligand MF)
mouse	thyroid-stimulating hormone signaling pathway (GO:0038194)	A	Cga: A (ligand MF); Tshb: A (ligand MF)
mouse	macrophage migration inhibitory factor signaling pathway (GO:0035691)	A	Mif: A (ligand MF)
mouse	phospholipase C-activating angiotensin-activated signaling pathway (GO:0086097)	B	Agt: B (ligand MF) on angiotensin-activated signaling pathway
mouse	galanin-activated signaling pathway (GO:0090663)	B	Gal: B (ligand MF) on neuropeptide signaling pathway; Galp: A (ligand MF)
mouse	adenylate cyclase-inhibiting opioid receptor signaling pathway (GO:0031635)	B	Pdyn: A (ligand MF); Penk: B (ligand MF) on G protein-coupled opioid receptor signaling pathway; Pomc: B (ligand MF) on adenylate cyclase-activating G protein-coupled receptor signaling pathway
mouse	positive regulation of MAPKKK cascade by fibroblast growth factor receptor signaling pathway (GO:0090080)	B	Fgf15: B (ligand MF) on fibroblast growth factor receptor signaling pathway; Fgf21: B (ligand MF) on fibroblast growth factor receptor signaling pathway; Fgf23: B (ligand MF) on fibroblast growth factor receptor signaling pathway
mouse	vascular endothelial growth factor receptor-1 signaling pathway (GO:0036323)	B	Pgf: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway; Vegfa: B (ligand MF) on positive regulation of endothelial cell chemotaxis by VEGF-activated vascular endothelial growth factor receptor signaling pathway; Vegfb: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway
mouse	Notch signaling involved in heart development (GO:0061314)	B	Dll4: B (ligand MF) on Notch signaling pathway; Jag1: B (ligand MF) on Notch signaling pathway
mouse	fibroblast growth factor receptor signaling pathway involved in negative regulation of apoptotic process in bone marrow cell (GO:0035602)	B	Fgf10: B (ligand MF) on fibroblast growth factor receptor signaling pathway; Fgf2: B (ligand MF) on fibroblast growth factor receptor signaling pathway; Fgf8: B (ligand MF) on fibroblast growth factor receptor signaling pathway
mouse	vascular endothelial growth factor receptor-3 signaling pathway (GO:0036325)	B	Vegfc: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway; Vegfd: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway
mouse	phospholipase C-activating endothelin receptor signaling pathway (GO:0160135)	B	Edn1: B (ligand MF) on endothelin receptor signaling pathway; Edn2: B (ligand MF) on endothelin receptor signaling pathway; Edn3: A (ligand MF)
mouse	collagen-activated signaling pathway (GO:0038065)	C	Col1a1: A; Col4a2: C
mouse	collagen-activated tyrosine kinase receptor signaling pathway (GO:0038063)	C	Col1a1: A; Col4a2: C
mouse	interleukin-34-mediated signaling pathway (GO:0061514)	C	Il34: C
mouse	fibroblast growth factor receptor apoptotic signaling pathway (GO:1902178)	C	Fgf2: B (ligand MF) on fibroblast growth factor receptor signaling pathway; Fgf8: B (ligand MF) on fibroblast growth factor receptor signaling pathway; Fgfr3: C
rat	G protein-coupled receptor signaling pathway, coupled to cyclic nucleotide second messenger (GO:0007187)	A	Agt: A (ligand MF); Ccl2: A (ligand MF); Npy: A (ligand MF)
rat	G protein-coupled acetylcholine receptor signaling pathway (GO:0007213)	A	Agrn: A (ligand MF)
rat	netrin-activated signaling pathway (GO:0038007)	A	Ntn1: A; Ntn3: A
rat	apolipoprotein A-I-mediated signaling pathway (GO:0038027)	A	Apoa1: A (ligand MF)
rat	brain-derived neurotrophic factor receptor signaling pathway (GO:0031547)	A	Bdnf: A (ligand MF)
rat	G protein-coupled receptor signaling pathway coupled to cGMP nucleotide second messenger (GO:0007199)	A	Agt: A (ligand MF)
rat	complement component C5a signaling pathway (GO:0038178)	A	C5: A (ligand MF)
rat	Fas signaling pathway (GO:0036337)	A	Faslg: A (ligand MF)
rat	bombesin receptor signaling pathway (GO:0031989)	A	Grp: A (ligand MF); Nmb: A (ligand MF)
rat	interleukin-22-mediated signaling pathway (GO:0140865)	A	Il22: A (ligand MF)
rat	Kit signaling pathway (GO:0038109)	A	Kitlg: A (ligand MF)
rat	thyroid-stimulating hormone signaling pathway (GO:0038194)	A	Cga: A (ligand MF); Tshb: A (ligand MF)
rat	macrophage migration inhibitory factor signaling pathway (GO:0035691)	A	Mif: A (ligand MF)
rat	phospholipase C-activating angiotensin-activated signaling pathway (GO:0086097)	B	Agt: B (ligand MF) on angiotensin-activated signaling pathway
rat	galanin-activated signaling pathway (GO:0090663)	B	Gal: B (ligand MF) on neuropeptide signaling pathway; Galp: A (ligand MF)
rat	Notch signaling involved in heart development (GO:0061314)	B	Dll4: B (ligand MF) on Notch signaling pathway; Jag1: B (ligand MF) on Notch signaling pathway
rat	adenylate cyclase-inhibiting opioid receptor signaling pathway (GO:0031635)	B	Pdyn: A (ligand MF); Penk: B (ligand MF) on G protein-coupled opioid receptor signaling pathway; Pomc: B (ligand MF) on adenylate cyclase-activating G protein-coupled receptor signaling pathway
rat	vascular endothelial growth factor receptor-1 signaling pathway (GO:0036323)	B	Pgf: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway; Vegfa: B (ligand MF) on positive regulation of endothelial cell chemotaxis by VEGF-activated vascular endothelial growth factor receptor signaling pathway; Vegfb: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway
rat	fibroblast growth factor receptor signaling pathway involved in negative regulation of apoptotic process in bone marrow cell (GO:0035602)	B	Fgf10: B (ligand MF) on fibroblast growth factor receptor signaling pathway; Fgf2: B (ligand MF) on fibroblast growth factor receptor signaling pathway; Fgf8: B (ligand MF) on fibroblast growth factor receptor signaling pathway
rat	vascular endothelial growth factor receptor-3 signaling pathway (GO:0036325)	B	Vegfc: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway; Vegfd: B (ligand MF) on vascular endothelial growth factor receptor signaling pathway
rat	phospholipase C-activating endothelin receptor signaling pathway (GO:0160135)	B	Edn1: B (ligand MF) on endothelin receptor signaling pathway; Edn2: B (ligand MF) on endothelin receptor signaling pathway; Edn3: A (ligand MF)
rat	platelet-derived growth factor receptor-alpha signaling pathway (GO:0035790)	B	Pdgfa: B (ligand MF) on platelet-derived growth factor receptor signaling pathway
rat	collagen-activated signaling pathway (GO:0038065)	C	Col1a1: A; Col4a2: C
rat	collagen-activated tyrosine kinase receptor signaling pathway (GO:0038063)	C	Col1a1: A; Col4a2: C
rat	interleukin-34-mediated signaling pathway (GO:0061514)	C	Il34: C
rat	urokinase plasminogen activator signaling pathway (GO:0038195)	C	Plau: A (ligand MF); Slurp1: C
zebrafish	phospholipase C-activating G protein-coupled receptor signaling pathway (GO:0007200)	A	agt: A; calca: A (ligand MF); ccka: A (ligand MF); cckb: A (ligand MF); edn2: A (ligand MF); hcrt: A (ligand MF); il2: A; kiss1: A; nmu: A; tac1: A
zebrafish	G protein-coupled receptor signaling pathway, coupled to cyclic nucleotide second messenger (GO:0007187)	A	agt: A; npy: A (ligand MF)
zebrafish	insulin receptor signaling pathway (GO:0008286)	A	igf1: A (ligand MF); igf2b: A (ligand MF); igf2a: A (ligand MF); insb: A (ligand MF); ins: A (ligand MF)
zebrafish	tumor necrosis factor-mediated signaling pathway (GO:0033209)	A	tnfa: A (ligand MF); tnfb: A (ligand MF); tnfsf11: A (ligand MF); tnfsf13b: A (ligand MF)
zebrafish	G protein-coupled acetylcholine receptor signaling pathway (GO:0007213)	A	agrn: A
zebrafish	lipopolysaccharide-mediated signaling pathway (GO:0031663)	A	il1b: A (ligand MF); tnfa: A (ligand MF); tnfb: A (ligand MF)
zebrafish	G protein-coupled opioid receptor signaling pathway (GO:0038003)	A	penkb: A (ligand MF); penka: A (ligand MF)
zebrafish	netrin-activated signaling pathway (GO:0038007)	A	ntn1b: A; ntn1a: A
zebrafish	endothelin receptor signaling pathway (GO:0086100)	A	edn1: A (ligand MF); edn2: A (ligand MF)
zebrafish	collagen-activated tyrosine kinase receptor signaling pathway (GO:0038063)	A	col1a1b: A; col1a1a: A; col4a2: A
zebrafish	interleukin-15-mediated signaling pathway (GO:0035723)	A	il15: A (ligand MF)
zebrafish	thyroid-stimulating hormone signaling pathway (GO:0038194)	A	cga: A (ligand MF); tshba: A (ligand MF); tshbb: A (ligand MF)
zebrafish	Kit signaling pathway (GO:0038109)	A	kitlga: A; kitlgb: A
zebrafish	interleukin-1-mediated signaling pathway (GO:0070498)	A	il1b: A (ligand MF)
zebrafish	reelin-mediated signaling pathway (GO:0038026)	A	reln: A
zebrafish	TRAIL-activated apoptotic signaling pathway (GO:0036462)	A	tnfsf10: A (ligand MF)
zebrafish	macrophage colony-stimulating factor signaling pathway (GO:0038145)	A	csf1b: A (ligand MF); csf1a: A (ligand MF)
zebrafish	Roundabout signaling pathway (GO:0035385)	A	slit2: A
zebrafish	CXCL12-activated CXCR4 signaling pathway (GO:0038160)	A	cxcl12a: A (ligand MF); cxcl12b: A (ligand MF)
zebrafish	gastric inhibitory peptide signaling pathway (GO:0038192)	A	gip: A
zebrafish	apolipoprotein A-I-mediated signaling pathway (GO:0038027)	A	apoa1b: A; apoa1a: A
zebrafish	complement component C5a signaling pathway (GO:0038178)	A	c5: A
zebrafish	Fas signaling pathway (GO:0036337)	A	faslg: A (ligand MF)
zebrafish	thrombopoietin-mediated signaling pathway (GO:0038163)	A	thpo: A (ligand MF)
zebrafish	follicle-stimulating hormone signaling pathway (GO:0042699)	A	cga: A (ligand MF); fshb: A (ligand MF); slit2: A
zebrafish	brain-derived neurotrophic factor receptor signaling pathway (GO:0031547)	A	bdnf: A (ligand MF)
zebrafish	cholecystokinin signaling pathway (GO:0038188)	A	ccka: A (ligand MF); cckb: A (ligand MF)
zebrafish	luteinizing hormone signaling pathway (GO:0042700)	A	lhb: A (ligand MF)
zebrafish	chemokine (C-C motif) ligand 19 signaling pathway (GO:0038115)	A	ccl19b: A (ligand MF)
zebrafish	growth hormone receptor signaling pathway via JAK-STAT (GO:0060397)	B	gh1: B (ligand MF) on growth hormone receptor signaling pathway
zebrafish	platelet-derived growth factor receptor-beta signaling pathway (GO:0035791)	B	pdgfaa: B (ligand MF) on platelet-derived growth factor receptor signaling pathway; pdgfab: B (ligand MF) on platelet-derived growth factor receptor signaling pathway; pdgfbb: B (ligand MF) on platelet-derived growth factor receptor signaling pathway; pdgfba: B on platelet-derived growth factor receptor signaling pathway
zebrafish	neurotrophin TRK receptor signaling pathway (GO:0048011)	B	ngfb: B (ligand MF) on nerve growth factor signaling pathway; ngfa: B (ligand MF) on nerve growth factor signaling pathway
zebrafish	galanin-activated signaling pathway (GO:0090663)	B	gal: B (ligand MF) on neuropeptide signaling pathway
zebrafish	Notch signaling involved in heart development (GO:0061314)	B	dll4: B on Notch signaling pathway; jag1a: B (ligand MF) on Notch signaling pathway; jag1b: B on Notch signaling pathway
zebrafish	platelet-derived growth factor receptor-alpha signaling pathway (GO:0035790)	B	pdgfaa: B (ligand MF) on platelet-derived growth factor receptor signaling pathway; pdgfab: B (ligand MF) on platelet-derived growth factor receptor signaling pathway
zebrafish	smoothened signaling pathway involved in ventral spinal cord patterning (GO:0021910)	B	shha: B on smoothened signaling pathway; shhb: B on smoothened signaling pathway
zebrafish	ephrin receptor signaling pathway (GO:0048013)	C	efna1a: C; efna1b: C; efna5a: C; efna5b: C; efnb2b: C; efnb2a: C; epha7: C
zebrafish	non-canonical Wnt signaling pathway (GO:0035567)	C	sfrp2: C; wnt11: B on Wnt signaling pathway; wnt3a: B (ligand MF) on Wnt signaling pathway; wnt4b: B (ligand MF) on Wnt signaling pathway; wnt4: B (ligand MF) on Wnt signaling pathway; wnt5a: B (ligand MF) on Wnt signaling pathway; wnt7aa: B (ligand MF) on Wnt signaling pathway; wnt7ab: B (ligand MF) on Wnt signaling pathway; wnt7ba: B (ligand MF) on Wnt signaling pathway; wnt7bb: B (ligand MF) on Wnt signaling pathway; wnt9b: B (ligand MF) on Wnt signaling pathway
zebrafish	tachykinin receptor signaling pathway (GO:0007217)	C	tac1: C; tac3a: C; tac3b: C; tac4: A
zebrafish	thrombin-activated receptor signaling pathway (GO:0070493)	C	f2: A; f2rl2: C
zebrafish	angiotensin-activated signaling pathway (GO:0038166)	C	agt: C
zebrafish	hepatocyte growth factor receptor signaling pathway (GO:0048012)	C	hgfa: C
zebrafish	Tie signaling pathway (GO:0048014)	C	angpt1: C; angpt2b: A; angpt2a: C; angpt4: A
zebrafish	interleukin-11-mediated signaling pathway (GO:0038154)	C	cntfr: C; il11a: A (ligand MF); il11b: A (ligand MF); il6st: A
fly	neurotrophin signaling pathway (GO:0038179)	A	NT1: A (ligand MF); spz: A (ligand MF); spz5: A (ligand MF)
fly	extrinsic apoptotic signaling pathway (GO:0097191)	A	egr: A (ligand MF)
fly	netrin-activated signaling pathway (GO:0038007)	A	NetA: A; NetB: A
fly	collagen-activated tyrosine kinase receptor signaling pathway (GO:0038063)	A	Col4a1: A; vkg: A
fly	insulin-like growth factor receptor signaling pathway (GO:0048009)	A	Ilp2: A (ligand MF); Ilp3: A (ligand MF); Ilp5: A (ligand MF)
fly	Frizzled Nuclear Import pathway (GO:0140709)	B	wg: B (ligand MF) on Wnt signaling pathway
fly	ephrin receptor signaling pathway (GO:0048013)	C	Ephrin: C
fly	Roundabout signaling pathway (GO:0035385)	C	sli: C

Ligand-activity genes in no signaling term

• human: ALKBH1, CCL17, CCL27, CCL28, CKLF, CXCL16, CXCL2, CXCL3, DEFB103A, DEFB104A, DEFB109B, DEFB110, DEFB114, DEFB130A, DEFB130B, DEFB133, DPP4, LGALS3, MT-RNR2, PF4V1, PPBP, S100A4
• mouse: AI182371, Alkbh1, Ccl17, Ccl28, Cd48, Chlsn, Cklf, Colec10, Cxcl1, Cxcl14, Cxcl15, Cxcl16, Cxcl2, Cxcl3, Defb14, Defb3, Defb33, Defb4, Defb46, Defb47, Defb48, Defb5, Defb6, Defb7, Defb8, Dpp4, Enho, Erfe, Fgf12, Fgf14, Gm6040, Hamp, Hc, Hmgb2, Itpripl1, Kitl, Lama1, Lgals3, Ppbp, Reg3a, Rln1, S100a4, S100a7a, Saa3, Sectm1a, Sectm1b, Stc2, mFGF13/iso:2
• rat: AABR07001348.1, AABR07016805.1, Alkbh1, C5, Ccl17, Ccl22, Ccl27, Ccl28, Cd48, Cklf, Colec10, Cxcl1, Cxcl14, Cxcl16, Cxcl2, Cxcl3, Defb14, Defb17, Defb3, Defb33, Defb4, Defb41, Defb42, Defb5, Dpp4, ENSRNOG00000070971, Fgf12, Fgf13, Fgf14, H60al, Hamp, Hmgb2, Hmgb2l1, Hmgb2l2, LOC108353456, LOC120093164, LOC120102174, Lama1, Lgals3, Mifl2, Mifl3, Mifl4, Ppbp, Rln1, S100a4, Stc2
• zebrafish: LOC567978, cckb, ccl19a.2, ccl19b, ccl20a.3, ccl20a.4, ccl20b, ccl20l, ccl25b, ccl27a, ccl27b, ccl32b.3, ccl33.2, ccl33.3, ccl34a.3, ccl34a.4, ccl34b.1, ccl34b.4, ccl34b.9, ccl35.1, ccl35.2, ccl36.1, ccl38.1, ccl38a.3, ccl38a.4, ccl38a.5, ccl39.1, ccl39.2, ccl39.3, ccl44, cxcl11.1, cxcl11.3, cxcl11.4, cxcl11.6, cxcl11.8, cxcl12a, cxcl12b, cxcl13, cxcl14, cxcl18a.1, cxcl18b, cxcl19, cxcl32b.1, cxcl8a, cxcl8b.3, cxl34b.11, cxl34c, edn3b, fam19a1, fbn3, fgl1, ghrh, grem2a, grem2b, igfbp7, izumo1, kazald2, osgn1, si:ch73-330k17.3, tafa1a, tnfsf13, vip
• fly: orion
• yeast: none
• pombe: none