USP10

UniProt ID: Q14694
Organism: Homo sapiens
Review Status: COMPLETE
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Gene Description

USP10 encodes a ubiquitin-specific cysteine deubiquitinase that removes ubiquitin from selected substrates including p53/TP53, CFTR, Beclin1-complex components, LC3B, and 40S ribosomal proteins. Its direct roles in protein homeostasis include protein deubiquitination, rescue of ubiquitinated stalled 40S ribosomal subunits, and regulation of autophagy/stress-granule signaling through LC3B, Beclin1, and G3BP contexts.

Proposed New Ontology Terms

ATG8-family protein deubiquitination

Definition: A protein deubiquitination process in which ubiquitin is removed from an ATG8-family protein, thereby modulating ATG8 protein stability, lipidated ATG8 abundance, or autophagic activity.

Justification: USP10 directly deubiquitinates LC3B and regulates LC3B abundance/autophagic activity, but current GOA can only capture broad protein deubiquitination or regulation of autophagy.

Parent term: protein deubiquitination

Supporting Evidence:

40S ribosomal protein deubiquitination during ribosome quality control

Definition: A monoubiquitinated protein deubiquitination process in which ubiquitin is removed from proteins of a cytosolic 40S ribosomal subunit during ribosome-associated quality control, preventing or reversing programmed degradation of the modified 40S subunit.

Justification: USP10 removes ubiquitin from RPS2/uS5, RPS3/uS3, and RPS10/eS10 in stalled or compromised 40S ribosomal-subunit contexts. Existing GO terms capture either broad monoubiquitinated protein deubiquitination or rescue of stalled cytosolic ribosome, but not the substrate-specific deubiquitination event.

Parent term: monoubiquitinated protein deubiquitination

Supporting Evidence:

Existing Annotations Review

GO Term Evidence Action Reason
GO:0010506 regulation of autophagy
IBA
GO_REF:0000033
KEEP AS NON CORE
Summary: USP10 regulates autophagy through Beclin1/Vps34 complexes and LC3B deubiquitination, but this is one substrate pathway of the broader DUB function.
Reason: Keep as non-core. The process is well supported and relevant to the PN ATG8 context, but the molecular function should remain protein deubiquitination rather than a generic autophagy-regulator identity.
Supporting Evidence:
PMID:21962518
two ubiquitin-specific peptidases, USP10 and USP13, that target the Beclin1 subunit of Vps34 complexes
PMID:33577797
LC3B ubiquitination is reversed by the action of the deubiquitinating enzyme USP10
PMID:33577797
LC3B and autophagic activity are controlled through cycles of LC3B ubiquitination and deubiquitination
GO:0030330 DNA damage response, signal transduction by p53 class mediator
IBA
GO_REF:0000033
KEEP AS NON CORE
Summary: USP10 activates p53 signaling after DNA damage by deubiquitinating/stabilizing p53 and translocating to the nucleus.
Reason: Keep as non-core. The annotation is supported but represents a substrate-specific signaling outcome of USP10 DUB activity.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0005769 early endosome
IBA
GO_REF:0000033
ACCEPT
Summary: USP10 localizes to early endosomes where it deubiquitinates CFTR and supports endocytic recycling.
Reason: Accept as a supported active compartment for a direct USP10 substrate context.
Supporting Evidence:
PMID:19398555
USP10 is located in early endosomes and regulates the deubiquitination of CFTR
PMID:19398555
facilitating the deubiquitination of CFTR in early endosomes and thereby enhancing the endocytic recycling of CFTR
GO:0004843 cysteine-type deubiquitinase activity
IEA
GO_REF:0000120
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
Reactome:R-HSA-5688426
Deubiquitinating enzymes (DUBs) catalyze the removal of Ub and regulate Ub-mediated pathways
GO:0005634 nucleus
IEA
GO_REF:0000044
ACCEPT
Summary: USP10 translocates to or acts in the nucleus in p53 and T-bet substrate-stability contexts.
Reason: Accept as a supported location, while treating the specific nuclear signaling outputs as substrate/context-specific rather than the sole core function.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0005737 cytoplasm
IEA
GO_REF:0000044
ACCEPT
Summary: USP10 is a cytoplasmic/cytosolic DUB and several core substrate contexts occur in the cytoplasm.
Reason: Accept as a supported cellular location for USP10 deubiquitination, including p53 homeostasis, RQC, and immune-signaling substrates.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0005769 early endosome
IEA
GO_REF:0000044
ACCEPT
Summary: USP10 localizes to early endosomes where it deubiquitinates CFTR and supports endocytic recycling.
Reason: Accept as a supported active compartment for a direct USP10 substrate context.
Supporting Evidence:
PMID:19398555
USP10 is located in early endosomes and regulates the deubiquitination of CFTR
PMID:19398555
facilitating the deubiquitination of CFTR in early endosomes and thereby enhancing the endocytic recycling of CFTR
GO:0016579 protein deubiquitination
IEA
GO_REF:0000002
ACCEPT
Summary: Protein deubiquitination is the broad biological process that best summarizes USP10 catalytic action across multiple substrates.
Reason: Accept as core. Substrate-specific evidence supports deubiquitination of p53, CFTR, ribosomal proteins, LC3B, and immune-signaling substrates.
Supporting Evidence:
Reactome:R-HSA-5688426
Deubiquitinating enzymes (DUBs) catalyze the removal of Ub and regulate Ub-mediated pathways
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:19398555
USP10 is located in early endosomes and regulates the deubiquitination of CFTR
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
GO:0005515 protein binding
IPI
PMID:19615732
Defining the human deubiquitinating enzyme interaction lands...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:21455491
A Pseudomonas aeruginosa toxin that hijacks the host ubiquit...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:24270572
USP10 inhibits genotoxic NF-ΞΊB activation by MCPIP1-facilita...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:24981860
Human-chromatin-related protein interactions identify a deme...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:25416956
A proteome-scale map of the human interactome network.
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:29892012
An interactome perturbation framework prioritizes damaging m...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:31515488
Extensive disruption of protein interactions by genetic vari...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:32296183
A reference map of the human binary protein interactome.
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:32814053
Interactome Mapping Provides a Network of Neurodegenerative ...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:33495715
SARS-CoV-2 nucleocapsid protein phase separates with G3BPs t...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:33961781
Dual proteome-scale networks reveal cell-specific remodeling...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:34799561
Large scale discovery of coronavirus-host factor protein int...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:34901782
SARS-CoV-2 nucleocapsid protein binds host mRNAs and attenua...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:35156780
CFTR interactome mapping using the mammalian membrane two-hy...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:35271311
OpenCell: Endogenous tagging for the cartography of human ce...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:36012204
Differential CFTR-Interactome Proximity Labeling Procedures ...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:40205054
Multimodal cell maps as a foundation for structural and func...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005737 cytoplasm
NAS
PMID:23279204
Both G3BP1 and G3BP2 contribute to stress granule formation.
ACCEPT
Summary: USP10 is a cytoplasmic/cytosolic DUB and several core substrate contexts occur in the cytoplasm.
Reason: Accept as a supported cellular location for USP10 deubiquitination, including p53 homeostasis, RQC, and immune-signaling substrates.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0005737 cytoplasm
IPI
PMID:31981475
The G3BP1-Family-USP10 Deubiquitinase Complex Rescues Ubiqui...
ACCEPT
Summary: USP10 is a cytoplasmic/cytosolic DUB and several core substrate contexts occur in the cytoplasm.
Reason: Accept as a supported cellular location for USP10 deubiquitination, including p53 homeostasis, RQC, and immune-signaling substrates.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
GO:0016579 protein deubiquitination
IDA
PMID:34469731
iRQC, a surveillance pathway for 40S ribosomal quality contr...
ACCEPT
Summary: Protein deubiquitination is the broad biological process that best summarizes USP10 catalytic action across multiple substrates.
Reason: Accept as core. Substrate-specific evidence supports deubiquitination of p53, CFTR, ribosomal proteins, LC3B, and immune-signaling substrates.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0016579 protein deubiquitination
NAS
PMID:34469731
iRQC, a surveillance pathway for 40S ribosomal quality contr...
ACCEPT
Summary: Protein deubiquitination is the broad biological process that best summarizes USP10 catalytic action across multiple substrates.
Reason: Accept as core. Substrate-specific evidence supports deubiquitination of p53, CFTR, ribosomal proteins, LC3B, and immune-signaling substrates.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0062030 negative regulation of stress granule assembly
IDA
PMID:27022092
G3BP-Caprin1-USP10 complexes mediate stress granule condensa...
KEEP AS NON CORE
Summary: USP10 binding to G3BP inhibits stress granule formation in a 40S-associated condensate context.
Reason: Keep as non-core. This is a supported stress-granule regulatory role but not the core catalytic DUB activity.
Supporting Evidence:
PMID:27022092
Caprin binding promotes, but USP10 binding inhibits, SG formation
PMID:27022092
G3BP interacts with 40S ribosomal subunits through its RGG motif
PMID:32302570
competitive binding of unconnected proteins disengages networks and prevents LLPS
GO:0062030 negative regulation of stress granule assembly
NAS
PMID:27022092
G3BP-Caprin1-USP10 complexes mediate stress granule condensa...
KEEP AS NON CORE
Summary: USP10 binding to G3BP inhibits stress granule formation in a 40S-associated condensate context.
Reason: Keep as non-core. This is a supported stress-granule regulatory role but not the core catalytic DUB activity.
Supporting Evidence:
PMID:27022092
Caprin binding promotes, but USP10 binding inhibits, SG formation
PMID:27022092
G3BP interacts with 40S ribosomal subunits through its RGG motif
PMID:32302570
competitive binding of unconnected proteins disengages networks and prevents LLPS
GO:0005654 nucleoplasm
IDA
GO_REF:0000052
ACCEPT
Summary: Nucleoplasm is a supported location for USP10 in PCNA/TLS and TP53 deubiquitination Reactome contexts.
Reason: Accept as a nuclear subcompartment location for substrate-specific USP10 activities.
Supporting Evidence:
Reactome:R-HSA-5653766
Ubiquitin protease USP10 binds doubly ISGylated and monoubiquitinated PCNA
Reactome:R-HSA-5653770
USP10 acts as a ubiquitin protease to remove ubiquitin from lysine K164 residue of doubly ISGylated PCNA
Reactome:R-HSA-5653770
Deubiquitination of PCNA by USP10 causes dissociation of Y family DNA damage bypass polymerases
GO:0005829 cytosol
IDA
GO_REF:0000052
ACCEPT
Summary: Cytosol is a supported compartment for USP10 deubiquitinase activity and substrate regulation.
Reason: Accept as a supported cytosolic location for USP10 activity; cytosolic p53 regulation and ribosome-quality-control evidence are consistent with this assignment.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
GO:0016579 protein deubiquitination
TAS
Reactome:R-HSA-5688426
ACCEPT
Summary: Protein deubiquitination is the broad biological process that best summarizes USP10 catalytic action across multiple substrates.
Reason: Accept as core. Substrate-specific evidence supports deubiquitination of p53, CFTR, ribosomal proteins, LC3B, and immune-signaling substrates.
Supporting Evidence:
Reactome:R-HSA-5688426
Deubiquitinating enzymes (DUBs) catalyze the removal of Ub and regulate Ub-mediated pathways
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:19398555
USP10 is located in early endosomes and regulates the deubiquitination of CFTR
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
GO:0019985 translesion synthesis
TAS
Reactome:R-HSA-110313
KEEP AS NON CORE
Summary: Reactome places USP10 in the PCNA deubiquitination step that terminates/limits translesion synthesis.
Reason: Keep as non-core. This is a specific nuclear DNA-damage-bypass substrate context, not the defining USP10 function.
Supporting Evidence:
Reactome:R-HSA-5653766
Ubiquitin protease USP10 binds doubly ISGylated and monoubiquitinated PCNA
Reactome:R-HSA-5653770
USP10 acts as a ubiquitin protease to remove ubiquitin from lysine K164 residue of doubly ISGylated PCNA
Reactome:R-HSA-5653770
Deubiquitination of PCNA by USP10 causes dissociation of Y family DNA damage bypass polymerases
GO:0004843 cysteine-type deubiquitinase activity
TAS
Reactome:R-HSA-5653770
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
Reactome:R-HSA-5653766
Ubiquitin protease USP10 binds doubly ISGylated and monoubiquitinated PCNA
Reactome:R-HSA-5653770
USP10 acts as a ubiquitin protease to remove ubiquitin from lysine K164 residue of doubly ISGylated PCNA
Reactome:R-HSA-5653770
Deubiquitination of PCNA by USP10 causes dissociation of Y family DNA damage bypass polymerases
GO:0004843 cysteine-type deubiquitinase activity
TAS
Reactome:R-HSA-5689973
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
Reactome:R-HSA-5689973
USP10 specifically deubiquitinate p53 and not MDM2
GO:0004843 cysteine-type deubiquitinase activity
TAS
Reactome:R-HSA-6782106
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
Reactome:R-HSA-6782106
USP10 deubiquitinates CFTR in early endosomes thereby enhancing its endocytic recycling
GO:0004843 cysteine-type deubiquitinase activity
EXP
PMID:32011234
Distinct regulatory ribosomal ubiquitylation events are reve...
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
Reactome:R-HSA-5688426
Deubiquitinating enzymes (DUBs) catalyze the removal of Ub and regulate Ub-mediated pathways
GO:0005737 cytoplasm
EXP
PMID:37582970
MAVS-loaded unanchored Lys63-linked polyubiquitin chains act...
ACCEPT
Summary: USP10 is a cytoplasmic/cytosolic DUB and several core substrate contexts occur in the cytoplasm.
Reason: Accept as a supported cellular location for USP10 deubiquitination, including p53 homeostasis, RQC, and immune-signaling substrates.
Supporting Evidence:
PMID:37582970
USP10 as a direct DUB that removes unanchored K63-linked polyubiquitin chains from MAVS
PMID:37582970
USP10 attenuates RIG-I-mediated MAVS aggregation and the production of type I interferon
GO:0045087 innate immune response
IDA
PMID:37023208
Immune evasion strategy involving propionylation by the KSHV...
KEEP AS NON CORE
Summary: USP10 modulates innate immune signaling by removing unanchored K63-linked ubiquitin chains from MAVS; the seeded KSHV/SIRT6 paper is indirect for this term.
Reason: Keep as non-core with added MAVS evidence. USP10 has a real RLR/MAVS immune-regulatory role, but the original reference mainly describes viral interference with SIRT6-USP10 regulation.
Supporting Evidence:
PMID:37582970
USP10 as a direct DUB that removes unanchored K63-linked polyubiquitin chains from MAVS
PMID:37582970
USP10 attenuates RIG-I-mediated MAVS aggregation and the production of type I interferon
GO:0004843 cysteine-type deubiquitinase activity
IDA
PMID:31981475
The G3BP1-Family-USP10 Deubiquitinase Complex Rescues Ubiqui...
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0004843 cysteine-type deubiquitinase activity
IDA
PMID:34348161
The E3 ubiquitin ligase RNF10 modifies 40S ribosomal subunit...
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0004843 cysteine-type deubiquitinase activity
IDA
PMID:34469731
iRQC, a surveillance pathway for 40S ribosomal quality contr...
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0022626 cytosolic ribosome
IDA
PMID:34348161
The E3 ubiquitin ligase RNF10 modifies 40S ribosomal subunit...
ACCEPT
Summary: USP10 acts at cytosolic 40S ribosomal subunits during ribosome-associated quality control.
Reason: Accept as an active site/context for the ribosomal deubiquitination function.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0022626 cytosolic ribosome
IDA
PMID:34469731
iRQC, a surveillance pathway for 40S ribosomal quality contr...
ACCEPT
Summary: USP10 acts at cytosolic 40S ribosomal subunits during ribosome-associated quality control.
Reason: Accept as an active site/context for the ribosomal deubiquitination function.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0035520 monoubiquitinated protein deubiquitination
IDA
PMID:34348161
The E3 ubiquitin ligase RNF10 modifies 40S ribosomal subunit...
ACCEPT
Summary: USP10 directly removes monoubiquitin from 40S ribosomal proteins in ribosome quality-control contexts.
Reason: Accept as a direct proteostasis function. RPS2/RPS3/RPS10 deubiquitination rescues modified 40S subunits from degradation.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0035520 monoubiquitinated protein deubiquitination
IDA
PMID:34469731
iRQC, a surveillance pathway for 40S ribosomal quality contr...
ACCEPT
Summary: USP10 directly removes monoubiquitin from 40S ribosomal proteins in ribosome quality-control contexts.
Reason: Accept as a direct proteostasis function. RPS2/RPS3/RPS10 deubiquitination rescues modified 40S subunits from degradation.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0072344 rescue of stalled cytosolic ribosome
IDA
PMID:31981475
The G3BP1-Family-USP10 Deubiquitinase Complex Rescues Ubiqui...
ACCEPT
Summary: USP10-containing G3BP complexes rescue ubiquitinated stalled 40S subunits from programmed degradation.
Reason: Accept as a direct ribosome-associated quality-control role in the proteostasis network.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0072344 rescue of stalled cytosolic ribosome
IDA
PMID:34348161
The E3 ubiquitin ligase RNF10 modifies 40S ribosomal subunit...
ACCEPT
Summary: USP10-containing G3BP complexes rescue ubiquitinated stalled 40S subunits from programmed degradation.
Reason: Accept as a direct ribosome-associated quality-control role in the proteostasis network.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0072344 rescue of stalled cytosolic ribosome
IDA
PMID:34469731
iRQC, a surveillance pathway for 40S ribosomal quality contr...
ACCEPT
Summary: USP10-containing G3BP complexes rescue ubiquitinated stalled 40S subunits from programmed degradation.
Reason: Accept as a direct ribosome-associated quality-control role in the proteostasis network.
Supporting Evidence:
PMID:31981475
G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
PMID:34348161
These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
PMID:34469731
USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
GO:0005515 protein binding
IPI
PMID:27022092
G3BP-Caprin1-USP10 complexes mediate stress granule condensa...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:32302570
Competing Protein-RNA Interaction Networks Control Multiphas...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:36279435
Yin and yang regulation of stress granules by Caprin-1.
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0062030 negative regulation of stress granule assembly
IDA
PMID:32302570
Competing Protein-RNA Interaction Networks Control Multiphas...
KEEP AS NON CORE
Summary: USP10 binding to G3BP inhibits stress granule formation in a 40S-associated condensate context.
Reason: Keep as non-core. This is a supported stress-granule regulatory role but not the core catalytic DUB activity.
Supporting Evidence:
PMID:27022092
Caprin binding promotes, but USP10 binding inhibits, SG formation
PMID:27022092
G3BP interacts with 40S ribosomal subunits through its RGG motif
PMID:32302570
competitive binding of unconnected proteins disengages networks and prevents LLPS
GO:0140678 molecular function inhibitor activity
IDA
PMID:27022092
G3BP-Caprin1-USP10 complexes mediate stress granule condensa...
KEEP AS NON CORE
Summary: USP10 acts as an inhibitor in the G3BP stress-granule assembly network.
Reason: Keep as non-core. The MF-inhibitor annotation captures a real G3BP regulatory effect, but the more informative biological-process annotation is negative regulation of stress granule assembly.
Supporting Evidence:
PMID:27022092
Caprin binding promotes, but USP10 binding inhibits, SG formation
PMID:27022092
G3BP interacts with 40S ribosomal subunits through its RGG motif
PMID:32302570
competitive binding of unconnected proteins disengages networks and prevents LLPS
GO:0140678 molecular function inhibitor activity
IDA
PMID:32302570
Competing Protein-RNA Interaction Networks Control Multiphas...
KEEP AS NON CORE
Summary: USP10 acts as an inhibitor in the G3BP stress-granule assembly network.
Reason: Keep as non-core. The MF-inhibitor annotation captures a real G3BP regulatory effect, but the more informative biological-process annotation is negative regulation of stress granule assembly.
Supporting Evidence:
PMID:27022092
Caprin binding promotes, but USP10 binding inhibits, SG formation
PMID:27022092
G3BP interacts with 40S ribosomal subunits through its RGG motif
PMID:32302570
competitive binding of unconnected proteins disengages networks and prevents LLPS
GO:0005515 protein binding
IPI
PMID:23279204
Both G3BP1 and G3BP2 contribute to stress granule formation.
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005515 protein binding
IPI
PMID:24845384
Deubiquitination and stabilization of T-bet by USP10.
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005634 nucleus
IDA
PMID:24845384
Deubiquitination and stabilization of T-bet by USP10.
ACCEPT
Summary: USP10 translocates to or acts in the nucleus in p53 and T-bet substrate-stability contexts.
Reason: Accept as a supported location, while treating the specific nuclear signaling outputs as substrate/context-specific rather than the sole core function.
Supporting Evidence:
PMID:24845384
USP10, a carboxyl-terminal ubiquitin-processing protease, could interact with T-bet in the nucleus
PMID:24845384
Overexpression of USP10 directly inhibited T-bet ubiquitination and increased the expression of T-bet
GO:0016579 protein deubiquitination
IMP
PMID:24845384
Deubiquitination and stabilization of T-bet by USP10.
ACCEPT
Summary: Protein deubiquitination is the broad biological process that best summarizes USP10 catalytic action across multiple substrates.
Reason: Accept as core. Substrate-specific evidence supports deubiquitination of p53, CFTR, ribosomal proteins, LC3B, and immune-signaling substrates.
Supporting Evidence:
PMID:24845384
USP10, a carboxyl-terminal ubiquitin-processing protease, could interact with T-bet in the nucleus
PMID:24845384
Overexpression of USP10 directly inhibited T-bet ubiquitination and increased the expression of T-bet
GO:0004843 cysteine-type deubiquitinase activity
IMP
PMID:25861989
TRAF Family Member-associated NF-ΞΊB Activator (TANK) Inhibit...
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
PMID:25861989
TANK formed a complex with MCPIP1 (also known as ZC3H12A) and a deubiquitinase, USP10
PMID:25861989
USP10-dependent deubiquitination of TRAF6 and the resolution of genotoxic NF-ΞΊB activation upon DNA damage
PMID:25861989
TANK-MCPIP1-USP10 complex also decreased TRAF6 ubiquitination in cells treated with IL-1Ξ² or LPS
GO:0006974 DNA damage response
IMP
PMID:25861989
TRAF Family Member-associated NF-ΞΊB Activator (TANK) Inhibit...
KEEP AS NON CORE
Summary: USP10 participates in DNA-damage-linked signaling through p53 and NF-kappaB pathway substrate deubiquitination.
Reason: Keep as non-core. The cited evidence supports DNA-damage-response modulation, but the direct molecular role is deubiquitination of signaling proteins.
Supporting Evidence:
PMID:25861989
TANK formed a complex with MCPIP1 (also known as ZC3H12A) and a deubiquitinase, USP10
PMID:25861989
USP10-dependent deubiquitination of TRAF6 and the resolution of genotoxic NF-ΞΊB activation upon DNA damage
PMID:25861989
TANK-MCPIP1-USP10 complex also decreased TRAF6 ubiquitination in cells treated with IL-1Ξ² or LPS
GO:0016579 protein deubiquitination
IMP
PMID:25861989
TRAF Family Member-associated NF-ΞΊB Activator (TANK) Inhibit...
ACCEPT
Summary: Protein deubiquitination is the broad biological process that best summarizes USP10 catalytic action across multiple substrates.
Reason: Accept as core. Substrate-specific evidence supports deubiquitination of p53, CFTR, ribosomal proteins, LC3B, and immune-signaling substrates.
Supporting Evidence:
PMID:25861989
TANK formed a complex with MCPIP1 (also known as ZC3H12A) and a deubiquitinase, USP10
PMID:25861989
USP10-dependent deubiquitination of TRAF6 and the resolution of genotoxic NF-ΞΊB activation upon DNA damage
PMID:25861989
TANK-MCPIP1-USP10 complex also decreased TRAF6 ubiquitination in cells treated with IL-1Ξ² or LPS
GO:0043124 negative regulation of canonical NF-kappaB signal transduction
IMP
PMID:25861989
TRAF Family Member-associated NF-ΞΊB Activator (TANK) Inhibit...
KEEP AS NON CORE
Summary: USP10 promotes TRAF6/NEMO deubiquitination in a TANK-MCPIP1 complex to dampen canonical NF-kappaB signaling.
Reason: Keep as non-core. The signaling effect is supported but is substrate- and pathway-specific.
Supporting Evidence:
PMID:25861989
TANK formed a complex with MCPIP1 (also known as ZC3H12A) and a deubiquitinase, USP10
PMID:25861989
USP10-dependent deubiquitination of TRAF6 and the resolution of genotoxic NF-ΞΊB activation upon DNA damage
PMID:25861989
TANK-MCPIP1-USP10 complex also decreased TRAF6 ubiquitination in cells treated with IL-1Ξ² or LPS
GO:0071347 cellular response to interleukin-1
IMP
PMID:25861989
TRAF Family Member-associated NF-ΞΊB Activator (TANK) Inhibit...
KEEP AS NON CORE
Summary: USP10 participates in the IL-1beta/LPS NF-kappaB response through TRAF6 deubiquitination in the TANK-MCPIP1-USP10 complex.
Reason: Keep as non-core. This is a supported immune-signaling response context, not the core DUB function itself.
Supporting Evidence:
PMID:25861989
TANK formed a complex with MCPIP1 (also known as ZC3H12A) and a deubiquitinase, USP10
PMID:25861989
USP10-dependent deubiquitination of TRAF6 and the resolution of genotoxic NF-ΞΊB activation upon DNA damage
PMID:25861989
TANK-MCPIP1-USP10 complex also decreased TRAF6 ubiquitination in cells treated with IL-1Ξ² or LPS
GO:0005515 protein binding
IPI
PMID:25861989
TRAF Family Member-associated NF-ΞΊB Activator (TANK) Inhibit...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0032991 protein-containing complex
IDA
PMID:25861989
TRAF Family Member-associated NF-ΞΊB Activator (TANK) Inhibit...
MARK AS OVER ANNOTATED
Summary: USP10 participates in a TANK-MCPIP1-USP10 complex, but GO:0032991 is too generic to be an informative gene-level annotation.
Reason: Mark as over-annotated. The evidence supports a specific complex context for NF-kappaB signaling, not the broad term protein-containing complex.
Supporting Evidence:
PMID:25861989
TANK formed a complex with MCPIP1 (also known as ZC3H12A) and a deubiquitinase, USP10
GO:0005829 cytosol
TAS
Reactome:R-HSA-6781779
ACCEPT
Summary: Cytosol is a supported compartment for USP10 deubiquitinase activity and substrate regulation.
Reason: Accept as a supported cytosolic location for USP10 activity; cytosolic p53 regulation and ribosome-quality-control evidence are consistent with this assignment.
Supporting Evidence:
Reactome:R-HSA-6781779
USP13 can deubiquitinate USP10, an essential regulator of TP53 stability
GO:0005829 cytosol
TAS
Reactome:R-HSA-6782106
ACCEPT
Summary: Cytosol is a supported compartment for USP10 deubiquitinase activity and substrate regulation.
Reason: Accept as a supported cytosolic location for USP10 activity; cytosolic p53 regulation and ribosome-quality-control evidence are consistent with this assignment.
Supporting Evidence:
Reactome:R-HSA-6782106
USP10 deubiquitinates CFTR in early endosomes thereby enhancing its endocytic recycling
GO:0003723 RNA binding
HDA
PMID:22658674
Insights into RNA biology from an atlas of mammalian mRNA-bi...
MARK AS OVER ANNOTATED
Summary: The RNA-binding annotations come from broad high-throughput mRNA-bound-proteome studies and do not establish RNA binding as an informative USP10 function.
Reason: Mark as over-annotated. USP10 has G3BP/40S/stress-granule contexts, but the stronger gene-specific evidence supports DUB and ribosome-quality-control roles rather than RNA binding as a core MF.
GO:0003723 RNA binding
HDA
PMID:22681889
The mRNA-bound proteome and its global occupancy profile on ...
MARK AS OVER ANNOTATED
Summary: The RNA-binding annotations come from broad high-throughput mRNA-bound-proteome studies and do not establish RNA binding as an informative USP10 function.
Reason: Mark as over-annotated. USP10 has G3BP/40S/stress-granule contexts, but the stronger gene-specific evidence supports DUB and ribosome-quality-control roles rather than RNA binding as a core MF.
GO:0005654 nucleoplasm
TAS
Reactome:R-HSA-5653766
ACCEPT
Summary: Nucleoplasm is a supported location for USP10 in PCNA/TLS and TP53 deubiquitination Reactome contexts.
Reason: Accept as a nuclear subcompartment location for substrate-specific USP10 activities.
Supporting Evidence:
Reactome:R-HSA-5653766
Ubiquitin protease USP10 binds doubly ISGylated and monoubiquitinated PCNA
Reactome:R-HSA-5653770
USP10 acts as a ubiquitin protease to remove ubiquitin from lysine K164 residue of doubly ISGylated PCNA
Reactome:R-HSA-5653770
Deubiquitination of PCNA by USP10 causes dissociation of Y family DNA damage bypass polymerases
GO:0005654 nucleoplasm
TAS
Reactome:R-HSA-5653770
ACCEPT
Summary: Nucleoplasm is a supported location for USP10 in PCNA/TLS and TP53 deubiquitination Reactome contexts.
Reason: Accept as a nuclear subcompartment location for substrate-specific USP10 activities.
Supporting Evidence:
Reactome:R-HSA-5653766
Ubiquitin protease USP10 binds doubly ISGylated and monoubiquitinated PCNA
Reactome:R-HSA-5653770
USP10 acts as a ubiquitin protease to remove ubiquitin from lysine K164 residue of doubly ISGylated PCNA
Reactome:R-HSA-5653770
Deubiquitination of PCNA by USP10 causes dissociation of Y family DNA damage bypass polymerases
GO:0005654 nucleoplasm
TAS
Reactome:R-HSA-5689973
ACCEPT
Summary: Nucleoplasm is a supported location for USP10 in PCNA/TLS and TP53 deubiquitination Reactome contexts.
Reason: Accept as a nuclear subcompartment location for substrate-specific USP10 activities.
Supporting Evidence:
Reactome:R-HSA-5689973
USP10 specifically deubiquitinate p53 and not MDM2
GO:0004197 cysteine-type endopeptidase activity
IMP
PMID:21962518
Beclin1 controls the levels of p53 by regulating the deubiqu...
MODIFY
Summary: The cited Beclin1/spautin-1 study supports ubiquitin-specific peptidase activity, not a generic cysteine-type endopeptidase function.
Reason: Modify to the more specific and biologically correct cysteine-type deubiquitinase activity term.
Supporting Evidence:
PMID:21962518
two ubiquitin-specific peptidases, USP10 and USP13, that target the Beclin1 subunit of Vps34 complexes
GO:0004843 cysteine-type deubiquitinase activity
IDA
PMID:21962518
Beclin1 controls the levels of p53 by regulating the deubiqu...
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
PMID:21962518
two ubiquitin-specific peptidases, USP10 and USP13, that target the Beclin1 subunit of Vps34 complexes
GO:0005515 protein binding
IPI
PMID:21962518
Beclin1 controls the levels of p53 by regulating the deubiqu...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0010506 regulation of autophagy
IDA
PMID:21962518
Beclin1 controls the levels of p53 by regulating the deubiqu...
KEEP AS NON CORE
Summary: USP10 regulates autophagy through Beclin1/Vps34 complexes and LC3B deubiquitination, but this is one substrate pathway of the broader DUB function.
Reason: Keep as non-core. The process is well supported and relevant to the PN ATG8 context, but the molecular function should remain protein deubiquitination rather than a generic autophagy-regulator identity.
Supporting Evidence:
PMID:21962518
two ubiquitin-specific peptidases, USP10 and USP13, that target the Beclin1 subunit of Vps34 complexes
PMID:33577797
LC3B ubiquitination is reversed by the action of the deubiquitinating enzyme USP10
PMID:33577797
LC3B and autophagic activity are controlled through cycles of LC3B ubiquitination and deubiquitination
GO:0002039 p53 binding
IPI
PMID:20096447
USP10 regulates p53 localization and stability by deubiquiti...
KEEP AS NON CORE
Summary: USP10 binds p53 as part of its p53 deubiquitination/stabilization pathway.
Reason: Keep as non-core. p53 binding is substrate-specific and informative, but USP10 core function is catalytic deubiquitination.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0004843 cysteine-type deubiquitinase activity
IMP
PMID:19398555
The deubiquitinating enzyme USP10 regulates the post-endocyt...
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
PMID:19398555
USP10 is located in early endosomes and regulates the deubiquitination of CFTR
PMID:19398555
facilitating the deubiquitination of CFTR in early endosomes and thereby enhancing the endocytic recycling of CFTR
GO:0004843 cysteine-type deubiquitinase activity
IDA
PMID:20096447
USP10 regulates p53 localization and stability by deubiquiti...
ACCEPT
Summary: USP10 is a ubiquitin-specific cysteine deubiquitinase; this MF captures the core catalytic activity even though individual papers test different substrates.
Reason: Accept as the core molecular function. USP10 repeatedly removes ubiquitin from protein substrates including p53, CFTR, LC3B, TRAF6/NEMO contexts, and 40S ribosomal proteins.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0005515 protein binding
IPI
PMID:19398555
The deubiquitinating enzyme USP10 regulates the post-endocyt...
MARK AS OVER ANNOTATED
Summary: Generic protein binding is not informative for USP10 and obscures more specific substrate/context annotations.
Reason: Mark as over-annotated. Supported specific interactions include p53 binding, CFTR/transporter binding, G3BP stress-granule regulation, and substrate-specific deubiquitination; GO:0005515 should not be retained as a functional conclusion.
GO:0005634 nucleus
IDA
PMID:20096447
USP10 regulates p53 localization and stability by deubiquiti...
ACCEPT
Summary: USP10 translocates to or acts in the nucleus in p53 and T-bet substrate-stability contexts.
Reason: Accept as a supported location, while treating the specific nuclear signaling outputs as substrate/context-specific rather than the sole core function.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0005737 cytoplasm
IDA
PMID:20096447
USP10 regulates p53 localization and stability by deubiquiti...
ACCEPT
Summary: USP10 is a cytoplasmic/cytosolic DUB and several core substrate contexts occur in the cytoplasm.
Reason: Accept as a supported cellular location for USP10 deubiquitination, including p53 homeostasis, RQC, and immune-signaling substrates.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0005769 early endosome
IDA
PMID:19398555
The deubiquitinating enzyme USP10 regulates the post-endocyt...
ACCEPT
Summary: USP10 localizes to early endosomes where it deubiquitinates CFTR and supports endocytic recycling.
Reason: Accept as a supported active compartment for a direct USP10 substrate context.
Supporting Evidence:
PMID:19398555
USP10 is located in early endosomes and regulates the deubiquitination of CFTR
PMID:19398555
facilitating the deubiquitination of CFTR in early endosomes and thereby enhancing the endocytic recycling of CFTR
GO:0016579 protein deubiquitination
IMP
PMID:19398555
The deubiquitinating enzyme USP10 regulates the post-endocyt...
ACCEPT
Summary: Protein deubiquitination is the broad biological process that best summarizes USP10 catalytic action across multiple substrates.
Reason: Accept as core. Substrate-specific evidence supports deubiquitination of p53, CFTR, ribosomal proteins, LC3B, and immune-signaling substrates.
Supporting Evidence:
PMID:19398555
USP10 is located in early endosomes and regulates the deubiquitination of CFTR
PMID:19398555
facilitating the deubiquitination of CFTR in early endosomes and thereby enhancing the endocytic recycling of CFTR
GO:0016579 protein deubiquitination
IDA
PMID:20096447
USP10 regulates p53 localization and stability by deubiquiti...
ACCEPT
Summary: Protein deubiquitination is the broad biological process that best summarizes USP10 catalytic action across multiple substrates.
Reason: Accept as core. Substrate-specific evidence supports deubiquitination of p53, CFTR, ribosomal proteins, LC3B, and immune-signaling substrates.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0030330 DNA damage response, signal transduction by p53 class mediator
IMP
PMID:20096447
USP10 regulates p53 localization and stability by deubiquiti...
KEEP AS NON CORE
Summary: USP10 activates p53 signaling after DNA damage by deubiquitinating/stabilizing p53 and translocating to the nucleus.
Reason: Keep as non-core. The annotation is supported but represents a substrate-specific signaling outcome of USP10 DUB activity.
Supporting Evidence:
PMID:20096447
USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
PMID:20096447
After DNA damage, USP10 is stabilized, and a fraction of USP10 translocates to the nucleus to activate p53
GO:0044325 transmembrane transporter binding
IDA
PMID:19398555
The deubiquitinating enzyme USP10 regulates the post-endocyt...
KEEP AS NON CORE
Summary: USP10 binds/regulates the transmembrane transporter CFTR in early endosomes.
Reason: Keep as non-core. The annotation reflects a supported CFTR substrate context, not broad transporter-binding specificity.
Supporting Evidence:
PMID:19398555
USP10 is located in early endosomes and regulates the deubiquitination of CFTR
PMID:19398555
facilitating the deubiquitination of CFTR in early endosomes and thereby enhancing the endocytic recycling of CFTR

Core Functions

USP10 is a ubiquitin-specific cysteine deubiquitinase that removes ubiquitin from protein substrates. This broad catalytic function underlies its p53, CFTR, LC3B/Beclin1, NF-kappaB, MAVS, and ribosome-quality-control activities.

Supporting Evidence:
  • Reactome:R-HSA-5688426
    Deubiquitinating enzymes (DUBs) catalyze the removal of Ub and regulate Ub-mediated pathways
  • PMID:20096447
    USP10, a cytoplasmic ubiquitin-specific protease, deubiquitinates p53
  • PMID:19398555
    USP10 is located in early endosomes and regulates the deubiquitination of CFTR
  • PMID:31981475
    G3BP1-family-USP10 complexes are required for deubiquitination of RPS2, RPS3, and RPS10 to rescue modified 40S subunits from programmed degradation
  • PMID:34348161
    These impeded ribosomes are tagged by ubiquitin at their 40S subunit for subsequent programmed degradation unless rescued by USP10
  • PMID:34469731
    USP10 as the deubiquitylating enzyme responsible for removing ubiquitin from uS3 and uS5
  • PMID:33577797
    LC3B ubiquitination is reversed by the action of the deubiquitinating enzyme USP10

References

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Suggested Questions for Experts

Q: Should USP10 and UCHL1 be annotated to a shared ATG8-family protein deubiquitination term for LC3/ATG8 substrate deubiquitination?

Suggested experts: GO autophagy editors, Juan S. Bonifacino, Ai Yamamoto

Q: Should 40S ribosomal protein deubiquitination during ribosome quality control be represented as a substrate-specific child of monoubiquitinated protein deubiquitination?

Suggested experts: Eric J. Bennett, Judith Frydman, GO proteostasis editors

Q: Which USP10 substrate contexts should remain non-core gene-level annotations versus annotation extensions on the core deubiquitinase activity?

Suggested experts: GO ubiquitin editors, GO-CAM editors

Suggested Experiments

Experiment: Map endogenous LC3A/LC3B/LC3C ubiquitination sites in USP10 knockout and catalytic-rescue cells, then test whether catalytically inactive USP10 fails to restore LC3 abundance and autophagy flux.

Hypothesis: USP10 directly deubiquitinates LC3-family ATG8 proteins to preserve LC3 abundance and stress-induced autophagy.

Type: substrate mapping and catalytic rescue assay

Experiment: Use ribosome profiling, ribosomal-protein ubiquitinomics, and 40S turnover assays in USP10 catalytic mutants to separate elongation RQC from initiation RQC substrates.

Hypothesis: USP10 rescues distinct classes of ubiquitinated 40S subunits by removing substrate-specific mono-ubiquitin marks from RPS2/RPS3/RPS10.

Type: ribosome quality-control ubiquitinomics

Experiment: Build substrate-specific USP10 GO-CAM models for p53, CFTR, LC3B, TRAF6/NEMO, MAVS, and 40S ribosomal proteins and compare which contexts are conserved across cell types.

Hypothesis: Most USP10 biological-process annotations are substrate-context outputs of a single core DUB activity rather than independent core functions.

Type: curation-focused comparative substrate model

πŸ“š Additional Documentation

Notes

(USP10-notes.md)

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Pn Notes

(USP10-pn-notes.md)

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πŸ“„ View Raw YAML

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